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<document id="59F0771B27DBFCCDE7D85980B730E041" ID-CLB-Dataset="28123" ID-DOI="10.11646/zootaxa.4568.3.2" ID-GBIF-Dataset="d40eb95b-5a98-418a-b607-7c2cbec7c897" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601607" ID-ZooBank="8FD0AC89-424E-4CAC-92A5-A5600A481140" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172834442" checkinUser="plazi" docAuthor="Kozłowska, Anna, Bates, Denis, Zalasiewicz, Jan &amp; Radzevičius, Sigitas" docDate="2019" docId="03A687FCFFFA985E8FA2D7AEFC44FE6E" docLanguage="en" docName="zootaxa.4568.3.2.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gothograptus auriculatus Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="454" masterDocId="FF9FFF84FFEA984D8F35D37CFD67FFA1" masterDocTitle="Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania" masterLastPageNumber="469" masterPageNumber="435" pageNumber="451" updateTime="1698703580205" updateUser="ExternalLinkService">
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<mods:title id="4696CED1DE805EB86FB1F67BE64D37E5">Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania</mods:title>
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<mods:namePart id="B6DF9516DDFFB0A42C03B6A4DD6B0C5F">Kozłowska, Anna</mods:namePart>
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<mods:namePart id="0921028CB15A94C36B97280405754FFF">Bates, Denis</mods:namePart>
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<mods:namePart id="7F26EFB13C3D2F4777A7B541A5F9D52A">Zalasiewicz, Jan</mods:namePart>
</mods:name>
<mods:name id="D2FA5F5797B60B6CA2E362963BDFD314" type="personal">
<mods:role id="B4ADA0875B2759E96BB6E760D7F5EB7F">
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<mods:namePart id="0FBD582A5AB4073AAC9A465F257AEFDE">Radzevičius, Sigitas</mods:namePart>
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<mods:title id="FFA4B263BD234BF2778D307D6B0E6041">Zootaxa</mods:title>
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<mods:date id="F594C289DF1AE0FF26964C95ABBF414D">2019</mods:date>
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<mods:number id="3772C0334941BB7FE0DF2E9B14F6EE39">2019-03-21</mods:number>
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<mods:number id="0C25A618187F375A66B39AC56B226F69">4568</mods:number>
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<mods:classification id="82E3A9C868E4AAB7D29963C5BD472593">journal article</mods:classification>
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<treatment id="03A687FCFFFA985E8FA2D7AEFC44FE6E" ID-DOI="http://doi.org/10.5281/zenodo.5935880" ID-GBIF-Taxon="154474978" ID-Zenodo-Dep="5935880" LSID="urn:lsid:plazi:treatment:03A687FCFFFA985E8FA2D7AEFC44FE6E" httpUri="http://treatment.plazi.org/id/03A687FCFFFA985E8FA2D7AEFC44FE6E" lastPageId="19" lastPageNumber="454" pageId="16" pageNumber="451">
<subSubSection id="C3156561FFFA985D8FA2D7AEFF7FFB4C" box="[151,536,1234,1261]" pageId="16" pageNumber="451" type="nomenclature">
<paragraph id="8BB036EAFFFA985D8FA2D7AEFF7FFB4C" blockId="16.[151,536,1234,1294]" box="[151,536,1234,1261]" pageId="16" pageNumber="451">
<heading id="D0F88186FFFA985D8FA2D7AEFF7FFB4C" bold="true" box="[151,536,1234,1261]" fontSize="11" level="1" pageId="16" pageNumber="451" reason="1">
<emphasis id="B97BEAF8FFFA985D8FA2D7AEFF7FFB4C" bold="true" box="[151,536,1234,1261]" pageId="16" pageNumber="451">
<taxonomicName id="4C0F4D69FFFA985D8FA2D7AEFCA9FB4D" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[151,462,1234,1260]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFFA985D8FA2D7AEFCA9FB4D" bold="true" box="[151,462,1234,1260]" italics="true" pageId="16" pageNumber="451">Gothograptus auriculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2485783FFFA985D8EE0D7AFFF7FFB4C" box="[469,536,1235,1261]" pageId="16" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FA2D789FEABFAF2" pageId="16" pageNumber="451" type="description">
<paragraph id="8BB036EAFFFA985D8FA2D789FC72FAAF" blockId="16.[151,536,1234,1294]" box="[151,277,1269,1294]" pageId="16" pageNumber="451">
<figureCitation id="13342A6FFFFA985D8FA2D789FC72FAAF" box="[151,277,1269,1294]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="17.[151,250,1830,1852]" captionStartId-1="18.[151,250,1778,1800]" captionTargetBox-0="[151,1436,265,1809]" captionTargetBox-1="[151,1435,556,1757]" captionTargetId-0="figure@17.[151,1436,265,1809]" captionTargetId-1="figure@18.[151,1435,556,1757]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." captionText-1="FIGURE 14. Gothograptus auriculatus n. sp., proximal ends of tubaria, Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. immature tubarium with the first three thecae, reverse view, ZPAL G.55/27. B. ventro-lateral side of tubarium, ZPAL G.55/28. C–E. reverse views of mature tubaria with very wide lists surrounding proximal lateral orifice and prongs; C. ZPAL G.55/23; D. lateral view, ZPAL G.55/29; F." httpUri-0="https://zenodo.org/record/2601645/files/figure.png" httpUri-1="https://zenodo.org/record/2601647/files/figure.png" pageId="16" pageNumber="451">Figs 13–14</figureCitation>
</paragraph>
<paragraph id="8BB036EAFFFA985D8FA2D641FEABFAF2" blockId="16.[151,972,1341,1363]" box="[151,972,1341,1363]" pageId="16" pageNumber="451">
2006
<taxonomicName id="4C0F4D69FFFA985D8FE5D641FE5FFAF2" authority=", Calner, Kozlowska, Masiak &amp; Schmitz" authorityName="Calner, Kozlowska, Masiak &amp; Schmitz" box="[208,824,1341,1363]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8FE5D641FCFAFAF2" box="[208,413,1341,1363]" italics="true" pageId="16" pageNumber="451">Gothograptus nassa</emphasis>
, Calner, Kozłowska, Masiak &amp; Schmitz
</taxonomicName>
, p. 80, fig. 1E.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FA2D6FCF948FA40" pageId="16" pageNumber="451" type="materials_examined">
<paragraph id="8BB036EAFFFA985D8FA2D6FCF948FA40" blockId="16.[151,1436,1408,2012]" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FA2D6FCFC59FA38" bold="true" box="[151,318,1408,1433]" pageId="16" pageNumber="451">Type material</emphasis>
.
<materialsCitation id="3B673CB7FFFA985D8E78D6FCF948FA40" ID-GBIF-Occurrence="2013965662" collectionCode="ZPAL" country="Poland" elevation="1645" location="Bartoszyce" pageId="16" pageNumber="451" specimenCount="2" typeStatus="holotype">
<typeStatus id="54B48848FFFA985D8E78D6FCFCD2FA38" box="[333,437,1408,1433]" pageId="16" pageNumber="451" type="holotype">Holotype</typeStatus>
<collectionCode id="ED1EAE2FFFFA985D8E88D6FDFF66FA38" box="[445,513,1409,1433]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="16" pageNumber="451">ZPAL</collectionCode>
G.55/23, mature specimen with six pairs of thecae and distal end broken off (
<figureCitation id="13342A6FFFFA985D8A5AD6FCFDA8FA1D" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="16" pageNumber="451">Fig. 13 B</figureCitation>
);
<typeStatus id="54B48848FFFA985D8FD1D6D9FC23FA1C" box="[228,324,1445,1469]" pageId="16" pageNumber="451" type="paratype">paratype</typeStatus>
<collectionCode id="ED1EAE2FFFFA985D8E7ED6D8FCE8FA1D" box="[331,399,1444,1468]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="16" pageNumber="451">ZPAL</collectionCode>
G.55/24, distal end of mature tubarium with five pairs of thecae and appendix (
<figureCitation id="13342A6FFFFA985D8A30D6D8F8E8FA1D" box="[1285,1423,1444,1469]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="16" pageNumber="451">Fig. 13 C, F</figureCitation>
).
<location id="8ED06031FFFA985D8FA2D6B5FC73FA40" LSID="urn:lsid:plazi:treatment:03A687FCFFFA985E8FA2D7AEFC44FE6E:8ED06031FFFA985D8FA2D6B5FC73FA40" box="[151,276,1481,1505]" country="Poland" name="Bartoszyce" pageId="16" pageNumber="451">Bartoszyce</location>
IG-1well, depth
<quantity id="4CF79B0FFFFA985D8EE6D6B4FF58FA41" box="[467,575,1480,1505]" metricMagnitude="3" metricUnit="m" metricValue="1.6452" pageId="16" pageNumber="451" unit="m" value="1645.2">
<elevation id="0022D1D9FFFA985D8EE6D6B4FF58FA41" box="[467,575,1480,1505]" metricMagnitude="3" metricUnit="m" metricValue="1.6452" pageId="16" pageNumber="451" unit="m" value="1645.2">1645.2 m</elevation>
</quantity>
,
<collectingCountry id="F318767AFFFA985D8D7FD6B4FFFCFA40" box="[586,667,1480,1505]" name="Poland" pageId="16" pageNumber="451">Poland</collectingCountry>
, lower part of
<taxonomicName id="4C0F4D69FFFA985D8C08D6B4F948FA40" authority="Biozone." authorityName="Biozone." box="[829,1071,1480,1505]" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFFA985D8C08D6B4FEA4FA41" box="[829,963,1480,1504]" italics="true" pageId="16" pageNumber="451">praedeubeli</emphasis>
Biozone.
</taxonomicName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FF2D690F9C7F9A5" box="[199,1184,1516,1541]" pageId="16" pageNumber="451" type="etymology">
<paragraph id="8BB036EAFFFA985D8FF2D690F9C7F9A5" blockId="16.[151,1436,1408,2012]" box="[199,1184,1516,1541]" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FF2D690FC20F9A4" bold="true" box="[199,327,1516,1541]" pageId="16" pageNumber="451">Etymology</emphasis>
. Latin
<taxonomicName id="4C0F4D69FFFA985D8EA3D690FF71F9A5" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[406,534,1516,1540]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="auriculatus">
<emphasis id="B97BEAF8FFFA985D8EA3D690FF71F9A5" box="[406,534,1516,1540]" italics="true" pageId="16" pageNumber="451">auriculatus</emphasis>
</taxonomicName>
—relating to the earlobe shape of the genicular structures.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FF2D56CFDB1F934" pageId="16" pageNumber="451" type="diagnosis">
<paragraph id="8BB036EAFFFA985D8FF2D56CFDB1F934" blockId="16.[151,1436,1408,2012]" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FF2D56CFC5DF988" bold="true" box="[199,314,1552,1577]" pageId="16" pageNumber="451">Diagnosis</emphasis>
. Tubarium with genicular structures formed by the
<taxonomicName id="4C0F4D69FFFA985D8CB0D56DFEA2F989" box="[901,965,1553,1576]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8CB0D56DFEA2F989" box="[901,965,1553,1576]" italics="true" pageId="16" pageNumber="451">nassa</emphasis>
</taxonomicName>
<typeStatus id="54B48848FFFA985D8CF8D56DFE9AF988" box="[973,1021,1553,1577]" pageId="16" pageNumber="451">type</typeStatus>
of hood, and paired vertical auriclelike structures bordering both sides of the thecal orifices. Auricle structures are characteristic only of
<taxonomicName id="4C0F4D69FFFA985D8A35D548FC70F9D1" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFFA985D8A35D548FC70F9D1" italics="true" pageId="16" pageNumber="451">Gothograptus auriculatus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFFA985D8E2BD524FC39F9D0" bold="true" box="[286,350,1624,1649]" pageId="16" pageNumber="451">
<taxonomicNameLabel id="A2485783FFFA985D8E2BD524FC39F9D0" box="[286,350,1624,1649]" pageId="16" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
In mature colonies the auricles bend towards the mid-ventral list below the thecal lip and
<taxonomicName id="4C0F4D69FFFA985D8A69D525F8FBF9D1" box="[1372,1436,1625,1648]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8A69D525F8FBF9D1" box="[1372,1436,1625,1648]" italics="true" pageId="16" pageNumber="451">nassa</emphasis>
</taxonomicName>
hood.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FF2D5DCFFE7F8A0" pageId="16" pageNumber="451" type="discussion">
<paragraph id="8BB036EAFFFA985D8FF2D5DCFFE7F8A0" blockId="16.[151,1436,1408,2012]" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FF2D5DCFC53F918" bold="true" box="[199,308,1696,1721]" pageId="16" pageNumber="451">Remarks</emphasis>
.
<taxonomicName id="4C0F4D69FFFA985D8E70D5DCFC82F919" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[325,485,1696,1720]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFFA985D8E70D5DCFC82F919" box="[325,485,1696,1720]" italics="true" pageId="16" pageNumber="451">G. auriculatus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFFA985D8EC5D5DCFF55F918" bold="true" box="[496,562,1696,1721]" pageId="16" pageNumber="451">
<taxonomicNameLabel id="A2485783FFFA985D8EC5D5DCFF55F918" box="[496,562,1696,1721]" pageId="16" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C0F4D69FFFA985D8DFAD5DCFE56F919" box="[719,817,1696,1720]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8DFAD5DCFE56F919" box="[719,817,1696,1720]" italics="true" pageId="16" pageNumber="451">G. nassa</emphasis>
</taxonomicName>
in having two lateral auricles, as additional elements located on the lateral sides of the thecal orifices and below the lips. Young colonies of the new species have
<taxonomicName id="4C0F4D69FFFA985D8A69D5B9F8FBF97D" box="[1372,1436,1733,1756]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8A69D5B9F8FBF97D" box="[1372,1436,1733,1756]" italics="true" pageId="16" pageNumber="451">nassa</emphasis>
</taxonomicName>
hoods; the auricles appear later in astogeny.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985D8FF2D470FC93F830" pageId="16" pageNumber="451" type="materials_examined">
<paragraph id="8BB036EAFFFA985D8FF2D470FC93F830" blockId="16.[151,1436,1408,2012]" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FF2D470FC48F884" bold="true" box="[199,303,1804,1829]" pageId="16" pageNumber="451">Material</emphasis>
. The material studied comes from
<collectingCountry id="F318767AFFFA985D8DE4D470FE45F884" box="[721,802,1804,1829]" name="Poland" pageId="16" pageNumber="451">Poland</collectingCountry>
,
<collectingCountry id="F318767AFFFA985D8C04D470FEFAF884" box="[817,925,1804,1829]" name="Lithuania" pageId="16" pageNumber="451">Lithuania</collectingCountry>
and
<collectingCountry id="F318767AFFFA985D8CE9D470F921F885" box="[988,1094,1804,1828]" name="Germany" pageId="16" pageNumber="451">Germany</collectingCountry>
. The most abundant material comes from the Bartoszyce IG- well, ranges from 1649.0 to
<quantity id="4CF79B0FFFFA985D8C74D44CFEC9F8E9" box="[833,942,1840,1865]" metricMagnitude="3" metricUnit="m" metricValue="1.6452" pageId="16" pageNumber="451" unit="m" value="1645.2">1645.2 m</quantity>
,
<taxonomicName id="4C0F4D69FFFA985D8C8FD44CF9C0F8E8" authority="Biozone" authorityName="Biozone" box="[954,1191,1840,1865]" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFFA985D8C8FD44CF927F8E9" box="[954,1088,1840,1864]" italics="true" pageId="16" pageNumber="451">praedeubeli</emphasis>
Biozone
</taxonomicName>
, yielding about 1300, mostly young, specimens. Two fragments of
<taxonomicName id="4C0F4D69FFFA985D8DA7D428FE56F8CD" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[658,817,1876,1900]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="auriculatus">
<emphasis id="B97BEAF8FFFA985D8DA7D428FE56F8CD" box="[658,817,1876,1900]" italics="true" pageId="16" pageNumber="451">G. auriculatus</emphasis>
</taxonomicName>
come from
<collectingCountry id="F318767AFFFA985D8C8FD428F94FF8CC" box="[954,1064,1876,1901]" name="Lithuania" pageId="16" pageNumber="451">Lithuania</collectingCountry>
, Pilviskiai-141 core, depth
<quantity id="4CF79B0FFFFA985D8A68D428FDC8F831" metricMagnitude="2" metricUnit="m" metricValue="8.422" pageId="16" pageNumber="451" unit="m" value="842.2">842.2 m</quantity>
,
<taxonomicName id="4C0F4D69FFFA985D8F8ED405FD9CF831" box="[187,251,1913,1936]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFFA985D8F8ED405FD9CF831" box="[187,251,1913,1936]" italics="true" pageId="16" pageNumber="451">nassa</emphasis>
</taxonomicName>
/
<taxonomicName id="4C0F4D69FFFA985D8E37D404FC93F830" authority="Biozone." authorityName="Biozone." box="[258,500,1912,1937]" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="16" pageNumber="451" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFFA985D8E37D404FCE0F831" box="[258,391,1912,1936]" italics="true" pageId="16" pageNumber="451">praedeubeli</emphasis>
Biozone.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFFA985E8FF2D4E2FE1CFEBC" lastPageId="19" lastPageNumber="454" pageId="16" pageNumber="451" type="description">
<paragraph id="8BB036EAFFFA985C8FF2D4E2FEF2FF74" blockId="16.[151,1436,1408,2012]" lastBlockId="17.[151,1436,151,213]" lastPageId="17" lastPageNumber="452" pageId="16" pageNumber="451">
<emphasis id="B97BEAF8FFFA985D8FF2D4E2FC36F816" bold="true" box="[199,337,1950,1975]" pageId="16" pageNumber="451">Description</emphasis>
. Tubaria slightly widening up to th1
<superScript id="7C7A9BA2FFFA985D8DDFD4E7FF95F808" attach="none" box="[746,754,1947,1961]" fontSize="6" pageId="16" pageNumber="451">2</superScript>
, then parallel-sided and gently tapering distally, ended by an appendix. Most of the specimens are represented by fragments of tubaria up to the 11th thecal pair. Tubarium width at the level of the first thecal pair is about
<quantity id="4CF79B0FFFFB985C8DB0D3EBFFB8FF0E" box="[645,735,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="452" unit="mm" value="1.5">1.5 mm</quantity>
, and in the middle part of the tubarium
<quantity id="4CF79B0FFFFB985C8B9AD3E4F86DFF0E" box="[1199,1290,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="452" unit="mm" value="1.2">1.2 mm</quantity>
. The longest fragment of tubarium, comprising 11 pairs of thecae, is
<quantity id="4CF79B0FFFFB985C8C33D3C0FE32FF75" box="[774,853,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="452" unit="mm" value="10.0">10 mm</quantity>
long.
</paragraph>
<caption id="DF706662FFFB985C8FA2D45AFCFDF877" httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="17" pageNumber="452" startId="17.[151,250,1830,1852]" targetBox="[151,1436,265,1809]" targetPageId="17">
<paragraph id="8BB036EAFFFB985C8FA2D45AFCFDF877" blockId="17.[151,1436,1830,2006]" pageId="17" pageNumber="452">
<emphasis id="B97BEAF8FFFB985C8FA2D45AFC78F89D" bold="true" box="[151,287,1830,1852]" pageId="17" pageNumber="452">FIGURE 13.</emphasis>
Mature tubaria of
<taxonomicName id="4C0F4D69FFFB985C8EEBD45AFF83F89D" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[478,740,1830,1852]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="17" pageNumber="452" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFFB985C8EEBD45AFF83F89D" box="[478,740,1830,1852]" italics="true" pageId="17" pageNumber="452">Gothograptus auriculatus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFFB985C8DDED45AFE43F89D" bold="true" box="[747,804,1830,1852]" pageId="17" pageNumber="452">
<taxonomicNameLabel id="A2485783FFFB985C8DDED45AFE43F89D" box="[747,804,1830,1852]" pageId="17" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Bartoszyce IG-1 core, depth 1645.2 m,
<taxonomicName id="4C0F4D69FFFB985C8BF7D45AF8FFF89D" authority="Biozone" authorityName="Biozone" box="[1218,1432,1830,1852]" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="17" pageNumber="452" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFFB985C8BF7D45AF85CF89D" box="[1218,1339,1830,1852]" italics="true" pageId="17" pageNumber="452">praedeubeli</emphasis>
Biozone
</taxonomicName>
, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end,
<emphasis id="B97BEAF8FFFB985C8D3AD418FF0BF8DB" bold="true" box="[527,620,1892,1914]" pageId="17" pageNumber="452">holotype</emphasis>
, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix,
<emphasis id="B97BEAF8FFFB985C8E1AD4FEFCF7F839" bold="true" box="[303,400,1922,1944]" pageId="17" pageNumber="452">paratype</emphasis>
ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26.
</paragraph>
</caption>
<paragraph id="8BB036EAFFF8985F8FF2D3EBFC0EFE29" blockId="18.[151,1437,151,503]" pageId="18" pageNumber="453">
The lists of the young stages of colony are very thin, about 15 µm across, while the lists of the mature tubarium are about five times thicker, at 75 µm. The thickening of lists starts from the proximal end. The looping meshes of the ancora umbrella are overgrown by a reticulum in the mature colonies. The ancora prongs below the lateral proximal orifices and the lists surrounding the proximal lateral orifices are extremely wide in mature specimens (
<figureCitation id="13342A6FFFF8985F8FAAD25BFD96FEE1" box="[159,241,295,320]" captionStart="FIGURE 14" captionStartId="18.[151,250,1778,1800]" captionTargetBox="[151,1435,556,1757]" captionTargetId="figure@18.[151,1435,556,1757]" captionTargetPageId="18" captionText="FIGURE 14. Gothograptus auriculatus n. sp., proximal ends of tubaria, Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. immature tubarium with the first three thecae, reverse view, ZPAL G.55/27. B. ventro-lateral side of tubarium, ZPAL G.55/28. C–E. reverse views of mature tubaria with very wide lists surrounding proximal lateral orifice and prongs; C. ZPAL G.55/23; D. lateral view, ZPAL G.55/29; F." httpUri="https://zenodo.org/record/2601647/files/figure.png" pageId="18" pageNumber="453">Fig. 14</figureCitation>
C–F).The reticular lists also become very wide in mature colonies, making the reticulum meshes very small. In the most mature specimens the reticulum lists reach similar thicknesses to those of the main lists of the tubaria (
<figureCitation id="13342A6FFFF8985F8FC1D213FC3BFE29" box="[244,348,367,392]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="18" pageNumber="453">Fig. 13D</figureCitation>
).
</paragraph>
<paragraph id="8BB036EAFFF8985F8FFAD2E8FF30FE56" blockId="18.[151,1437,151,503]" pageId="18" pageNumber="453">
Based on well-preserved specimens representing the young tubaria, the sicula length is estimated at
<quantity id="4CF79B0FFFF8985F8A0AD2E8F8FFFE0D" box="[1343,1432,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="453" unit="mm" value="1.4">1.4 mm</quantity>
, the length of the prosicula at 400 µm (
<figureCitation id="13342A6FFFF8985F8D75D2C6FFE2FE72" box="[576,645,442,467]" captionStart="FIGURE 2" captionStartId="4.[151,250,1897,1919]" captionTargetBox="[152,1435,193,1875]" captionTargetId="figure@4.[152,1435,193,1875]" captionTargetPageId="4" captionText="FIGURE 2. Immature tubaria of Gothograptus auriculatus n. sp. from the Bartoszyce IG-1 core, depth 1645.6 m, praedeubeli Biozone, Poland. A. tubarium with four pairs of thecae, ZPAL G.55/1. B. tubarium with two pairs of thecae, ZPAL G.55/2. C– E. tubarium with first pair of thecae, ZPAL G.55/3; C. ventro-lateral view, D. enlargement of distal part of virga with trace of prosicular apex, E. enlargement of distal end showing virga and fragment of virgella. Abbreviation: pros.—prosicular." httpUri="https://zenodo.org/record/2601615/files/figure.png" pageId="18" pageNumber="453">Fig. 2</figureCitation>
). The nema starts at the level of the th1
<superScript id="7C7A9BA2FFF8985F8B0ED2CBF924FE64" attach="left" box="[1083,1091,439,453]" fontSize="6" pageId="18" pageNumber="453">2</superScript>
geniculum and is connected to the obverse lateral wall of the tubarium.
</paragraph>
<caption id="DF706662FFF8985F8FA2D58EFF9DF8C5" httpUri="https://zenodo.org/record/2601647/files/figure.png" pageId="18" pageNumber="453" startId="18.[151,250,1778,1800]" targetBox="[151,1435,556,1757]" targetPageId="18">
<paragraph id="8BB036EAFFF8985F8FA2D58EFF9DF8C5" blockId="18.[151,1436,1778,1892]" pageId="18" pageNumber="453">
<emphasis id="B97BEAF8FFF8985F8FA2D58EFC46F8A9" bold="true" box="[151,289,1778,1800]" pageId="18" pageNumber="453">FIGURE 14.</emphasis>
<taxonomicName id="4C0F4D69FFF8985F8E1DD58EFF57F8A9" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[296,560,1778,1800]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="18" pageNumber="453" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFF8985F8E1DD58EFF57F8A9" box="[296,560,1778,1800]" italics="true" pageId="18" pageNumber="453">Gothograptus auriculatus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFF8985F8D0DD58EFF15F8A9" bold="true" box="[568,626,1778,1800]" pageId="18" pageNumber="453">
<taxonomicNameLabel id="A2485783FFF8985F8D0DD58EFF15F8A9" box="[568,626,1778,1800]" pageId="18" pageNumber="453" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, proximal ends of tubaria, Bartoszyce IG-1 core, depth 1645.2 m,
<taxonomicName id="4C0F4D69FFF8985F8A16D58EFD89F886" authority="Biozone" authorityName="Biozone" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="18" pageNumber="453" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFF8985F8A16D58EF8FBF8A9" box="[1315,1436,1778,1800]" italics="true" pageId="18" pageNumber="453">praedeubeli</emphasis>
Biozone
</taxonomicName>
, Poland. A. immature tubarium with the first three thecae, reverse view, ZPAL G.55/27. B. ventro-lateral side of tubarium, ZPAL G.55/28. C–E. reverse views of mature tubaria with very wide lists surrounding proximal lateral orifice and prongs; C. ZPAL G.55/23; D. lateral view, ZPAL G.55/29; F.
</paragraph>
</caption>
<paragraph id="8BB036EAFFF8985E8FF2D4EDF9C9FF75" blockId="18.[151,1436,1937,2034]" lastBlockId="19.[151,1436,151,464]" lastPageId="19" lastPageNumber="454" pageId="18" pageNumber="453">
The typical genicular structures are well developed in mature specimens. During astogeny the auricles, typical of this species, are first developed in more distal thecae (
<figureCitation id="13342A6FFFF8985F8C2DD4C8FE00F86C" box="[792,871,1972,1997]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="18" pageNumber="453">Fig. 13</figureCitation>
A–C). The longest hoods in mature specimens are about 370 µm long, and extend below the lips. As a consequence the thecal orifices are not visible. The hoods of astogenetically old tubaria may be elongated in the centre of their lower parts (
<figureCitation id="13342A6FFFF9985E8B1BD3EBF9F1FF11" box="[1070,1174,151,176]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="19" pageNumber="454">Fig. 13E</figureCitation>
). The distal thecae are shorter and their genicular structures are usually irregular and shorter horizontally (
<figureCitation id="13342A6FFFF9985E8B09D3C0F9C7FF75" box="[1084,1184,188,213]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="19" pageNumber="454">Fig. 13E</figureCitation>
).
</paragraph>
<paragraph id="8BB036EAFFF9985E8FF2D39CFE1CFEBC" blockId="19.[151,1436,151,464]" pageId="19" pageNumber="454">
In mature specimens, the nema widens distally to about 67 µm across at the most distal end of the tubarium and in the appendix (
<figureCitation id="13342A6FFFF9985E8E67D278FCFCFEBD" box="[338,411,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,766,788]" captionTargetBox="[151,1435,218,715]" captionTargetId="figure@5.[151,1435,218,715]" captionTargetPageId="5" captionText="FIGURE 3. Gothograptus auriculatus n. sp., fragments of distal parts of tubaria showing a thick nema, Bartoszyce IG-1, depth 1647.2 m, praedeubeli Biozone, Poland. A–B, distal end of tubarium with broken appendix, ZPAL G.55/4, A, fragment with beginning of appendix, B, enlargement of last thecal region showing thick nema. C, lateral view of fragment of medial part of tubarium, ZPAL G.55/5." httpUri="https://zenodo.org/record/2601617/files/figure.png" pageId="19" pageNumber="454">Figs 3</figureCitation>
,
<figureCitation id="13342A6FFFF9985E8E92D278FCBEFEBD" box="[423,473,260,284]" captionStart="FIGURE 13" captionStartId="17.[151,250,1830,1852]" captionTargetBox="[151,1436,265,1809]" captionTargetId="figure@17.[151,1436,265,1809]" captionTargetPageId="17" captionText="FIGURE 13. Mature tubaria of Gothograptus auriculatus n. sp., Bartoszyce IG-1 core, depth 1645.2 m, praedeubeli Biozone, Poland. A. lateral-ventral tubarium aspect with six pairs of thecae, with the distal end broken off, ZPAL G.55/22. B. obverse side of tubarium with proximal end, holotype, ZPAL G.55/23. C, F. reverse views of distal fragments with six pairs of thecae and appendix, paratype ZPAL G.55/24; C. whole specimen; F. enlargement of distal part showing thick lists and auricle. D. part of tubarium with proximal end, reverse view, ZPAL G.55/25. E. ventral view of distal part of tubarium with beginning of appendix, ZPAL G.55/26." httpUri="https://zenodo.org/record/2601645/files/figure.png" pageId="19" pageNumber="454">13C</figureCitation>
). A nematularium was not identified.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF9985E8FF2D25BFC44FE6E" pageId="19" pageNumber="454" type="discussion">
<paragraph id="8BB036EAFFF9985E8FF2D25BFC44FE6E" blockId="19.[151,1436,151,464]" pageId="19" pageNumber="454">
<emphasis id="B97BEAF8FFF9985E8FF2D25BFC5CFEE1" bold="true" box="[199,315,295,320]" pageId="19" pageNumber="454">Remarks.</emphasis>
<taxonomicName id="4C0F4D69FFF9985E8E76D254FF01FEE1" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[323,614,296,320]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="auriculatus" status="sp. nov.">
<emphasis id="B97BEAF8FFF9985E8E76D254FF01FEE1" box="[323,614,296,320]" italics="true" pageId="19" pageNumber="454">Gothograptus auriculatus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFF9985E8D58D25BFFCAFEE1" bold="true" box="[621,685,295,320]" pageId="19" pageNumber="454">
<taxonomicNameLabel id="A2485783FFF9985E8D58D25BFFCAFEE1" box="[621,685,295,320]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has been described from the
<taxonomicName id="4C0F4D69FFF9985E8CCFD254F982FEE1" authority="Biozone" authorityName="Biozone" box="[1018,1253,295,320]" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFF9985E8CCFD254F9E7FEE1" box="[1018,1152,296,320]" italics="true" pageId="19" pageNumber="454">praedeubeli</emphasis>
Biozone
</taxonomicName>
of
<collectingCountry id="F318767AFFF9985E8A39D25BF80FFEE1" box="[1292,1384,295,320]" name="Sweden" pageId="19" pageNumber="454">Sweden</collectingCountry>
, and published as
<taxonomicName id="4C0F4D69FFF9985E8E19D230FCEBFEC5" box="[300,396,332,356]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF9985E8E19D230FCEBFEC5" box="[300,396,332,356]" italics="true" pageId="19" pageNumber="454">G. nassa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9E4B1BFFF9985E8EA9D230FF08FEC4" author="Calner, M. &amp; Kozlowska, A. &amp; Masiak, M. &amp; Schmitz, B." box="[412,623,332,357]" pageId="19" pageNumber="454" pagination="79 - 84" refId="ref10672" refString="Calner, M., Kozlowska, A., Masiak, M. &amp; Schmitz, B. (2006) A shoreline to deep basin correlation chart for the middle Silurian coupled extinction-stable isotopic event. GFF, 128, 79 - 84. https: // doi. org / 10.1080 / 11035890601282079" type="journal article" year="2006">
Calner
<emphasis id="B97BEAF8FFF9985E8EC4D231FF4CFEC5" box="[497,555,332,356]" italics="true" pageId="19" pageNumber="454">et al.</emphasis>
2006
</bibRefCitation>
, fig. 1E). This taxon, also found in erratic boulders from
<collectingCountry id="F318767AFFF9985E8A35D230F80EFEC5" box="[1280,1385,332,356]" name="Germany" pageId="19" pageNumber="454">Germany</collectingCountry>
, has been illustrated and identified by Jaeger as
<taxonomicName id="4C0F4D69FFF9985E8D48D20CFE07FE29" box="[637,864,368,392]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF9985E8D48D20CFE07FE29" box="[637,864,368,392]" italics="true" pageId="19" pageNumber="454">Gothograptus nassa</emphasis>
</taxonomicName>
late form (
<bibRefCitation id="EF9E4B1BFFF9985E8CE8D213F9BEFE29" author="Maletz, J." box="[989,1241,367,392]" pageId="19" pageNumber="454" pagination="501 - 522" refId="ref12051" refString="Maletz, J. (2010) Retiolitid graptolites from the collection of Hermann Jaeger II: Cometograptus, Spinograptus and Plectograptus. Palaontologische Zeitschrift, 84, 501 - 522. https: // doi. org / 10.1007 / s 12542 - 010 - 0065 - x" type="journal article" year="2010">Maletz 2010, figs 1, 2</bibRefCitation>
). The specimens, stored in the Museum für Naturkunde,
<collectingRegion id="49CBF808FFF9985E8D63D2E8FFF8FE0C" box="[598,671,404,429]" country="Germany" name="Berlin" pageId="19" pageNumber="454">Berlin</collectingRegion>
, come from a fauna containing the
<typeStatus id="54B48848FFF9985E8B02D2E9F900FE0C" box="[1079,1127,405,429]" pageId="19" pageNumber="454">type</typeStatus>
material of
<taxonomicName id="4C0F4D69FFF9985E8BC1D2E8FC7AFE71" class="Graptolithina" genus="Colonograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="praedeubeli">
<emphasis id="B97BEAF8FFF9985E8BC1D2E8FC7AFE71" italics="true" pageId="19" pageNumber="454">Colonograptus praedeubeli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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