159 lines
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159 lines
11 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.186.62018" ID-GBIF-Dataset="8becf6a0-f9c4-495b-9620-2040bd691f43" ID-PMC="PMC8671706" ID-Pensoft-Pub="1314-2003-186-93" ID-Pensoft-UUID="51C402FAC1635A0FB18EA14A1C2FE5F8" ID-PubMed="34975277" ModsDocID="1314-2003-186-93" checkinTime="1638850659404" checkinUser="pensoft" docAuthor="Kiew, Ruth & Kamin, Imin" docDate="2021" docId="8CA3B4DAB04F5703A7A2893C9418F14F" docLanguage="en" docName="PhytoKeys 186: 93-110" docOrigin="PhytoKeys 186" docPubDate="2021-12-06" docSource="http://dx.doi.org/10.3897/phytokeys.186.62018" docTitle="Coleus kunstleri A. J. Paton in Paton et al., Phytokeys 129 2019" docType="treatment" docVersion="4" id="51C402FAC1635A0FB18EA14A1C2FE5F8" lastPageNumber="93" masterDocId="51C402FAC1635A0FB18EA14A1C2FE5F8" masterDocTitle="Coleus (Lamiaceae) in Peninsular Malaysia including two new species" masterLastPageNumber="110" masterPageNumber="93" pageNumber="93" updateTime="1668140546889" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Coleus (Lamiaceae) in Peninsular Malaysia including two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kiew, Ruth</mods:namePart>
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<mods:affiliation>Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
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<mods:nameIdentifier type="email">ruth@frim.gov.my</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kamin, Imin</mods:namePart>
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<mods:affiliation>Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>186</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>93</mods:start>
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<mods:end>110</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.186.62018</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.186.62018</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-186-93</mods:identifier>
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<mods:identifier type="Pensoft-UUID">51C402FAC1635A0FB18EA14A1C2FE5F8</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191162306" LSID="urn:lsid:plazi:treatment:8CA3B4DAB04F5703A7A2893C9418F14F" httpUri="http://treatment.plazi.org/id/8CA3B4DAB04F5703A7A2893C9418F14F" lastPageNumber="93" pageId="0" pageNumber="93">
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<subSubSection pageId="0" pageNumber="93" type="nomenclature">
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<paragraph pageId="0" pageNumber="93">
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2.
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<taxonomicName LSID="8CA3B4DA-B04F-5703-A7A2-893C9418F14F" authority="(Prain) A. J. Paton in Paton et al., Phytokeys 129 (2019) 63." authorityName="A. J. Paton in Paton et al., Phytokeys 129" authorityYear="2019" baseAuthorityName="Prain" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus kunstleri" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="kunstleri">Coleus kunstleri (Prain) A.J. Paton in Paton et al., Phytokeys 129 (2019) 63.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="reference_group">
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<paragraph pageId="0" pageNumber="93">
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<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus kunstleri" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="kunstleri">Plectranthus kunstleri</taxonomicName>
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Homotypic synonym:
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<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus kunstleri" order="Lamiales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="kunstleri">Plectranthus kunstleri</taxonomicName>
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Prain, J. Asiat. Soc. Bengal 74, 2: 706 (1907); Ridley, Fl. Mal. Penin. 2: 646 (1923); Keng, Gard. Bull. Singapore 24: 151 (1969), Fl. Malesiana Ser. I, 8: 392 (1978); Chin, Gard. Bull. Singapore 32: 160 (1979); Turner, Gard. Bull. Singapore. 47: 272 (1996
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<normalizedToken originalValue="‘1995’">'1995'</normalizedToken>
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); Bramley, Fl. Malesiana Ser. I, 23: 293 (2019).Type: Malaysia. Peninsular Malaysia: Perak, Kuala Dipang, top of limestone hills, 130-200 m. Sept. 1885.
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<emphasis italics="true" pageId="0" pageNumber="93">Kunstler</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="93">
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<normalizedToken originalValue="King’s">King's</normalizedToken>
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collector
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</emphasis>
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)
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<emphasis italics="true" pageId="0" pageNumber="93">8240</emphasis>
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(holotype CAL; isotypes BM!, K!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="description">
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<paragraph pageId="0" pageNumber="93">Description.</paragraph>
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<paragraph pageId="0" pageNumber="93">
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Erect herb, almost shrubby, 0.5-1.2(-1.5) m high.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Stem</emphasis>
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and branches finely puberulous, angular.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Leaves</emphasis>
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with petiole 1-4.5 cm; lamina membranaceous, ovate to broadly ovate, 6-10(-13)
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<normalizedToken originalValue="×">x</normalizedToken>
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4-6 cm, base truncate or shortly cuneate, margin elsewhere remotely crenate, apex acute, very sparsely pubescent on the main and secondary veins on both surfaces, sometimes with a whitish line down midrib.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Inflorescence</emphasis>
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terminal, paniculate, 10-20 cm long, the lower branches of the inflorescence axis 5-6 cm, decreasing in length towards apex. Bracts ovate, acute, 2-3 mm long, caducous. Pedicels 3-4 mm long, glandular-puberulent.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Flowers</emphasis>
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spreading, (not pendent);
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<emphasis italics="true" pageId="0" pageNumber="93">calyx</emphasis>
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obliquely campanulate, 1-2(-2.5) mm long, in fruit 5-7 mm, shortly hirsute and sparingly gland-dotted, unequally 5-lobed; upper lip ovate, broadly ovate or sub-rounded, lower lip with two lateral ovate-acute lobes shorter (about
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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) than the lower central lobes but later almost equal in fruit, the two central lobes subulate, connate beneath;
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<emphasis italics="true" pageId="0" pageNumber="93">corolla</emphasis>
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waxy white or greenish white (
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<emphasis italics="true" pageId="0" pageNumber="93">Kunstler 8240</emphasis>
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) or purplish pink (
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<emphasis italics="true" pageId="0" pageNumber="93">Chin 1760</emphasis>
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), 7-8(-10) mm long, puberulent, tube sigmoid, decurved, only slightly gibbous near the base, posterior lip short, erect, anterior lip concave, boat-shaped, c. 4 mm long;
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<emphasis italics="true" pageId="0" pageNumber="93">stamens</emphasis>
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not exserted, lying within the anterior lip, free above point of insertion on the corolla tube.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Nutlets</emphasis>
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oblong ovoid, black,
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<emphasis italics="true" pageId="0" pageNumber="93">ca.</emphasis>
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1 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="distribution">
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<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="93">
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Endemic in Peninsular Malaysia, Perak - Kinta (no specific locality) and Kuala Dipang; and Kedah - Langkawi, Pulau Langgun (Map
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<figureCitation captionStart="Map 1" captionStartId="F1" captionText="Map 1. Distribution of Coleus hairulii (circle), C. kunstleri (triangle) and C. rafidahiae (star) in Peninsular Malaysia." figureDoi="10.3897/phytokeys.186.62018.map1" httpUri="https://binary.pensoft.net/fig/619121" pageId="0" pageNumber="93">1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="provisional conservation status">
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<paragraph pageId="0" pageNumber="93">Provisional conservation status.</paragraph>
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<paragraph pageId="0" pageNumber="93">Critically Endangered B1ab(i,iii). Limestone hills in Perak are regularly visited for botanical collecting but this species has not been recollected there for over a hundred years and the majority of karst limestone hills in the Kinta Valley are heavily disturbed by quarries. The species was last collected in the 1970s from Pulau Langgun. (Assessed by A.R. Rafidah).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="ecology">
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<paragraph pageId="0" pageNumber="93">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="93">
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Restricted to limestone rocks, in shade, at 50-200 m elevation, either in open habitats on top of a limestone hill to 200 m (
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<emphasis italics="true" pageId="0" pageNumber="93">Kunstler 8240</emphasis>
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) or on huge boulders rooting in rock crevices and on humus accumulated on boulders under shade of scattered trees at ca. 50 m (
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<emphasis italics="true" pageId="0" pageNumber="93">Chin 1760</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="etymology">
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<paragraph pageId="0" pageNumber="93">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="93">
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H.H. Kunstler, a German botanist, who collected plants in 1880-1886 mostly from Perak, Peninsular Malaysia, for Sir George King, Director of Calcutta Botanic Garden. He is also recorded on herbarium labels as
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<normalizedToken originalValue="‘King’s">'King's</normalizedToken>
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<normalizedToken originalValue="Collector’">Collector'</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="93" type="materials_examined">
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<paragraph pageId="0" pageNumber="93">Additional specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="93">
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<emphasis bold="true" pageId="0" pageNumber="93">
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Peninsular
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Malaysia" location="Perak" specimenCount="1" stateProvince="Perak">
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<collectingCountry name="Malaysia">Malaysia</collectingCountry>
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.
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<collectingRegion country="Malaysia" name="Perak">Perak</collectingRegion>
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: Kinta (
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<emphasis italics="true" pageId="0" pageNumber="93">Kunstler 7143</emphasis>
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, BM!)
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</materialsCitation>
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</emphasis>
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.
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<collectingRegion country="Malaysia" name="Kedah">Kedah</collectingRegion>
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: Langkawi, Pulau Langgun (
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<emphasis italics="true" pageId="0" pageNumber="93">Chin 1760</emphasis>
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, KLU!).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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