<documentid="78E33C5272982C5507C01005AA19346E"ID-DOI="10.12651/JSR.2024.13.2.185"ID-ISSN="2713-8615"ID-Zenodo-Dep="13344859"IM.bibliography_approvedBy="carolina"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722550107885"checkinUser="felipe"docAuthor="Lee, Il-Hoi Kim and Taekjun"docDate="2024"docId="ED1887CBFFE0FFC8FF7FB537B2646154"docLanguage="en"docName="JSpeciesRes.13.2.185-254.pdf"docOrigin="Journal of Species Research 13 (2)"docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article"docStyleId="4D398BFF45FF8CA9E08F546BB06AE372"docStyleName="JSpeciesRes.2012-.journal_article"docStyleVersion="2"docTitle="Asterocheres spiniventer Lee, 2024, n. sp."docType="treatment"docVersion="3"lastPageNumber="220"masterDocId="1121FFB3FFFEFFEBFFD1B25AB0306275"masterDocTitle="Copepods of the genus Asterocheres Boeck, 1859 (Siphonostomatoida: Asterocheridae) from Korean waters"masterLastPageNumber="254"masterPageNumber="185"pageNumber="215"updateTime="1724091014337"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="799D6D2D4DE811590011B02E5C2132D8">Copepods of the genus Asterocheres Boeck, 1859 (Siphonostomatoida: Asterocheridae) from Korean waters</mods:title>
<figureCitationid="FD8A2A58FFE0FFF5FDDAB534B2BE65F3"box="[523,654,1902,1926]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="32.[174,207,1907,1929]"captionStartId-1="33.[144,177,1719,1741]"captionStartId-2="34.[174,207,1755,1777]"captionTargetBox-0="[212,1349,267,1873]"captionTargetBox-1="[228,1333,270,1684]"captionTargetBox-2="[244,1380,276,1717]"captionTargetId-0="figure-13@32.[185,1446,254,1892]"captionTargetId-1="figure-106@33.[141,1417,254,1700]"captionTargetId-2="figure-85@34.[171,1447,254,1729]"captionTargetPageId-0="32"captionTargetPageId-1="33"captionTargetPageId-2="34"captionText-0="Fig. 18. Asterocheres spiniventer n. sp., female.A, habitus, dorsal; B, urosome, dorsal; C, left caudal ramus, dorsal; D, rostral area, ventral; E, antennule; F, antenna; G, oral siphon; H, mandible; I, maxillule; J, maxilla. Scale bars:A, 0.1mm; B, D, E, G, 0.05mm; C, F, H-J, 0.02mm."captionText-1="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."captionText-2="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi-0="http://doi.org/10.5281/zenodo.13344897"figureDoi-1="http://doi.org/10.5281/zenodo.13344899"figureDoi-2="http://doi.org/10.5281/zenodo.13344901"httpUri-0="https://zenodo.org/record/13344897/files/figure.png"httpUri-1="https://zenodo.org/record/13344899/files/figure.png"httpUri-2="https://zenodo.org/record/13344901/files/figure.png"pageId="30"pageNumber="215">Figs. 18-20</figureCitation>
<materialsCitationid="D5D93C80FFE0FFF4FBD3B534B2836035"collectingDate="2022-07-20"collectingDateMax="2023-04-06"collectingDateMin="2022-07-20"collectionCode="SCUBA, T, HNIBR"collectorName="W. Jung & I. - H. Kim"country="South Korea"county="Lee"elevation="20"lastPageId="31"lastPageNumber="216"latitude="33.228085"location="Honam National Institute of Biological Resources"longLatPrecision="1"longitude="126.568306"municipality="Island"pageId="30"pageNumber="215"specimenCount="22"specimenCount-female="17"specimenCount-male="5"stateProvince="Jeju"typeStatus="holotype">
<collectionCodeid="03A0AE18FFE1FFF4FF18B31CB0EB6328"box="[201,219,326,349]"name="Tavera, Department of Geology and Geophysics"pageId="31"pageNumber="216">T</collectionCode>
<collectingDateid="014BE9F5FFE1FFF4FE85B31CB1DB632B"box="[340,491,325,350]"pageId="31"pageNumber="216"value="2022-07-20">20 July 2022</collectingDate>
</date>
. Intact type specimens have been deposited in the
<locationid="606E6006FFE1FFF4FE6AB33CB14663EB"LSID="urn:lsid:plazi:treatment:ED1887CBFFE0FFC8FF7FB537B2646154:606E6006FFE1FFF4FE6AB33CB14663EB"country="South Korea"county="Lee"latitude="33.228085"longLatPrecision="1"longitude="126.568306"municipality="Island"name="Honam National Institute of Biological Resources"pageId="31"pageNumber="216"stateProvince="Jeju">Honam National Institute of Biological Resources</location>
<collectionCodeid="03A0AE18FFE1FFF4FE16B072B1E9604A"box="[455,473,552,575]"name="Tavera, Department of Geology and Geophysics"pageId="31"pageNumber="216">T</collectionCode>
<collectingDateid="014BE9F5FFE1FFF4FDC9B07DB29F6035"box="[536,687,551,576]"pageId="31"pageNumber="216"value="2023-04-06">06 April 2023</collectingDate>
) moderately broad. Body length of figured and described specimen 807 μm. Prosome ovoid, 552 μm long, representing about 68% of body length. Cephalothorax wider than long (356 × 418 μm). All prosomal somite bearing rounded lateral corners. Urosome (
) 4-segmented. Fifth pedigerous somite 114 μm wide. Genital double-somite 1.06 times longer than wide (114 × 108 μm); genital apertures positioned dorsolaterally at widest, 40% region of double-somite; postgenital part tapering posteriorly; postgenital lateral margins each bearing about 27 small spinules grouped as 8, 6-8, 7-8, and 4 from proximal to distal (
<figureCitationid="FD8A2A58FFE1FFF4FF23B616B1626611"box="[242,338,1100,1124]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19F</figureCitation>
) 1.52 times longer than wide (35 × 23 μm), armed with 6 setae, and ornamented with 5 transverse row of minute spinules on both sides; distal margin smooth, lacking any spinule.
) bearing short, unarmed coxa; basis unarmed but bearing longitudinal row of fine spinules; exopodal segment 11 × 5 μm, bearing 3 setae; first endopodal segment 49 μm long, unarmed but ornamented with fine spinules (or setules) along outer margin; short second endopodal segment tipped with 1 small seta; third endopodal segment bearing 3 unequal setae; terminal claw of 49 μm long.
) stout; proximal segment bearing row of thin, needle-like spinules at proximal region; distal segment longer than proximal segment, with few spinules and setules in distal region. Maxilliped (
<figureCitationid="FD8A2A58FFE1FFF4FC61B07DB4256035"box="[944,1045,551,576]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19G</figureCitation>
) 6-segmented; armature formula 1, 1, 2, 1, 1, and 1 + claw; second segment with spinules on outer margin; terminal segment and claw 24 and 53 μm long, respectively.
<figureCitationid="FD8A2A58FFE1FFF4FC48B0F2B43060B4"box="[921,1024,680,705]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19A</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FBE3B0F2B4A760B4"box="[1074,1175,680,705]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19B</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FAC9B0F2B54D60B4"box="[1304,1405,680,705]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19D</figureCitation>
) biramous with 3-segmented rami. Distal margin of intercoxal plate spinulose in legs 1 and 2, but smooth in legs 3 and 4. Inner distal seta on basis of leg 1 thin and naked. Leg 3 similar to leg 2, except bearing 1 spine and 5 setae on third endopodal segment (
<figureCitationid="FD8A2A58FFE1FFF4FB6CB110B5126117"box="[1213,1314,842,866]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19C</figureCitation>
). Second endopodal segment of all swimming legs bearing bicuspid outer distal corner. Inner coxal seta of leg 4 small but distinct. Armature formula for legs 1-4 as in
<figureCitationid="FD8A2A58FFE1FFF4FA8BB651B3656631"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19E</figureCitation>
) 1.85 times longer than wide (37 × 20 μm), with 3 setae and spinulose outer and inner margins; lengths of setae 55, 48, and 20 from outer to inner. Leg 6 (
<figureCitationid="FD8A2A58FFE1FFF4FAF3B636B54D66F1"box="[1314,1405,1132,1156]"captionStart="Fig"captionStartId="33.[144,177,1719,1741]"captionTargetBox="[228,1333,270,1684]"captionTargetId="figure-106@33.[141,1417,254,1700]"captionTargetPageId="33"captionText="Fig. 19. Asterocheres spiniventer n. sp., female.A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars:A-D, 0.05mm; E-G, 0.02mm."figureDoi="http://doi.org/10.5281/zenodo.13344899"httpUri="https://zenodo.org/record/13344899/files/figure.png"pageId="31"pageNumber="216">Fig. 19F</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FC6EB697B4176690"box="[959,1063,1229,1253]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="31"pageNumber="216">Fig. 20A</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FC08B774B40E6733"box="[985,1086,1326,1350]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="31"pageNumber="216">Fig. 20B</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FB6FB7B5B512647D"box="[1214,1314,1519,1544]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="31"pageNumber="216">Fig. 20C</figureCitation>
) 17-segmented, geniculate between 15th and 16th segments; 9th, 12th, 15th, 16th, and terminal segments bearing 5 setae, 4 setae, 4 setae, 2 setae + aesthetasc, and 11 setae, respectively; all other segments each with 2 setae; penultimate segment with pointed anterodistal corner; all setae naked. Antenna as in female.
<figureCitationid="FD8A2A58FFE1FFF4FC3CB4ABB460657F"box="[1005,1104,1777,1802]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="31"pageNumber="216">Fig. 20D</figureCitation>
<figureCitationid="FD8A2A58FFE1FFF4FC5FB568B3DF653F"box="[910,1007,1842,1866]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="31"pageNumber="216">Fig. 20E</figureCitation>
) segmented and armed as in female, but endopod different from that of female; second endopodal segment bearing unicuspid outer distal corner and thick setules on outer distal region; third endopodal segment distally with 1 strong, spiniform outer process bearing spinules on its outer margin and 1 smaller, curved inner process bearing truncate tip. Legs 2-4 as in female.
, female. A, leg 1; B, leg 2; C, endopod of leg 3; D, leg 4; E, exopod of leg 5; F, left side of genital double-somite, dorsal; G, maxilliped. Scale bars: A- D, 0.05 mm; E- G, 0.02 mm.
<figureCitationid="FD8A2A58FFDFFFCAFF27B5D4B16865D3"box="[246,344,1934,1958]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="33"pageNumber="218">Fig. 20F</figureCitation>
<figureCitationid="FD8A2A58FFDFFFCAFB05B514B5076513"box="[1236,1335,1870,1894]"captionStart="Fig"captionStartId="34.[174,207,1755,1777]"captionTargetBox="[244,1380,276,1717]"captionTargetId="figure-85@34.[171,1447,254,1729]"captionTargetPageId="34"captionText="Fig. 20. Asterocheres spiniventer n. sp., male. A, habitus, dorsal; B, urosome, dorsal; C, antenna; D, maxilliped; E, leg 1; F, leg 5; G, leg 6. Scale bars: A, 0.1 mm; B-G, 0.02 mm."figureDoi="http://doi.org/10.5281/zenodo.13344901"httpUri="https://zenodo.org/record/13344901/files/figure.png"pageId="33"pageNumber="218">Fig. 20G</figureCitation>
, as the new species exhibits higher similarity values with these species (more than 75% values) than with other species (
<tableCitationid="28330366FFDCFFC9FD1CB528B08E65DE"captionStart="Table 2"captionStartId="2.[92,145,474,499]"captionText="Table 2. A character matrix of Asterocheres species (“-” indicates missing data)."pageId="34"pageNumber="219">Table 2</tableCitation>
<taxonomicNameid="A2B14D5EFFDCFFC9FE64B5E9B43C651F"authority="Crescenti, Baviera & Zaccone, 2010"authorityName="Crescenti, Baviera & Zaccone"authorityYear="2010"box="[437,1036,1874,1995]"class="Copepoda"family="Asterocheridae"genus="Asterocheres"kingdom="Animalia"order="Siphonostomatoida"pageId="34"pageNumber="219"phylum="Arthropoda"rank="species"species="faroensis">
<bibRefCitationid="01204B2CFFDCFFC9FD90B5E9B43C651F"author="Crescenti, N. & C. Baviera & D. Zaccone"box="[577,1036,1874,1995]"pageId="34"pageNumber="219"pagination="299 - 311"refId="ref37122"refString="Crescenti, N., C. Baviera and D. Zaccone. 2010. Asterocheres faroensis, a new species (Copepoda, Siphonostomatoida, Asterocheridae) associated with Hymeniacidon perlevis (Montagu, 1812) (Porifera) from Lake Faro, Mediterranean Sea. Crustaceana 83 (3): 299 - 311. https: // doi. org / 10.1 163 / 001121609 X 12596543952414"type="journal article"year="2010">Crescenti, Baviera & Zaccone, 2010</bibRefCitation>
<bibRefCitationid="01204B2CFFDCFFC9FB2BB508B5A7651F"author="Hansen, H. J."box="[1274,1431,1874,1898]"pageId="34"pageNumber="219"pagination="1 - 92"refId="ref37363"refString="Hansen, H. J. 1923. Crustacea Copepoda II. Copepoda Parasita and Hemiparasita. The Danish Ingolf-Expedition 3: 1 - 92."type="journal article"year="1923">Hansen, 1923</bibRefCitation>
)
</taxonomicName>
. However, they are clearly distinguished from the new species by their diagnostic features exhibited in the mandibular palp and oral siphon, since the first three species have a 2-segmented mandibular palp (vs. 1-segmented in
<taxonomicNameid="A2B14D5EFFDDFFC8FD04B37FB0C16328"authorityName="Crescenti, Baviera & Zaccone"authorityYear="2010"class="Copepoda"family="Asterocheridae"genus="Asterocheres"kingdom="Animalia"order="Siphonostomatoida"pageId="35"pageNumber="220"phylum="Arthropoda"rank="species"species="faroensis">
, have a longer oral siphon that approaches over the insertion of the maxilliped in the former species (
<bibRefCitationid="01204B2CFFDDFFC8FEB5B3DCB27F63EB"author="Crescenti, N. & C. Baviera & D. Zaccone"box="[356,591,390,414]"pageId="35"pageNumber="220"pagination="299 - 311"refId="ref37122"refString="Crescenti, N., C. Baviera and D. Zaccone. 2010. Asterocheres faroensis, a new species (Copepoda, Siphonostomatoida, Asterocheridae) associated with Hymeniacidon perlevis (Montagu, 1812) (Porifera) from Lake Faro, Mediterranean Sea. Crustaceana 83 (3): 299 - 311. https: // doi. org / 10.1 163 / 001121609 X 12596543952414"type="journal article"year="2010">Crescenti et al., 2010</bibRefCitation>
<bibRefCitationid="01204B2CFFDDFFC8FEA3B3FCB23B63CB"author="Hansen, H. J."box="[370,523,422,446]"pageId="35"pageNumber="220"pagination="1 - 92"refId="ref37363"refString="Hansen, H. J. 1923. Crustacea Copepoda II. Copepoda Parasita and Hemiparasita. The Danish Ingolf-Expedition 3: 1 - 92."type="journal article"year="1923">Hansen, 1923</bibRefCitation>
) (vs. the oral siphon approaches to the insertion of the maxilliped in
, the caudal ramus is 1.5 times longer than wide, two distal setae on the mandibular palp are extremely unequal in length (longer seta six times longer than shorter one), and the genital double-somite of the female is ornamented with about 25 spinules arranged along the entire postgenital lateral margin. Most of all, the nature of the sexually dimorphic morphological transformation of the endopod of male leg 1 is unique to the new species.