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245 lines
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<document id="8B60D7BC4F4D04E9B7166ACCF6090E72" ID-DOI="10.11646/zootaxa.5152.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6630873" ID-ZooBank="92E9DD85-6CC6-4602-BD7C-C51F49CEEF47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654852147308" checkinUser="plazi" docAuthor="Márquez, Juan & Asiain, Julieta" docDate="2022" docId="038E878977187F3B81AFA202FA82FA79" docLanguage="en" docName="zootaxa.5152.1.1.pdf" docOrigin="Zootaxa 5152 (1)" docSource="http://zoobank.org/92e9dd85-6cc6-4602-bd7c-c51f49ceef47" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Belonuchus mixtecus Márquez & Asiain 2022, sp. nov." docType="treatment" docUuid="341F9562-23CC-447B-82D0-8848B80967CF" docUuidSource="ZooBank" docVersion="3" lastPageNumber="57" masterDocId="FFB7FFF1772F7F038138A63CFF9FFFE4" masterDocTitle="Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)" masterLastPageNumber="129" masterPageNumber="1" pageNumber="56" updateTime="1699335993931" updateUser="plazi">
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<mods:titleInfo id="40D76D8B4FEFF4E0DB15BF4184595256">
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<mods:title id="962FB1B52E02C3F008BEE5C22AA93CF5">Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)</mods:title>
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<mods:name id="27BCFBB28E7786BB343B63E8E9B719C8" type="personal">
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<mods:namePart id="04FBA5CB500CFBF60E1E1F948104581B">Márquez, Juan</mods:namePart>
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<mods:nameIdentifier id="FE80B5A14D8F6CBC61DA2C7B4A203685" type="ORCID">0000-0001-5423-3545</mods:nameIdentifier>
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<mods:nameIdentifier id="466ACFFC323F23169635109EFCE4789F" type="email">marquezorum@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="752232CB4CB78506BE3635893C05F006">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7791E6D1A0A79761A8112D2DAF68273A">Asiain, Julieta</mods:namePart>
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<mods:nameIdentifier id="E0629032F89357DE2714F3D6C5B2023A" type="ORCID">0000-0002-7958-4779</mods:nameIdentifier>
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<mods:nameIdentifier id="6538CB9F76B0C60ED6F2070188ABF8F9" type="email">asiainae@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="03DACCB42553C4C2E82E17C5F2695D07">text</mods:typeOfResource>
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<mods:title id="07D66A36C983747E2840A21440B250FA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F967AEBB289A38539D23B8C7A447A871">
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<mods:date id="07DCF1C792BE6BF9B77D0C43D1C7FD00">2022</mods:date>
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<mods:number id="90F968FBD9C5F3CDE1803C8DCA3406A8">2022-06-10</mods:number>
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<mods:number id="BA48AB7F811E329B3AA32A16B675D4A5">5152</mods:number>
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<mods:number id="D618691E0101F991FABC53E8B08188A3">1</mods:number>
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<mods:start id="37E2DB995F91D273AE9F098CA0719EDF">1</mods:start>
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<mods:url id="F33FE494C1E581C0BCC7A7B6439FC118">http://zoobank.org/92e9dd85-6cc6-4602-bd7c-c51f49ceef47</mods:url>
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<mods:classification id="98852EE5FDBF46F106A880E624B31A9B">journal article</mods:classification>
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<mods:identifier id="6AE8D03A20201384750B6968E2D546C2" type="DOI">10.11646/zootaxa.5152.1.1</mods:identifier>
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<mods:identifier id="E4F235479893D0D50DE13E77E91C5856" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="158D39D31E643ED193626CC827D351FA" type="Zenodo-Dep">6630873</mods:identifier>
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<treatment id="038E878977187F3B81AFA202FA82FA79" ID-DOI="http://doi.org/10.5281/zenodo.6630861" ID-Zenodo-Dep="6630861" LSID="urn:lsid:zoobank.org:act:341F9562-23CC-447B-82D0-8848B80967CF" httpUri="http://treatment.plazi.org/id/038E878977187F3B81AFA202FA82FA79" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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<subSubSection id="C33D651477187F3481AFA202FBBAFB99" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph id="8B98369F77187F3481AFA202FD9FFBBC" blockId="55.[151,1061,1086,1185]" box="[151,512,1086,1113]" pageId="55" pageNumber="56">
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<heading id="D0D081F377187F3481AFA202FD9FFBBC" bold="true" box="[151,512,1086,1113]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="4">
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<emphasis id="B953EA8D77187F3481AFA202FD9FFBBC" bold="true" box="[151,512,1086,1113]" pageId="55" pageNumber="56">
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<taxonomicName id="4C274D1C77187F3481AFA202FE03FBBC" ID-CoL="9GKJ3" authority="Márquez & Asiain, 2022" authorityName="Márquez & Asiain" authorityYear="2022" box="[151,412,1086,1113]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="mixtecus" status="sp. nov.">
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<emphasis id="B953EA8D77187F3481AFA202FE03FBBC" bold="true" box="[151,412,1086,1113]" italics="true" pageId="55" pageNumber="56">Belonuchus mixtecus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26057F677187F34809AA202FD9FFBBC" box="[418,512,1086,1112]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B98369F77187F3481AFA25EFBBAFB99" blockId="55.[151,1061,1086,1185]" box="[151,1061,1122,1149]" pageId="55" pageNumber="56">http://zoobank.org/NomenclaturalActs/341f9562-23cc-447b-82d0-8848b80967cf</paragraph>
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</subSubSection>
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<subSubSection id="C33D651477187F3B81AFA2BAFDC1FB21" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="description">
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<paragraph id="8B98369F77187F3481AFA2BAFD99FB44" blockId="55.[151,1061,1086,1185]" box="[151,518,1158,1185]" pageId="55" pageNumber="56">
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<figureCitation id="131C2A1A77187F3481AFA2BAFF6DFB45" box="[151,242,1158,1185]" captionStart="FIGURE 5" captionStartId="99.[151,250,1721,1745]" captionTargetBox="[175,1411,194,1684]" captionTargetId="figure-19@99.[163,1424,181,1697]" captionTargetPageId="99" captionText="FIGURE 5. Dorsal view of males (except figure g) of: a) B. mixtecus (holotype), b) B. pollens (syntype, NHM©), c) B. pollens, d) B. rufipennis, e) B. basiventris (syntype, NHM©), f) B. basiventris, g) B. connexus (female holotype of B. xanthopus, synonym of B. connexus, shared by A. Kovalev©), h) B. connexus. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630893" httpUri="https://zenodo.org/record/6630893/files/figure.png" pageId="55" pageNumber="56">Figs. 5a</figureCitation>
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,
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<figureCitation id="131C2A1A77187F3481C6A2BAFEB6FB44" box="[254,297,1158,1184]" captionStart="FIGURE 13" captionStartId="107.[151,250,1930,1954]" captionTargetBox="[165,1424,192,1892]" captionTargetId="figure-19@107.[151,1436,181,1907]" captionTargetPageId="107" captionText="FIGURE 13. Ventral view of the left posterior leg of males of: a) B. trochanterinus, b) B. tenuistylus, c) B. zunilensis, d) B. rufipennis, e) B. mixtecus, f) B. pollens, g) B. ephippiatus, h) B. aenigmaticus, i) B. iridescens, j) B. viridipennis. Tarsomeres of posterior leg of males of: k) B. colon, l) B. ephippiatus. Yellow ovals: median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6630917" httpUri="https://zenodo.org/record/6630917/files/figure.png" pageId="55" pageNumber="56">13e</figureCitation>
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, 17o,
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<quantity id="4CDF9B7A77187F348054A2BAFE07FB44" box="[364,408,1158,1184]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="55" pageNumber="56" unit="g" value="19.0">19g</quantity>
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, 23d, 27b
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</paragraph>
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<paragraph id="8B98369F77187F3481AFA2F2FE21FAB0" blockId="55.[151,1437,1230,2013]" pageId="55" pageNumber="56">
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Total body length
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<quantity id="4CDF9B7A77187F34805EA2F2FE50FB0D" box="[358,463,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="55" pageNumber="56" unit="mm" value="12.5">12.5 mm</quantity>
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. Black on head, antennae, mouthparts, pronotum, scutellum, prosternum, anterior half of mesoventrite, from fifth visible abdominal segment (except 1/4 to 1/5 anterior) to genital segment and abdominal styli. Reddish on elytra, posterior half of mesoventrite, metaventrite, legs, visible abdominal segments 1–4 and anterior 1/4 to 1/5 of fifth.
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</paragraph>
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<paragraph id="8B98369F77187F3481FFA362FB9FF9B5" blockId="55.[151,1437,1230,2013]" pageId="55" pageNumber="56">
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<emphasis id="B953EA8D77187F3481FFA362FE98FA9C" bold="true" box="[199,263,1374,1400]" pageId="55" pageNumber="56">Head</emphasis>
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: transverse, ratio length/width 0.65, postero-dorsal area and posterior corners convex. Dorsal surface with sulcate longitudinal midline visible in anterior half of head; front foveate between antennal insertions. Eyes less than 0.5 times the cephalic lateral length, not protruding laterally. Antennomere 2 almost as long as 3, 4–6 elongate, 7 as long as wide, 8–10 transverse. Mandibles very long, 1.75 times longer than head; each with two very close teeth, basal tooth bigger than middle tooth; mandibular channel moderately developed, external margin slightly carinate, internal margin carinate, extending beyond level of middle tooth. Apical palpomere of maxillary and labial palpi almost as long as preapical palpomere. Head 1.23 times wider than pronotum.
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</paragraph>
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<paragraph id="8B98369F77187F3B81FFA066FDF2FF31" blockId="55.[151,1437,1230,2013]" lastBlockId="56.[151,1437,151,1437]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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<emphasis id="B953EA8D77187F3481FFA066FEBFF990" bold="true" box="[199,288,1626,1652]" pageId="55" pageNumber="56">Thorax</emphasis>
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: left dorsal row with seven punctures and right dorsal row with six punctures; pronotum 1.12 times longer than wide and 1.13 times wider at anterior corners than at posterior corners. Scutellum with punctures slightly denser than on elytra, where it is moderately dense. Prosternum slightly elevated transversally before anterior margin. Intercoxal process of mesoventrite shield-shaped; transverse discal ridge well developed, broadly Ushaped, not meeting margin of intercoxal process laterally. Profemur slightly widened toward apex (less noticeable than in
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<taxonomicName id="4C274D1C77187F3481D4A133FECAF8CC" authorityName="Sharp" authorityYear="1885" box="[236,341,1806,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="bidens">
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<emphasis id="B953EA8D77187F3481D4A133FECAF8CC" box="[236,341,1806,1832]" italics="true" pageId="55" pageNumber="56">B. bidens</emphasis>
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</taxonomicName>
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); external margin with row of short, dark and dense spines that initiate near base and end at apex where margin is narrow; internal margin with row of spines even shorter (difficult to see), initiating further apicad than external row and reaching apex, where four to six spines are longer than previous spines; with a few spines at base of femur between both rows. Metafemur somewhat widened and flattened in basal half of external surface (
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<figureCitation id="131C2A1A77187F348457A146FF5DF85C" captionStart="FIGURE 13" captionStartId="107.[151,250,1930,1954]" captionTargetBox="[165,1424,192,1892]" captionTargetId="figure-19@107.[151,1436,181,1907]" captionTargetPageId="107" captionText="FIGURE 13. Ventral view of the left posterior leg of males of: a) B. trochanterinus, b) B. tenuistylus, c) B. zunilensis, d) B. rufipennis, e) B. mixtecus, f) B. pollens, g) B. ephippiatus, h) B. aenigmaticus, i) B. iridescens, j) B. viridipennis. Tarsomeres of posterior leg of males of: k) B. colon, l) B. ephippiatus. Yellow ovals: median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6630917" httpUri="https://zenodo.org/record/6630917/files/figure.png" pageId="55" pageNumber="56">Fig. 13e</figureCitation>
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); with two rows of spines not clearly aligned at base, where there are some additional spines, spines are best aligned more distally; spines extending close to 3/4 of length of femur; spines of external side stronger and longer than spines of internal side, and also longer than spines in
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<taxonomicName id="4C274D1C77177F3B820EA6ABFC56FF55" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[822,969,151,177]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B953EA8D77177F3B820EA6ABFC56FF55" box="[822,969,151,177]" italics="true" pageId="56" pageNumber="57">B. rufipennis</emphasis>
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</taxonomicName>
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. Metatibiae weakly curved at midlength. Tarsomeres moderately flattened dorsally.
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</paragraph>
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<paragraph id="8B98369F77177F3B81FFA6E2FE4DFE11" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
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<emphasis id="B953EA8D77177F3B81FFA6E2FEA3FF1C" bold="true" box="[199,316,222,248]" pageId="56" pageNumber="57">Abdomen</emphasis>
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: first three visible abdominal tergites with posterior basal transverse carina well developed; adjacent area a little depressed and with sparse, wide-superficial punctures combined with fine punctures moderately dense; visible tergite 4 with punctures as on tergites 1–3; visible tergites 5–6 only with fine punctures. Visible sternites 1–4 with punctures as on tergites 1–3, but wide-superficial punctures slightly denser; sternites 5–6 only with fine punctures. Male pregenital sternite with deep, narrow U-shaped emargination at posterior margin, as deep as nearly 1/5 of length of sternite (Fig. 17–o). Male genital sternite short (1.94 times longer than wide), moderately asymmetrical, anterior portion occupying 39% and posterior portion 61% of its length, apical emargination deep (
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<figureCitation id="131C2A1A77177F3B8457A78BFF5AFE11" captionStart="FIGURE 19" captionStartId="113.[151,250,1823,1847]" captionTargetBox="[165,1426,181,1793]" captionTargetId="figure-19@113.[161,1426,181,1800]" captionTargetPageId="113" captionText="FIGURE 19. Male genital sternite of: a) Belonuchus tepoztecus, b) B. trochanterinus, c) B. zunilensis, d) B. bidens, e) B. erichsoni, f) B. jalappensis, g) B. mixtecus, h) B. pollens, i) B. rufipennis, j) B. basiventris, k) B. connexus, l) B. longitarsus, m) B. rufiventris, n) B. xanthomelas, o) B. ephippiatus, p) B. moroni. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630927" httpUri="https://zenodo.org/record/6630927/files/figure.png" pageId="56" pageNumber="57">
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Fig.
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<quantity id="4CDF9B7A77177F3B81AFA7E7FF5AFE11" box="[151,197,475,501]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="56" pageNumber="57" unit="g" value="19.0">19g</quantity>
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</figureCitation>
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). Abdominal styli large.
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</paragraph>
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<paragraph id="8B98369F77177F3B81FFA7C2FF4DFD62" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
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<emphasis id="B953EA8D77177F3B81FFA7C2FEA5FDFC" bold="true" box="[199,314,510,536]" pageId="56" pageNumber="57">Aedeagus</emphasis>
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: length
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<quantity id="4CDF9B7A77177F3B80A1A7C3FE6BFDFD" box="[409,500,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="56" pageNumber="57" unit="mm" value="1.9">1.9 mm</quantity>
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; oval shape; apex of median lobe narrow and rounded, median lobe parallel sided and narrow, compared to rest of aedeagus which is wider; apical half of aedeagus becoming narrow to base of median lobe, basal half of aedeagus with sides almost homogeneous in width; internal sac visible in anterior half (
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<figureCitation id="131C2A1A77177F3B8457A47BFF59FD62" captionStart="FIGURE 23" captionStartId="117.[151,250,1878,1902]" captionTargetBox="[158,1426,187,1844]" captionTargetId="figure-19@117.[151,1436,181,1854]" captionTargetPageId="117" captionText="FIGURE 23. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus bidens, b) B. erichsoni, c) B. jalappensis, d) B. mixtecus, e) B. pollens, f) B. rufipennis, g) B. basiventris, h) B. connexus (at center, detail of the apex of the median lobe), i) B. longitarsus (at center, detail of the apex of the median lobe). Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630939" httpUri="https://zenodo.org/record/6630939/files/figure.png" pageId="56" pageNumber="57">Fig. 23d</figureCitation>
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).
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</paragraph>
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<paragraph id="8B98369F77177F3B81FFA4B2FBB1FC2D" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
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<emphasis id="B953EA8D77177F3B81FFA4B2FED8FD4C" bold="true" box="[199,327,654,680]" pageId="56" pageNumber="57">Variability</emphasis>
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. Total body length in males
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<quantity id="4CDF9B7A77177F3B83ACA4B3FD66FD4D" box="[660,761,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="56" pageNumber="57" unit="mm" value="11.9">11.9 mm</quantity>
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(range
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<quantity id="4CDF9B7A77177F3B826AA4B3FC63FD4D" box="[850,1020,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.27" metricValueMin="1.0699999999999998" pageId="56" pageNumber="57" unit="mm" value="11.7" valueMax="12.7" valueMin="10.7">10.7–12.7 mm</quantity>
|
|||
|
), in females 13.0 (range 12.6–13.5); ratio length/width of head in males 0.75 (range 0.67–0.86), in females 0.87 (range 0.86–0.88); ratio mandibular length/cephalic length in males 1.35 (range 0.96–1.67), in females 1.0 (without variation); ratio cephalic width/pronotal width in males 1.18 (range 1.08–1.26), in females 1.05 (range 1.03–1.06); ratio length/width of pronotum in males 1.12 (range 1.10–1.16), in females 1.2 (range 1.18–1.22); ratio anterior width/posterior width of pronotum in males 1.12 (range 1.07–1.15), in females 1.08 (range 1.07–1.10). Dorsal rows of punctures on pronotum vary from 7:7, 6:6 to 6:7; anterior half of prosternum may be reddish-brown; in small males, spines of posterior femora may be better aligned and, those at its base smaller than they are in large specimens; fifth visible abdominal segment may be black in posterior half and reddish in anterior half, or reddish until anterior 1/4.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B98369F77177F3B81FFA5EEFDC1FB21" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
|
|||
|
<emphasis id="B953EA8D77177F3B81FFA5EEFE55FC09" bold="true" box="[199,458,978,1005]" pageId="56" pageNumber="57">Taxonomic comments</emphasis>
|
|||
|
.
|
|||
|
<taxonomicName id="4C274D1C77177F3B80E0A5EFFD5CFC08" authorityName="Márquez & Asiain" authorityYear="2022" box="[472,707,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="mixtecus">
|
|||
|
<emphasis id="B953EA8D77177F3B80E0A5EFFD5CFC08" box="[472,707,979,1005]" italics="true" pageId="56" pageNumber="57">Belonuchus mixtecus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
has a body coloration similar to
|
|||
|
<taxonomicName id="4C274D1C77177F3B8509A5EFFB3FFC09" authorityName="Sharp" authorityYear="1885" box="[1073,1184,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pollens">
|
|||
|
<emphasis id="B953EA8D77177F3B8509A5EFFB3FFC09" box="[1073,1184,979,1005]" italics="true" pageId="56" pageNumber="57">B. pollens</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and
|
|||
|
<taxonomicName id="4C274D1C77177F3B85EEA5EFFAF8FC09" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1238,1383,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rufipennis">
|
|||
|
<emphasis id="B953EA8D77177F3B85EEA5EFFAF8FC09" box="[1238,1383,979,1005]" italics="true" pageId="56" pageNumber="57">B. rufipennis</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, and it is distinguished from both species by the dorsal rows of pronotum formed by six or seven punctures, the more robust body, the male pregenital sternite deeply emarginate at posterior margin and aedeagus large and oval. While in
|
|||
|
<taxonomicName id="4C274D1C77177F3B818EA203FEB6FBBD" authorityName="Sharp" authorityYear="1885" box="[182,297,1087,1113]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pollens">
|
|||
|
<emphasis id="B953EA8D77177F3B818EA203FEB6FBBD" box="[182,297,1087,1113]" italics="true" pageId="56" pageNumber="57">B. pollens</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and
|
|||
|
<taxonomicName id="4C274D1C77177F3B805BA203FE68FBBD" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[355,503,1087,1113]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rufipennis">
|
|||
|
<emphasis id="B953EA8D77177F3B805BA203FE68FBBD" box="[355,503,1087,1113]" italics="true" pageId="56" pageNumber="57">B. rufipennis</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
there are five punctures in each dorsal row of pronotum, the body is less robust, male pregenital sternite is slightly to moderately emarginate at posterior margin, and their aedeagus is smaller and elongate.
|
|||
|
<taxonomicName id="4C274D1C77177F3B803CA2BBFE6FFB44" authorityName="Márquez & Asiain" authorityYear="2022" box="[260,496,1159,1185]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="mixtecus">
|
|||
|
<emphasis id="B953EA8D77177F3B803CA2BBFE6FFB44" box="[260,496,1159,1185]" italics="true" pageId="56" pageNumber="57">Belonuchus mixtecus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
can also be confused with
|
|||
|
<taxonomicName id="4C274D1C77177F3B8227A2BBFC39FB45" authorityName="Bernhauer" authorityYear="1917" box="[799,934,1159,1185]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="erichsoni">
|
|||
|
<emphasis id="B953EA8D77177F3B8227A2BBFC39FB45" box="[799,934,1159,1185]" italics="true" pageId="56" pageNumber="57">B. erichsoni</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, and their differences have been indicated in the taxonomic comments of this species.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C33D651477177F3B81FFA2F3FD8EFAE9" pageId="56" pageNumber="57" type="etymology">
|
|||
|
<paragraph id="8B98369F77177F3B81FFA2F3FD8EFAE9" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
|
|||
|
<emphasis id="B953EA8D77177F3B81FFA2F3FED7FB0D" bold="true" box="[199,328,1231,1257]" pageId="56" pageNumber="57">Etymology</emphasis>
|
|||
|
. The name of this species refers to the geographical place where it was collected, which is called the “Mixteca Alta” region of
|
|||
|
<collectingRegion id="49E3F87D77177F3B808DA2CFFD92FAE9" box="[437,525,1267,1293]" country="Mexico" name="Oaxaca" pageId="56" pageNumber="57">Oaxaca</collectingRegion>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C33D651477177F3B81FFA32BFA82FA79" pageId="56" pageNumber="57" type="materials_examined">
|
|||
|
<paragraph id="8B98369F77177F3B81FFA32BFA02FAB1" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
|
|||
|
<emphasis id="B953EA8D77177F3B81FFA32BFEE3FAD4" bold="true" box="[199,380,1302,1329]" pageId="56" pageNumber="57">Type material (</emphasis>
|
|||
|
<materialsCitation id="3B4F3CC277177F3B8044A32AFA02FAB1" collectingDate="2001-08-06" collectionCode="CC-UAEH" collectorName="S. Bautista & J. Asiain & J. Marquez" country="Mexico" elevation="1788" latitude="17.151943" location="San Mateo Penasco" longLatPrecision="21" longitude="-97.06056" pageId="56" pageNumber="57" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="holotype">
|
|||
|
<emphasis id="B953EA8D77177F3B8044A32AFC89FAD5" bold="true" box="[380,790,1302,1329]" pageId="56" pageNumber="57">
|
|||
|
<specimenCount id="9D21FD1677177F3B8044A32AFE68FAD5" box="[380,503,1302,1329]" pageId="56" pageNumber="57" type="male">five males</specimenCount>
|
|||
|
,
|
|||
|
<specimenCount id="9D21FD1677177F3B833BA32BFD09FAD4" box="[515,662,1302,1329]" pageId="56" pageNumber="57" type="female">two females</specimenCount>
|
|||
|
):
|
|||
|
<typeStatus id="549C883D77177F3B8391A32BFC89FAD5" box="[681,790,1303,1329]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
|
|||
|
</emphasis>
|
|||
|
(male,
|
|||
|
<collectionCode id="ED36AE5A77177F3B8253A32BFC71FAD5" box="[875,1006,1303,1329]" pageId="56" pageNumber="57">CC-UAEH</collectionCode>
|
|||
|
): “
|
|||
|
<collectingCountry id="F330760F77177F3B8531A32BFBFAFAD5" box="[1033,1125,1303,1329]" name="Mexico" pageId="56" pageNumber="57">México</collectingCountry>
|
|||
|
:
|
|||
|
<collectingRegion id="49E3F87D77177F3B8548A32BFB58FAD5" box="[1136,1223,1303,1329]" country="Mexico" name="Oaxaca" pageId="56" pageNumber="57">Oaxaca</collectingRegion>
|
|||
|
,
|
|||
|
<location id="8EF8604477177F3B85EAA32BFF2BFAB1" LSID="urn:lsid:plazi:treatment:038E878977187F3B81AFA202FA82FA79:8EF8604477177F3B85EAA32BFF2BFAB1" country="Mexico" latitude="17.151943" longLatPrecision="21" longitude="-97.06056" name="San Mateo Penasco" pageId="56" pageNumber="57" stateProvince="Oaxaca">San Mateo Peñasco</location>
|
|||
|
, en jardín,
|
|||
|
<quantity id="4CDF9B7A77177F3B8017A307FE19FAB1" box="[303,390,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.788" pageId="56" pageNumber="57" unit="m" value="1788.0">
|
|||
|
<elevation id="000AD1AC77177F3B8017A307FE19FAB1" box="[303,390,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.788" pageId="56" pageNumber="57" unit="m" value="1788.0">1788 m</elevation>
|
|||
|
</quantity>
|
|||
|
,
|
|||
|
<geoCoordinate id="EE13505877177F3B80AAA307FD90FAB1" box="[402,527,1339,1365]" degrees="17" direction="north" minutes="09" orientation="latitude" pageId="56" pageNumber="57" precision="15" seconds="7" value="17.151943">N17°09’7”</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate id="EE13505877177F3B8322A307FD01FAB1" box="[538,670,1339,1365]" degrees="97" direction="west" minutes="3" orientation="longitude" pageId="56" pageNumber="57" precision="15" seconds="38" value="-97.06056">W97°3’38”</geoCoordinate>
|
|||
|
, en nopal podrido,
|
|||
|
<date id="FF99105F77177F3B824CA307FB9FFAB1" box="[884,1024,1339,1365]" pageId="56" pageNumber="57" value="2001-08-06">
|
|||
|
<collectingDate id="EFDDE9B777177F3B824CA307FB9FFAB1" box="[884,1024,1339,1365]" pageId="56" pageNumber="57" value="2001-08-06">6-VIII-2001</collectingDate>
|
|||
|
</date>
|
|||
|
,
|
|||
|
<collectorName id="26D2534977177F3B8532A307FB18FAB1" box="[1034,1159,1339,1365]" pageId="56" pageNumber="57">S. Bautista</collectorName>
|
|||
|
,
|
|||
|
<collectorName id="26D2534977177F3B85A9A307FB6EFAB1" box="[1169,1265,1339,1365]" pageId="56" pageNumber="57">J. Asiain</collectorName>
|
|||
|
y
|
|||
|
<collectorName id="26D2534977177F3B8435A307FA10FAB1" box="[1293,1423,1339,1365]" pageId="56" pageNumber="57">J. Márquez</collectorName>
|
|||
|
”.
|
|||
|
</materialsCitation>
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B98369F77177F3B81AFA363FA82FA79" blockId="56.[151,1437,151,1437]" pageId="56" pageNumber="57">
|
|||
|
<materialsCitation id="3B4F3CC277177F3B81AFA363FC9BFA9C" box="[151,772,1374,1401]" collectingDate="2001-08-06" collectionCode="MAAS" collectorName="S. Bautista & J. Asiain & J. Marquez" country="Mexico" elevation="1788" latitude="17.151943" location="San Mateo Penasco" longLatPrecision="21" longitude="-97.06056" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Paratypes" typeStatus="paratype">
|
|||
|
<emphasis id="B953EA8D77177F3B81AFA363FE90FA9D" bold="true" box="[151,271,1375,1401]" pageId="56" pageNumber="57">
|
|||
|
<typeStatus id="549C883D77177F3B81AFA363FE90FA9D" box="[151,271,1375,1401]" pageId="56" pageNumber="57" type="paratype">Paratypes</typeStatus>
|
|||
|
</emphasis>
|
|||
|
: same data as holotype (5,
|
|||
|
<collectionCode id="ED36AE5A77177F3B8379A363FD08FA9D" box="[577,663,1375,1401]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="56" pageNumber="57" type="Herbarium">MAAS</collectionCode>
|
|||
|
).
|
|||
|
<collectingCountry id="F330760F77177F3B8391A362FC9BFA9C" box="[681,772,1374,1400]" name="Mexico" pageId="56" pageNumber="57">
|
|||
|
<emphasis id="B953EA8D77177F3B8391A362FC9BFA9C" bold="true" box="[681,772,1374,1400]" pageId="56" pageNumber="57">Mexico</emphasis>
|
|||
|
</collectingCountry>
|
|||
|
</materialsCitation>
|
|||
|
<emphasis id="B953EA8D77177F3B823CA362FC96FA9C" bold="true" box="[772,777,1374,1400]" pageId="56" pageNumber="57">:</emphasis>
|
|||
|
<materialsCitation id="3B4F3CC277177F3B8228A362FA86FA79" collectingDate="1999-11-16" collectionCode="MAAS" collectorName="J. Marquez" country="Mexico" elevation="1436" location="San Juan Chicomesuchitl" municipality="Rio Grande" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
<emphasis id="B953EA8D77177F3B8228A362FCEBFA9C" bold="true" box="[784,884,1374,1400]" pageId="56" pageNumber="57">
|
|||
|
<collectingRegion id="49E3F87D77177F3B8228A362FCF0FA9C" box="[784,879,1374,1400]" country="Mexico" name="Oaxaca" pageId="56" pageNumber="57">Oaxaca</collectingRegion>
|
|||
|
:
|
|||
|
</emphasis>
|
|||
|
“
|
|||
|
<location id="8EF8604477177F3B82BDA363FB39FA9D" LSID="urn:lsid:plazi:treatment:038E878977187F3B81AFA202FA82FA79:8EF8604477177F3B82BDA363FB39FA9D" box="[901,1190,1375,1401]" country="Mexico" municipality="Rio Grande" name="San Juan Chicomesuchitl" pageId="56" pageNumber="57" stateProvince="Oaxaca">San Juan Chicomesuchitl</location>
|
|||
|
,
|
|||
|
<collectingMunicipality id="6BFCACE577177F3B8589A363FAAAFA9D" box="[1201,1333,1375,1401]" pageId="56" pageNumber="57">Rio Grande</collectingMunicipality>
|
|||
|
,
|
|||
|
<quantity id="4CDF9B7A77177F3B8478A363FA08FA9D" box="[1344,1431,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="56" pageNumber="57" unit="m" value="1436.0">
|
|||
|
<elevation id="000AD1AC77177F3B8478A363FA08FA9D" box="[1344,1431,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="56" pageNumber="57" unit="m" value="1436.0">1436 m</elevation>
|
|||
|
</quantity>
|
|||
|
, N17°18’729”, W96°30’600”, en nopal y mandarina podrida,
|
|||
|
<date id="FF99105F77177F3B827CA3BFFC55FA79" box="[836,970,1411,1437]" pageId="56" pageNumber="57" value="1999-11-16">
|
|||
|
<collectingDate id="EFDDE9B777177F3B827CA3BFFC55FA79" box="[836,970,1411,1437]" pageId="56" pageNumber="57" value="1999-11-16">16-XI-1999</collectingDate>
|
|||
|
</date>
|
|||
|
,
|
|||
|
<collectorName id="26D2534977177F3B82EEA3BFFBCCFA79" box="[982,1107,1411,1437]" pageId="56" pageNumber="57">J. Márquez</collectorName>
|
|||
|
col.” (1,
|
|||
|
<collectionCode id="ED36AE5A77177F3B8583A3BFFA8EFA79" box="[1211,1297,1411,1437]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="56" pageNumber="57" type="Herbarium">MAAS</collectionCode>
|
|||
|
)
|
|||
|
</materialsCitation>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|