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<document id="7E6F0975C4C14E06BC11E76214A62975" ID-DOI="http://dx.doi.org/10.3897/zse.95.32793" ID-GBIF-Dataset="7887c2f9-a00f-484e-93c2-595204b8e28d" ID-Pensoft-Pub="1860-0743-1-133" ID-ZooBank="3447C0590FE0482F81D09F91BC1ED7EC" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1860-0743-1-133" ModsDocOrigin="Zoosystematics and Evolution 1" ModsDocTitle="Tadpoles of three western African frog genera: Astylosternus Werner, 1898, Nyctibates Boulenger, 1904, and Scotobleps Boulenger, 1900 (Amphibia, Anura, Arthroleptidae)" checkinTime="1554510561182" checkinUser="pensoft" docAuthor="Griesbaum, Frederic, Hirschfeld, Mareike, F. Barej, Michael, Schmitz, Andreas, Rohrmoser, Mariam, Dahmen, Matthias, Muehlberger, Fabian, Christoph Liedtke, H., L. Gonwouo, Nono, Doumbia, Joseph &amp; Roedel, Mark-Oliver" docDate="2019" docId="961823580DB22D9D32D830E6C8FD981D" docLanguage="en" docName="ZoosystEvol 95(1): 133-160" docOrigin="Zoosystematics and Evolution 95 (1)" docSource="http://dx.doi.org/10.3897/zse.95.32793" docTitle="Astylosternus fallax Amiet 1978" docType="treatment" docVersion="4" lastPageNumber="138" masterDocId="7B4FFF9EFF8EFFA7275BFFDEFFA0A36C" masterDocTitle="Tadpoles of three western African frog genera: Astylosternus Werner, 1898, Nyctibates Boulenger, 1904, and Scotobleps Boulenger, 1900 (Amphibia, Anura, Arthroleptidae)" masterLastPageNumber="160" masterPageNumber="133" pageNumber="137" updateTime="1701316964302" updateUser="plazi">
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<mods:title id="F19FFA5A75DF943B72DFA405202D1D24">Tadpoles of three western African frog genera: Astylosternus Werner, 1898, Nyctibates Boulenger, 1904, and Scotobleps Boulenger, 1900 (Amphibia, Anura, Arthroleptidae)</mods:title>
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<mods:namePart id="9A26ED780FC53364B8B2F0A8E42A54F9">Griesbaum, Frederic</mods:namePart>
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<mods:namePart id="60D0FD556BA073361AF563E73C24612C">F. Barej, Michael</mods:namePart>
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<mods:namePart id="B1687457E309F4969B8858C416E08543">Schmitz, Andreas</mods:namePart>
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<mods:namePart id="106643F1605FC13A96EE731F895CD93E">Dahmen, Matthias</mods:namePart>
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<mods:namePart id="151745BAD0B5A050398CDB0238BC5113">Muehlberger, Fabian</mods:namePart>
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<mods:namePart id="1E475DEE06A9D75AF95BC61CF9D3118C">Christoph Liedtke, H.</mods:namePart>
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<mods:namePart id="8F65CF472B50EC615C10F8FB61F50007">L. Gonwouo, Nono</mods:namePart>
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<mods:namePart id="06A85A5FB218409B832E700D6FB448E3">Doumbia, Joseph</mods:namePart>
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<mods:namePart id="92BA067A52CE4F48CAD7AB0EE3047200">Roedel, Mark-Oliver</mods:namePart>
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<treatment id="961823580DB22D9D32D830E6C8FD981D" ID-GBIF-Taxon="156108983" LSID="urn:lsid:plazi:treatment:961823580DB22D9D32D830E6C8FD981D" httpUri="http://treatment.plazi.org/id/961823580DB22D9D32D830E6C8FD981D" lastPageId="5" lastPageNumber="138" pageId="4" pageNumber="137">
<subSubSection id="E7EE433DA09FBEE93C07FFDB192FB487" pageId="4" pageNumber="137" type="nomenclature">
<paragraph id="920816683DFCAD92CF99C79476A68F7C" pageId="4" pageNumber="137">
<taxonomicName id="6AE6326600679A06E12CEE33903A164D" ID-CoL="J643" authority="Amiet, 1978" authorityName="Amiet" authorityYear="1978" class="Amphibia" family="Arthroleptidae" genus="Astylosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Astylosternus fallax" order="Anura" pageId="4" pageNumber="137" phylum="Chordata" rank="species" species="fallax">Astylosternus cf. fallax Amiet, 1978</taxonomicName>
</paragraph>
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<subSubSection id="79CE1D3D40D0AA31674CB26F78939871" pageId="4" pageNumber="137" type="materials_examined">
<paragraph id="65E2774BADB098308E8551D50A9F7746" pageId="4" pageNumber="137">Material examined.</paragraph>
<paragraph id="DC743F7A6C054C774406C3EBC216DC3F" pageId="4" pageNumber="137">
ZMB 82867 (GenBank MK318852), Gosner stage 25, Cameroon, foot of Mt Nlonako, Ekomtolo, 477 m,
<geoCoordinate id="DD36678D5AE5B821CDB6696B6CA70A98" direction="north" orientation="latitude" precision="1" value="4.8329053">4°49'58.46&quot;N</geoCoordinate>
,
<geoCoordinate id="4138DD7A08EA1FA10DDB6B9A039A5884" direction="east" orientation="longitude" precision="1" value="9.92595">9°55'33.42&quot;E</geoCoordinate>
, 24 November 2011, leg. M.F. Barej, H.C. Liedtke, N.L. Gonwouo &amp; M. Hirschfeld.
</paragraph>
<paragraph id="9F481DB41185C591C01F6E5AC9538753" pageId="4" pageNumber="137">
The tadpole was caught in a forest stream. The genotyped tadpole has been compared with the two adults from Mount Nlonako, Cameroon (ZFMK 81164 and ZFMK 81608; GenBank MK318853, MK318854). The tadpole was genetically identical to these adults (compare Taxonomic remarks in the description of the
<taxonomicName id="CD73C785B7A0D461D83B81A8EC48BFB7" lsidName="A. fallax" pageId="4" pageNumber="137" rank="species" species="fallax">A. fallax</taxonomicName>
tadpole).
</paragraph>
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<subSubSection id="9C37F61CBD4E1E3DC21F0E42E3885017" lastPageId="5" lastPageNumber="138" pageId="4" pageNumber="137" type="description">
<paragraph id="30853084E8827DB9CBC936B877427467" pageId="4" pageNumber="137">Description.</paragraph>
<paragraph id="3434857B94F4283F2F0E14F09031C0FE" lastPageId="5" lastPageNumber="138" pageId="4" pageNumber="137">
Long slender tadpole with long, muscular tail (Fig. 5); body oval in dorsal view, bullet-shaped in
<pageBreakToken id="9FB7C37051136D2E55E2A10A3F5A67B2" pageId="5" pageNumber="138" start="start">lateral</pageBreakToken>
view; snout rounded in dorsal and lateral view; body length 31.7% of total length; body height 34.9% of body length; body width 48.4% of body length; eyes positioned dorsolaterally, eye diameter 10.7% of body length; nostrils positioned dorsolaterally, closer to snout tip than to eyes; inter-nostril distance 86.7% of interorbital distance; tail fins narrow, dorsal and ventral fin originating from tail base; ventral fin height narrower, 73.2% of dorsal fin height; highest part of tail approximately at two-thirds of the tail; body height 72.3% of maximum tail height; tail axis width 52.6% of body width; tail axis height 65.3% of maximum tail height; tail tip pointed; vent tube dextral; body with lateral sacs, extending from spiracle to end of body; short spiracle, sinistral; mouth positioned rostroventral, narrower than interorbital distance; keratodont formula 1:2+2+2/2+2:1; anterior lip with lateral papillae and big rostral gap; posterior lip with 2 or 3 rows of ca 20 massive papillae, papillae of marginal row are distinctly longer than of inner rows (Fig. 5c); black jaw sheaths massive and serrated; upper jaw sheath with a small medial fang, lower jaw U-shaped with a broad medial notch.
</paragraph>
<caption id="B8660F1D9A244C1A1E2F03604AF84E3D" pageId="5" pageNumber="138">
<paragraph id="79427A29F42929F9D1F027CB38CFA826" pageId="5" pageNumber="138">
Figure 5.
<taxonomicName id="F29BC54F02E3660D8EAAE8ECDA2824F4" class="Amphibia" family="Arthroleptidae" genus="Astylosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Astylosternus fallax" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="fallax">Astylosternus cf. fallax</taxonomicName>
(ZMB 82867, Gosner stage 25); a lateral, and b dorsal view; c oral disc; d keratodont arrangement; compare Fig. 6; black bars = 1 cm, white bar = 1 mm.
</paragraph>
</caption>
<paragraph id="E455E86C8155A6E4FC7C972E6FE2D881" pageId="5" pageNumber="138">The tadpole (Gosner stage 25) measures 97.4 mm in total length.</paragraph>
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<subSubSection id="548E71B36195A55DBCBBE5F470335D57" pageId="5" pageNumber="138" type="coloration in preservation">
<paragraph id="8B697B21AD9318B502EC2D2066FF7E45" pageId="5" pageNumber="138">Coloration in preservation.</paragraph>
<paragraph id="7978345DF61530BDA55ACDE34156846D" pageId="5" pageNumber="138">Basic color light brown with darker and speckled dorsal surface; venter light and grayish with few or no dark speckling; tail fins beige, semitransparent with irregular tiny dark brown spots. In life the color was of a duller, darker brown (Fig. 6).</paragraph>
<caption id="D1E463DD54F76E992BB18B064AADB805" pageId="5" pageNumber="138">
<paragraph id="C519A5B00D745C916DFDA14F649CA9EA" pageId="5" pageNumber="138">
Figure 6.
<taxonomicName id="FC542FCA17951FAA2D0811F0EA404B7B" class="Amphibia" family="Arthroleptidae" genus="Astylosternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Astylosternus fallax" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="fallax">Astylosternus cf. fallax</taxonomicName>
in life coloration (ZMB 82867, Gosner stage 25).
</paragraph>
</caption>
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