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<document id="B7CE9FE3774760F7D3F0CCCEB325FBE3" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FFB9FFEFFF20592138E1FA22" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius gracilis Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="48" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="F944A5CD15704AC06A17971CD972B332">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="7E2A8EE353F02AB4416E17284D1C11DB">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="B7338448D9BED5F8DA09FF13ACDE4502">Feng, Zegang</mods:namePart>
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<mods:namePart id="37FC8FADD617A1EFC8557177A841ECC4">Zhang, Feng</mods:namePart>
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<treatment id="FD7487B1FFB9FFEFFF20592138E1FA22" ID-DOI="http://doi.org/10.5281/zenodo.7795419" ID-GBIF-Taxon="209487180" ID-Zenodo-Dep="7795419" LSID="urn:lsid:plazi:treatment:FD7487B1FFB9FFEFFF20592138E1FA22" httpUri="http://treatment.plazi.org/id/FD7487B1FFB9FFEFFF20592138E1FA22" lastPageId="53" lastPageNumber="54" pageId="47" pageNumber="48">
<subSubSection id="3DC7652CFFB9FFF5FF2059213902F9FA" box="[151,786,1585,1612]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="756236A7FFB9FFF5FF2059213902F9FA" blockId="47.[151,786,1585,1647]" box="[151,786,1585,1612]" pageId="47" pageNumber="48">
<heading id="2E2A81CBFFB9FFF5FF2059213902F9FA" bold="true" box="[151,786,1585,1612]" fontSize="11" level="2" pageId="47" pageNumber="48" reason="4">
<emphasis id="47A9EAB5FFB9FFF5FF2059213902F9FA" bold="true" box="[151,786,1585,1612]" pageId="47" pageNumber="48">
<taxonomicName id="B2DD4D24FFB9FFF5FF2059213BC8F9FF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,472,1585,1612]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFB9FFF5FF2059213BC8F9FF" bold="true" box="[151,472,1585,1612]" italics="true" pageId="47" pageNumber="48">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C9A57CEFFB9FFF5FE685921382DF9F8" box="[479,573,1585,1611]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
‹KḦƟṁae›
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFB9FFF5FF2059453B04F9DC" box="[151,276,1621,1647]" pageId="47" pageNumber="48" type="description">
<paragraph id="756236A7FFB9FFF5FF2059453B04F9DC" blockId="47.[151,786,1585,1647]" box="[151,276,1621,1647]" pageId="47" pageNumber="48">
<figureCitation id="EDE62A22FFB9FFF5FF2059453B04F9DC" box="[151,276,1621,1647]" captionStart-0="FIGURE 37" captionStart-1="FIGURE38" captionStart-2="FIGURE 39" captionStart-3="FIGURE 40" captionStart-4="FIGURE 41" captionStartId-0="48.[151,250,1328,1352]" captionStartId-1="49.[151,250,1197,1221]" captionStartId-2="50.[151,250,1891,1915]" captionStartId-3="51.[151,250,1943,1967]" captionStartId-4="52.[151,250,1943,1967]" captionTargetBox-0="[182,1406,344,1304]" captionTargetBox-1="[166,1436,288,1162]" captionTargetBox-2="[151,1436,181,1868]" captionTargetBox-3="[179,1388,181,1916]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-86@48.[182,1406,344,1304]" captionTargetId-1="figure-62@49.[151,1437,273,1174]" captionTargetId-2="figure-20@50.[151,1436,181,1868]" captionTargetId-3="figure-18@51.[179,1408,181,1919]" captionTargetId-4="figure-20@52.[179,1408,181,1919]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-2="50" captionTargetPageId-3="51" captionTargetPageId-4="52" captionText-0="FIGURE 37. Xianren Cave, type locality of Tyrannochthonius gracilis sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where T. gracilis sp. nov. specimens were collected (red arrow); D. Live female of T. gracilis sp. nov. in its natural environment." captionText-1="FIGURE38.Tyrannochthoniusgracilissp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." captionText-2="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." captionText-3="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." captionText-4="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795244" figureDoi-1="http://doi.org/10.5281/zenodo.7795247" figureDoi-2="http://doi.org/10.5281/zenodo.7795249" figureDoi-3="http://doi.org/10.5281/zenodo.7795251" figureDoi-4="http://doi.org/10.5281/zenodo.7795253" httpUri-0="https://zenodo.org/record/7795244/files/figure.png" httpUri-1="https://zenodo.org/record/7795247/files/figure.png" httpUri-2="https://zenodo.org/record/7795249/files/figure.png" httpUri-3="https://zenodo.org/record/7795251/files/figure.png" httpUri-4="https://zenodo.org/record/7795253/files/figure.png" pageId="47" pageNumber="48">Figs 37–41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFB9FFF5FF2059883B11F81D" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="756236A7FFB9FFF5FF2059883B11F81D" blockId="47.[151,1437,1688,2038]" pageId="47" pageNumber="48">
<emphasis id="47A9EAB5FFB9FFF5FF2059883B58F901" bold="true" box="[151,328,1688,1714]" pageId="47" pageNumber="48">Type material.</emphasis>
<materialsCitation id="C5B53CFAFFB9FFF5FEE559883995F8AD" ID-GBIF-Occurrence="4071438361" collectingDate="2022-07-30" collectionCode="MHBU-HBUARA" collectorName="Yanmeng Hou &amp; Lu Zhang &amp; Jianzhou Sun &amp; Wenlong Fan" country="China" county="Hezhang County" elevation="1446" latitude="27.13096" location="Jiangnan Village" longLatPrecision="1" longitude="104.95568" municipality="Pingshan Town" pageId="47" pageNumber="48" specimenCode="MHBU-HBUARA#2022-490-01" specimenCount="1" stateProvince="Guizhou Province" typeStatus="holotype">
<typeStatus id="AA668805FFB9FFF5FEE559883BAAF901" box="[338,442,1688,1714]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
♂ (Ps.-
<specimenCode id="257B9EDCFFB9FFF5FDA75988399FF901" box="[528,911,1688,1714]" collectionCode="MHBU-HBUARA" pageId="47" pageNumber="48">MHBU-HBUARA#2022-490-01</specimenCode>
):
<collectingCountry id="0DCA7637FFB9FFF5FC13598839FAF901" box="[932,1002,1688,1714]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
,
<collectingRegion id="B719F845FFB9FFF5FC4F59883ED9F901" box="[1016,1225,1688,1714]" country="China" name="Guizhou" pageId="47" pageNumber="48">Guizhou Province</collectingRegion>
,
<collectingCounty id="9C034E2BFFB9FFF5FB6059883F89F901" box="[1239,1433,1688,1714]" pageId="47" pageNumber="48">Hezhang County</collectingCounty>
,
<collectingMunicipality id="9506ACDDFFB9FFF5FF2059AC3B55F964" box="[151,325,1724,1751]" pageId="47" pageNumber="48">Pingshan Town</collectingMunicipality>
,
<location id="7002607CFFB9FFF5FEE759AC381DF965" LSID="urn:lsid:plazi:treatment:FD7487B1FFB9FFEFFF20592138E1FA22:7002607CFFB9FFF5FEE759AC381DF965" box="[336,525,1724,1750]" country="China" county="Hezhang County" latitude="27.13096" longLatPrecision="1" longitude="104.95568" municipality="Pingshan Town" name="Jiangnan Village" pageId="47" pageNumber="48" stateProvince="Guizhou Province">Jiangnan Village</location>
,
<location id="7002607CFFB9FFF5FDA159AC38A2F965" LSID="urn:lsid:plazi:treatment:FD7487B1FFB9FFEFFF20592138E1FA22:7002607CFFB9FFF5FDA159AC38A2F965" box="[534,690,1724,1750]" country="China" county="Hezhang County" latitude="27.13096" longLatPrecision="1" longitude="104.95568" municipality="Pingshan Town" name="Xianren Cave" pageId="47" pageNumber="48" stateProvince="Guizhou Province">Xianren Cave</location>
, under stones, detritus and on the cave walls within 20–200 m from the cave entrance (Temperature: 14°C, Humidity: 85%) [
<geoCoordinate id="10E95060FFB9FFF5FC9159F139D6F94F" box="[806,966,1761,1788]" degrees="27" direction="north" minutes="7" orientation="latitude" pageId="47" pageNumber="48" precision="1" seconds="51.46" value="27.13096">27°7′51.46″N</geoCoordinate>
,
<geoCoordinate id="10E95060FFB9FFF5FC6359F13E9CF94F" box="[980,1164,1761,1788]" degrees="104" direction="east" minutes="57" orientation="longitude" pageId="47" pageNumber="48" precision="1" seconds="20.45" value="104.95568">104°57′20.45″E</geoCoordinate>
],
<elevation id="FEF0D194FFB9FFF5FB1659F03F28F949" box="[1185,1336,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.446" pageId="47" pageNumber="48" unit="m" value="1446.0">
<quantity id="B2259B42FFB9FFF5FB1659F03EE9F948" box="[1185,1273,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.446" pageId="47" pageNumber="48" unit="m" value="1446.0">1446 m</quantity>
a.s.l.
</elevation>
,
<date id="01631067FFB9FFF5FAFF59F03AC2F8AD" pageId="47" pageNumber="48" value="2022-07-30">
<collectingDate id="1127E98FFFB9FFF5FAFF59F03AC2F8AD" pageId="47" pageNumber="48" value="2022-07-30">30 July 2022</collectingDate>
</date>
,
<collectorName id="D8285371FFB9FFF5FF6B58153B90F8AC" box="[220,384,1797,1823]" pageId="47" pageNumber="48">Yanmeng Hou</collectorName>
,
<collectorName id="D8285371FFB9FFF5FE3C58153BEBF8AD" box="[395,507,1796,1823]" pageId="47" pageNumber="48">Lu Zhang</collectorName>
,
<collectorName id="D8285371FFB9FFF5FDB15814388DF8AD" box="[518,669,1796,1822]" pageId="47" pageNumber="48">Jianzhou Sun</collectorName>
&amp;
<collectorName id="D8285371FFB9FFF5FD0858143944F8AC" box="[703,852,1796,1823]" pageId="47" pageNumber="48">Wenlong Fan</collectorName>
leg.
</materialsCitation>
<materialsCitation id="C5B53CFAFFB9FFF5FC3C58153806F8F1" ID-GBIF-Occurrence="4071438314" collectionCode="HBUARA" country="China" location="Ps." pageId="47" pageNumber="48" specimenCode="MHBU-HBUARA#2022- 490-02, HBUARA#2022-490-14" specimenCount="13" specimenCount-male="13" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="AA668805FFB9FFF5FC3C581539ECF8AC" box="[907,1020,1797,1823]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="63DBFD2EFFB9FFF5FBB058143E2EF893" box="[1031,1086,1796,1824]" pageId="47" pageNumber="48" type="male">13 ♂</specimenCount>
(Ps.-
<specimenCode id="257B9EDCFFB9FFF5FBC558143AF6F8F1" pageId="47" pageNumber="48">MHBU-HBUARA#2022- 490-02</specimenCode>
–
<specimenCode id="257B9EDCFFB9FFF5FF415838381EF8F1" box="[246,526,1832,1858]" pageId="47" pageNumber="48">HBUARA#2022-490-14</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="C5B53CFAFFB9FFF5FD9558383BF1F8D5" ID-GBIF-Occurrence="4071438358" collectingDate="2022-07-30" collectionCode="HBUARA" collectorName="Yanmeng Hou &amp; Lu Zhang &amp; Jianzhou Sun &amp; Wenlong Fan" country="China" county="Hezhang County" elevation="110" latitude="27.13096" location="Ps." longLatPrecision="1" longitude="104.95568" municipality="Pingshan Town" pageId="47" pageNumber="48" specimenCode="MHBU-HBUARA#2022-490-15, HBUARA#2022-490-29" specimenCount="15" specimenCount-female="15" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="63DBFD2EFFB9FFF5FD955838384CF8F0" box="[546,604,1832,1859]" pageId="47" pageNumber="48" type="female">15 ♀</specimenCount>
(Ps.-
<specimenCode id="257B9EDCFFB9FFF5FD2658383E1AF8F0" box="[657,1034,1832,1859]" pageId="47" pageNumber="48">MHBU-HBUARA#2022-490-15</specimenCode>
–
<specimenCode id="257B9EDCFFB9FFF5FBAF58383F22F8F0" box="[1048,1330,1832,1859]" pageId="47" pageNumber="48">HBUARA#2022-490-29</specimenCode>
), all with the same data as the holotype
</materialsCitation>
;
<materialsCitation id="C5B53CFAFFB9FFF5FE5C585C3E22F8D5" ID-GBIF-Occurrence="4071438319" box="[491,1074,1868,1896]" collectionCode="GZC" country="China" location="Ps." pageId="47" pageNumber="48" specimenCode="MSWU- GZC190808-01, GZC190808-04" specimenCount="4" specimenCount-male="4" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="63DBFD2EFFB9FFF5FE5C585C3804F8DB" box="[491,532,1868,1896]" pageId="47" pageNumber="48" type="male">4 ♂</specimenCount>
(Ps.-
<specimenCode id="257B9EDCFFB9FFF5FDFF585C3977F8D5" box="[584,871,1868,1894]" pageId="47" pageNumber="48">MSWU- GZC190808-01</specimenCode>
–
<specimenCode id="257B9EDCFFB9FFF5FCC1585C3E3BF8D5" box="[886,1067,1868,1894]" collectionCode="GZC" pageId="47" pageNumber="48">GZC190808-04</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="C5B53CFAFFB9FFF5FB8B585C3AEDF81D" ID-GBIF-Occurrence="4071438324" collectingDate="2019-08-08" collectionCode="GZC" collectorName="Zegang Feng &amp; Zhaoyi Li &amp; Chen Zhang" country="China" county="Hezhang County" elevation="110" latitude="27.13096" location="Ps." longLatPrecision="1" longitude="104.95568" municipality="Pingshan Town" pageId="47" pageNumber="48" specimenCode="MSWU- GZC190808- 05, GZC190808-12" specimenCount="8" specimenCount-female="8" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="63DBFD2EFFB9FFF5FB8B585C3E76F8D4" box="[1084,1126,1868,1895]" pageId="47" pageNumber="48" type="female">8 ♀</specimenCount>
(Ps.-
<specimenCode id="257B9EDCFFB9FFF5FB2D585C3AA3F838" pageId="47" pageNumber="48">MSWU- GZC190808- 05</specimenCode>
–
<specimenCode id="257B9EDCFFB9FFF5FF7558603B66F838" box="[194,374,1904,1931]" collectionCode="GZC" pageId="47" pageNumber="48">GZC190808-12</specimenCode>
),
<date id="01631067FFB9FFF5FE3D58603820F839" box="[394,560,1904,1930]" pageId="47" pageNumber="48" value="2019-08-08">
<collectingDate id="1127E98FFFB9FFF5FE3D58603820F839" box="[394,560,1904,1930]" pageId="47" pageNumber="48" value="2019-08-08">8 August 2019</collectingDate>
</date>
,
<collectorName id="D8285371FFB9FFF5FD8C586138C1F838" box="[571,721,1905,1931]" pageId="47" pageNumber="48">Zegang Feng</collectorName>
,
<collectorName id="D8285371FFB9FFF5FD6C5860395BF839" box="[731,843,1904,1930]" pageId="47" pageNumber="48">Zhaoyi Li</collectorName>
&amp;
<collectorName id="D8285371FFB9FFF5FCD9586039E8F839" box="[878,1016,1904,1930]" pageId="47" pageNumber="48">Chen Zhang</collectorName>
leg., all with the same location as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFB9FFF5FF7058A83B8CF845" pageId="47" pageNumber="48" type="etymology">
<paragraph id="756236A7FFB9FFF5FF7058A83B8CF845" blockId="47.[151,1437,1688,2038]" pageId="47" pageNumber="48">
<emphasis id="47A9EAB5FFB9FFF5FF7058A83B5EF861" bold="true" box="[199,334,1976,2002]" pageId="47" pageNumber="48">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="B2DD4D24FFB9FFF5FC2058A839FDF861" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[919,1005,1976,2002]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="47A9EAB5FFB9FFF5FC2058A839FDF861" box="[919,1005,1976,2002]" italics="true" pageId="47" pageNumber="48">gracilis</emphasis>
</taxonomicName>
”, meaning gracile, which refers to the elongated coxal spines.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFA6FFEAFF705F863B9DFEF3" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="756236A7FFA6FFEAFF705F863B9DFEF3" blockId="48.[151,1437,150,320]" pageId="48" pageNumber="49">
<emphasis id="47A9EAB5FFA6FFEAFF705F863B97FF07" bold="true" box="[199,391,150,180]" pageId="48" pageNumber="49">
Diagnosis (J
<specimenCount id="63DBFD2EFFA6FFEAFED75F883B66FF07" box="[352,374,152,180]" pageId="48" pageNumber="49" type="female">♀</specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with 2 setae. Pedipalps slender, femur 8.50–8.88 (♂), 8.05–8.53 (
<specimenCount id="63DBFD2EFFA6FFEAFF155E123AA4FEAF" box="[162,180,258,284]" pageId="48" pageNumber="49" type="female">♀</specimenCount>
) times longer than broad; chela 8.79–9.04 (♂), 8.41–8.81 (
<specimenCount id="63DBFD2EFFA6FFEAFCE25E123977FEAF" box="[853,871,258,284]" pageId="48" pageNumber="49" type="female">♀</specimenCount>
) times longer than broad; only fixed chelal finger with intercalary teeth.
</paragraph>
</subSubSection>
<caption id="21A2662FFFA6FFEAFF205A203AE3FA23" ID-DOI="http://doi.org/10.5281/zenodo.7795244" ID-Zenodo-Dep="7795244" httpUri="https://zenodo.org/record/7795244/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1328,1352]" targetBox="[182,1406,344,1304]" targetPageId="48">
<paragraph id="756236A7FFA6FFEAFF205A203AE3FA23" blockId="48.[151,1437,1328,1424]" pageId="48" pageNumber="49">
<emphasis id="47A9EAB5FFA6FFEAFF205A203B31FAFB" bold="true" box="[151,289,1328,1352]" pageId="48" pageNumber="49">FIGURE 37.</emphasis>
Xianren Cave, type locality of
<taxonomicName id="B2DD4D24FFA6FFEAFDD15A203961FAFB" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[614,881,1328,1352]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA6FFEAFDD15A203961FAFB" box="[614,881,1328,1352]" italics="true" pageId="48" pageNumber="49">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA6FFEAFCCE5A2039DBFAFB" bold="true" box="[889,971,1328,1352]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="5C9A57CEFFA6FFEAFCCE5A2039DBFAFB" box="[889,971,1328,1352]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. Area where
<taxonomicName id="B2DD4D24FFA6FFEAFEA45A453B69FADF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[275,377,1364,1388]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA6FFEAFEA45A453B69FADF" box="[275,377,1364,1388]" italics="true" pageId="48" pageNumber="49">T. gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA6FFEAFE375A443BC1FADF" bold="true" box="[384,465,1364,1388]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="5C9A57CEFFA6FFEAFE375A443BC1FADF" box="[384,465,1364,1388]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected (red arrow); D. Live female of
<taxonomicName id="B2DD4D24FFA6FFEAFBA95A453E94FADF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1054,1156,1364,1388]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA6FFEAFBA95A453E94FADF" box="[1054,1156,1364,1388]" italics="true" pageId="48" pageNumber="49">T. gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA6FFEAFB3C5A443ECCFADF" bold="true" box="[1163,1244,1364,1388]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="5C9A57CEFFA6FFEAFB3C5A443ECCFADF" box="[1163,1244,1364,1388]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<subSubSection id="3DC7652CFFA6FFEFFF705AAE3F67FBE7" lastPageId="53" lastPageNumber="54" pageId="48" pageNumber="49" type="description">
<paragraph id="756236A7FFA6FFEAFF705AAE39E7FA6B" blockId="48.[151,1436,1470,2037]" box="[199,1015,1470,1496]" pageId="48" pageNumber="49">
<emphasis id="47A9EAB5FFA6FFEAFF705AAE3B49FA6B" bold="true" box="[199,345,1470,1496]" pageId="48" pageNumber="49">Description.</emphasis>
Males (
<typeStatus id="AA668805FFA6FFEAFE045AAE3807FA6B" box="[435,535,1470,1496]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
and
<typeStatus id="AA668805FFA6FFEAFDFA5AAE38AAFA6B" box="[589,698,1470,1496]" pageId="48" pageNumber="49" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFA6FFEAFD675AAE392CFA6B" box="[720,828,1470,1496]" captionStart="FIGURE38" captionStartId="49.[151,250,1197,1221]" captionTargetBox="[166,1436,288,1162]" captionTargetId="figure-62@49.[151,1437,273,1174]" captionTargetPageId="49" captionText="FIGURE38.Tyrannochthoniusgracilissp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795247" httpUri="https://zenodo.org/record/7795247/files/figure.png" pageId="48" pageNumber="49">Figs 38A</figureCitation>
,
<figureCitation id="EDE62A22FFA6FFEAFCF05AAE3986FA6B" box="[839,918,1470,1496]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="48" pageNumber="49">39A–F</figureCitation>
,
<figureCitation id="EDE62A22FFA6FFEAFC165AAE39D0FA6B" box="[929,960,1470,1496]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="48" pageNumber="49">40</figureCitation>
,
<figureCitation id="EDE62A22FFA6FFEAFC7C5AAE39FAFA6B" box="[971,1002,1470,1496]" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="48" pageNumber="49">41</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFA6FFEAFF705AF23EFFFA4F" blockId="48.[151,1436,1470,2037]" box="[199,1263,1506,1533]" pageId="48" pageNumber="49">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="756236A7FFA6FFEAFF7059163F7AF8F2" blockId="48.[151,1436,1470,2037]" pageId="48" pageNumber="49">
Cephalothorax (
<figureCitation id="EDE62A22FFA6FFEAFEC859163BFDF993" box="[383,493,1542,1568]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="48" pageNumber="49">Figs 39C</figureCitation>
,
<figureCitation id="EDE62A22FFA6FFEAFE4B59163821F993" box="[508,561,1542,1568]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="48" pageNumber="49">40A</figureCitation>
): carapace 0.99–1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exter ior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11–12 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FFA6FFEAFD7958363926F8F3" box="[718,822,1830,1856]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="48" pageNumber="49">Fig. 40C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="756236A7FFA6FFEBFF70585A3879FF4A" blockId="48.[151,1436,1470,2037]" lastBlockId="49.[151,1436,150,249]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
Chelicera (
<figureCitation id="EDE62A22FFA6FFEAFEF5585A3BBEF8D7" box="[322,430,1866,1892]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="48" pageNumber="49">Figs 39D</figureCitation>
,
<figureCitation id="EDE62A22FFA6FFEAFE0D585A3BFEF8D7" box="[442,494,1866,1892]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="48" pageNumber="49">40B</figureCitation>
): large, about as long as carapace, 2.50–2.53 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15–18 teeth, distal one largest; movable finger with 18–21 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="EDE62A22FFA6FFEAFBC658CA3EC9F847" box="[1137,1241,2010,2036]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="48" pageNumber="49">Fig. 40B</figureCitation>
). Serrula exterior with 21–22 and serrula interior with 14–15 blades. Rallum with 6 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FFA7FFEBFE415FCF384DFF4A" box="[502,605,223,249]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="49" pageNumber="50">Fig. 40E</figureCitation>
).
</paragraph>
<caption id="21A2662FFFA7FFEBFF205BBD3F8DFB75" ID-DOI="http://doi.org/10.5281/zenodo.7795247" ID-Zenodo-Dep="7795247" box="[151,1437,1197,1222]" httpUri="https://zenodo.org/record/7795247/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,250,1197,1221]" targetBox="[166,1436,288,1162]" targetPageId="49">
<paragraph id="756236A7FFA7FFEBFF205BBD3F8DFB75" blockId="49.[151,1437,1197,1222]" box="[151,1437,1197,1222]" pageId="49" pageNumber="50">
<emphasis id="47A9EAB5FFA7FFEBFF205BBD3B0CFB76" bold="true" box="[151,284,1197,1221]" pageId="49" pageNumber="50">FIGURE38.</emphasis>
<taxonomicName id="B2DD4D24FFA7FFEBFEA95BBE3834FB75" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[286,548,1198,1222]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA7FFEBFEA95BBE3834FB75" box="[286,548,1198,1222]" italics="true" pageId="49" pageNumber="50">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA7FFEBFD915BBE3863FB75" bold="true" box="[550,627,1198,1222]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="5C9A57CEFFA7FFEBFD915BBE3863FB75" box="[550,627,1198,1222]" pageId="49" pageNumber="50" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
,A.Holotype male,habitus (dorsal view);B.Paratype female,habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="756236A7FFA7FFEBFF705BE43B44F8FD" blockId="49.[151,1437,1268,2014]" pageId="49" pageNumber="50">
Pedipalp (
<figureCitation id="EDE62A22FFA7FFEBFE8B5BE43BE1FABD" box="[316,497,1268,1294]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="49" pageNumber="50">Figs 39A, B, E</figureCitation>
,
<figureCitation id="EDE62A22FFA7FFEBFDB65BE43824FABD" box="[513,564,1268,1294]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="49" pageNumber="50">40D</figureCitation>
,
<figureCitation id="EDE62A22FFA7FFEBFDF45BE4388BFABD" box="[579,667,1268,1294]" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="49" pageNumber="50">41A, B</figureCitation>
): long and slender, trochanter 1.24–1.40, femur 8.50–8.88, patella 2.70–3.05, chela 8.79–9.04, hand 3.08–3.30 times longer than broad; femur 2.47–2.48 times longer than patella; movable chelal finger 1.63–1.76 times longer than hand and 0.60–0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FFA7FFEBFD585A703948FAC9" box="[751,856,1376,1402]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="49" pageNumber="50">Figs 39E</figureCitation>
,
<figureCitation id="EDE62A22FFA7FFEBFCD35A703989FAC9" box="[868,921,1376,1402]" captionStart="FIGURE 40" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[179,1388,181,1916]" captionTargetId="figure-18@51.[179,1408,181,1919]" captionTargetPageId="51" captionText="FIGURE 40. Tyrannochthonius gracilis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="49" pageNumber="50">40D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="47A9EAB5FFA7FFEBFAE15AB83F7CFA71" box="[1366,1388,1448,1474]" italics="true" pageId="49" pageNumber="50">ib</emphasis>
and
<emphasis id="47A9EAB5FFA7FFEBFF205ADC3AA8FA55" box="[151,184,1484,1510]" italics="true" pageId="49" pageNumber="50">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="47A9EAB5FFA7FFEBFC335ADC398EFA55" box="[900,926,1484,1510]" italics="true" pageId="49" pageNumber="50">eb</emphasis>
,
<emphasis id="47A9EAB5FFA7FFEBFC1A5ADC39C2FA55" box="[941,978,1484,1510]" italics="true" pageId="49" pageNumber="50">esb</emphasis>
and
<emphasis id="47A9EAB5FFA7FFEBFBBE5ADD3E33FA55" box="[1033,1059,1485,1510]" italics="true" pageId="49" pageNumber="50">ist</emphasis>
forming a nearly straight oblique row at base of fixed chelal finger;
<emphasis id="47A9EAB5FFA7FFEBFDAA5AE1383DF9B9" box="[541,557,1521,1546]" italics="true" pageId="49" pageNumber="50">it</emphasis>
slightly distal to
<emphasis id="47A9EAB5FFA7FFEBFD455AE13901F9B9" box="[754,785,1521,1546]" italics="true" pageId="49" pageNumber="50">est</emphasis>
, situated subdistally;
<emphasis id="47A9EAB5FFA7FFEBFBB05AE13E0BF9B9" box="[1031,1051,1521,1546]" italics="true" pageId="49" pageNumber="50">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="47A9EAB5FFA7FFEBFDB35904380EF99D" box="[516,542,1556,1582]" italics="true" pageId="49" pageNumber="50">dx</emphasis>
situated distal to
<emphasis id="47A9EAB5FFA7FFEBFD69590538E2F99D" box="[734,754,1557,1582]" italics="true" pageId="49" pageNumber="50">et</emphasis>
;
<emphasis id="47A9EAB5FFA7FFEBFCB759043909F99D" box="[768,793,1556,1582]" italics="true" pageId="49" pageNumber="50">sb</emphasis>
closer to
<emphasis id="47A9EAB5FFA7FFEBFC3459043981F99D" box="[899,913,1556,1582]" italics="true" pageId="49" pageNumber="50">b</emphasis>
than to
<emphasis id="47A9EAB5FFA7FFEBFC5F590539EBF99D" box="[1000,1019,1557,1582]" italics="true" pageId="49" pageNumber="50">st</emphasis>
;
<emphasis id="47A9EAB5FFA7FFEBFBBF59043E06F99D" box="[1032,1046,1556,1582]" italics="true" pageId="49" pageNumber="50">b</emphasis>
and
<emphasis id="47A9EAB5FFA7FFEBFBFE59053E41F99D" box="[1097,1105,1557,1582]" italics="true" pageId="49" pageNumber="50">t</emphasis>
situated subdistally,
<emphasis id="47A9EAB5FFA7FFEBFA8F59053F50F99D" box="[1336,1344,1557,1582]" italics="true" pageId="49" pageNumber="50">t</emphasis>
situated distal to
<emphasis id="47A9EAB5FFA7FFEBFF4E59283B17F9E1" box="[249,263,1592,1618]" italics="true" pageId="49" pageNumber="50">b</emphasis>
and slightly proximal to
<emphasis id="47A9EAB5FFA7FFEBFD9159293855F9E1" box="[550,581,1593,1618]" italics="true" pageId="49" pageNumber="50">est</emphasis>
(
<figureCitation id="EDE62A22FFA7FFEBFDE2592838AEF9E1" box="[597,702,1592,1618]" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="49" pageNumber="50">Fig. 41A</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="47A9EAB5FFA7FFEBFED1594D3B90F9C5" box="[358,384,1629,1654]" italics="true" pageId="49" pageNumber="50">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28–30 macrodenticles, long and pointed, plus 14–
<specimenCount id="63DBFD2EFFA7FFEBFC5659B43E6CF90D" box="[993,1148,1700,1727]" pageId="49" pageNumber="50" type="intercalary">15 intercalary</specimenCount>
microdenticles,
<quantity id="B2259B42FFA7FFEBFA8F59B43F8DF90D" box="[1336,1437,1700,1727]" metricMagnitude="0" metricUnit="m" metricValue="1.1049" metricValueMax="1.143" metricValueMin="1.0668" pageId="49" pageNumber="50" unit="in" value="43.5" valueMax="45.0" valueMin="42.0">42–45 in</quantity>
total; movable chelal finger with 34–37 macrodenticles (smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 5–6 vestigial, rounded and contiguous basal teeth,
<quantity id="B2259B42FFA7FFEBFC2859FC3E13F8B5" box="[927,1027,1772,1798]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.0668" metricValueMin="1.016" pageId="49" pageNumber="50" unit="in" value="41.0" valueMax="42.0" valueMin="40.0">40–42 in</quantity>
total, without intercalary teeth (
<figureCitation id="EDE62A22FFA7FFEBFAD859FC3ADCF899" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="49" pageNumber="50">Fig. 41A</figureCitation>
). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FFA7FFEBFF2858243B19F8FD" box="[159,265,1844,1870]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="49" pageNumber="50">Figs 39B</figureCitation>
,
<figureCitation id="EDE62A22FFA7FFEBFEA358243B57F8FD" box="[276,327,1844,1870]" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="49" pageNumber="50">41B</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFA7FFEBFF7058483ED9F86D" blockId="49.[151,1437,1268,2014]" pageId="49" pageNumber="50">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 4: T2T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III–XII: 11–12: 10: 8: 7–8: 6–7: 7: 7: 7: 0: 2. Anterior genital operculum with 10–12 setae, genital opening slit-like, with 9–11 marginal setae on each side,
<quantity id="B2259B42FFA7FFEBFC1A58D43E00F86D" box="[941,1040,1988,2014]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="8.382" metricValueMin="7.874" pageId="49" pageNumber="50" unit="in" value="32.0" valueMax="33.0" valueMin="31.0">31–33 in</quantity>
total (
<figureCitation id="EDE62A22FFA7FFEBFBEF58D43EADF86D" box="[1112,1213,1988,2014]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="49" pageNumber="50">Fig. 39F</figureCitation>
).
</paragraph>
<caption id="21A2662FFFA4FFE8FF2058733994F877" ID-DOI="http://doi.org/10.5281/zenodo.7795249" ID-Zenodo-Dep="7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,250,1891,1915]" targetBox="[151,1436,181,1868]" targetPageId="50">
<paragraph id="756236A7FFA4FFE8FF2058733994F877" blockId="50.[151,1436,1891,1988]" pageId="50" pageNumber="51">
<emphasis id="47A9EAB5FFA4FFE8FF2058733B31F8C8" bold="true" box="[151,289,1891,1915]" pageId="50" pageNumber="51">FIGURE 39.</emphasis>
<taxonomicName id="B2DD4D24FFA4FFE8FE9E58743824F8CF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[297,564,1892,1916]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA4FFE8FE9E58743824F8CF" box="[297,564,1892,1916]" italics="true" pageId="50" pageNumber="51">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA4FFE8FD8B5874389EF8CF" bold="true" box="[572,654,1892,1916]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5C9A57CEFFA4FFE8FD8B5874389EF8CF" box="[572,654,1892,1916]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view).
</paragraph>
</caption>
<caption id="21A2662FFFA5FFE9FF2058873851F84B" ID-DOI="http://doi.org/10.5281/zenodo.7795251" ID-Zenodo-Dep="7795251" httpUri="https://zenodo.org/record/7795251/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1943,1967]" targetBox="[179,1388,181,1916]" targetPageId="51">
<paragraph id="756236A7FFA5FFE9FF2058873851F84B" blockId="51.[151,1436,1943,2040]" pageId="51" pageNumber="52">
<emphasis id="47A9EAB5FFA5FFE9FF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="51" pageNumber="52">FIGURE 40.</emphasis>
<taxonomicName id="B2DD4D24FFA5FFE9FE9E58883824F803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[297,564,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA5FFE9FE9E58883824F803" box="[297,564,1944,1968]" italics="true" pageId="51" pageNumber="52">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA5FFE9FD8A5888389FF803" bold="true" box="[573,655,1944,1968]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="5C9A57CEFFA5FFE9FD8A5888389FF803" box="[573,655,1944,1968]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E).
</paragraph>
</caption>
<caption id="21A2662FFFA2FFEEFF2058873F70F860" ID-DOI="http://doi.org/10.5281/zenodo.7795253" ID-Zenodo-Dep="7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="52">
<paragraph id="756236A7FFA2FFEEFF2058873F70F860" blockId="52.[151,1436,1943,2004]" pageId="52" pageNumber="53">
<emphasis id="47A9EAB5FFA2FFEEFF2058873B32F81C" bold="true" box="[151,290,1943,1967]" pageId="52" pageNumber="53">FIGURE 41.</emphasis>
<taxonomicName id="B2DD4D24FFA2FFEEFE9D58883826F803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[298,566,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA2FFEEFE9D58883826F803" box="[298,566,1944,1968]" italics="true" pageId="52" pageNumber="53">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA2FFEEFD8858883881F803" bold="true" box="[575,657,1944,1968]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="5C9A57CEFFA2FFEEFD8858883881F803" box="[575,657,1944,1968]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm.
</paragraph>
</caption>
<paragraph id="756236A7FFA3FFEFFF705F873992FED6" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">
Legs (
<figureCitation id="EDE62A22FFA3FFEFFEBB5F873B84FF02" box="[268,404,151,177]" captionStart="FIGURE 41" captionStartId="52.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@52.[179,1408,181,1919]" captionTargetPageId="52" captionText="FIGURE 41. Tyrannochthonius gracilis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795253" httpUri="https://zenodo.org/record/7795253/files/figure.png" pageId="53" pageNumber="54">Fig. 41C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76–1.77 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.37–2.41 times longer than tibia. Femoropatella of leg IV 3.93–4.41 times longer than deep; tibia 6.75–6.83 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.80–4.00 times longer than deep (TS= 0.23–0.26), tarsus 14.14–14.57 times longer than deep and 2.48–2.68 times longer than metatarsus (TS= 0.27). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FFA3FFEFFF705E7F3EF0FDAA" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">
Adult females (
<typeStatus id="AA668805FFA3FFEFFEC55E7F3BF1FE3A" box="[370,481,367,393]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFA3FFEFFE405E7F3872FE3A" box="[503,610,367,393]" captionStart="FIGURE 37" captionStartId="48.[151,250,1328,1352]" captionTargetBox="[182,1406,344,1304]" captionTargetId="figure-86@48.[182,1406,344,1304]" captionTargetPageId="48" captionText="FIGURE 37. Xianren Cave, type locality of Tyrannochthonius gracilis sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where T. gracilis sp. nov. specimens were collected (red arrow); D. Live female of T. gracilis sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7795244" httpUri="https://zenodo.org/record/7795244/files/figure.png" pageId="53" pageNumber="54">Figs 37D</figureCitation>
,
<figureCitation id="EDE62A22FFA3FFEFFDDB5E7F388DFE3A" box="[620,669,367,393]" captionStart="FIGURE38" captionStartId="49.[151,250,1197,1221]" captionTargetBox="[166,1436,288,1162]" captionTargetId="figure-62@49.[151,1437,273,1174]" captionTargetPageId="49" captionText="FIGURE38.Tyrannochthoniusgracilissp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795247" httpUri="https://zenodo.org/record/7795247/files/figure.png" pageId="53" pageNumber="54">38B</figureCitation>
,
<figureCitation id="EDE62A22FFA3FFEFFD1F5E7F38CCFE3A" box="[680,732,367,393]" captionStart="FIGURE 39" captionStartId="50.[151,250,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-20@50.[151,1436,181,1868]" captionTargetPageId="50" captionText="FIGURE 39. Tyrannochthonius gracilis sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795249" httpUri="https://zenodo.org/record/7795249/files/figure.png" pageId="53" pageNumber="54">39G</figureCitation>
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV–XII: 12–14: 8: 7–8: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10–12 marginal setae,
<quantity id="B2259B42FFA3FFEFFB675EA73F23FE62" box="[1232,1331,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="53" pageNumber="54" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.56 times longer than deep (TS= 0.22–0.27), tarsus 13.25–14.57 times longer than deep and 2.49–2.59 times longer than metatarsus (TS= 0.21–0.26).
</paragraph>
<paragraph id="756236A7FFA3FFEFFF705D333BFDFCEE" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.96–2.06. Pedipalps: trochanter 0.26–0.28/0.20–0.21 (1.24–1.40), femur 1.51–1.53/0.17–0.18 (8.50– 8.88), patella 0.61–0.62/0.20–0.23 (2.70–3.05), chela 2.08–2.11/0.23–0.24 (8.79–9.04), hand 0.74–0.76/0.23–0.24 (3.08–3.30), movable chelal finger length 1.24–1.30. Chelicera 0.80–0.86/0.32–0.34 (2.50–2.53), movable finger length 0.43–0.45. Carapace 0.68–0.71/0.68–0.72 (0.99–1.00). Leg I: trochanter 0.23/0.18–0.19 (1.21–1.28), femur 0.85–0.88/0.09–0.10 (8.80–9.44), patella 0.48–0.50/0.09 (5.33–5.56), tibia 0.38–0.39/0.06–0.08 (4.88–6.33), tarsus 0.90–0.94/0.06–0.07 (13.43–15.00). Leg IV: trochanter 0.32–0.33/0.18–0.20 (1.65–1.78), femoropatella 1.18–1.19/0.27–0.30 (3.93–4.41), tibia 0.81–0.82/0.12 (6.75–6.83), metatarsus 0.38–0.40/0.10 (3.80–4.00), tarsus 0.99–1.02/0.07 (14.14–14.57).</paragraph>
<paragraph id="756236A7FFA3FFEFFF705C763F67FBE7" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">Females: body length 2.12–2.58. Pedipalps: trochanter 0.27–0.30/0.21 (1.29–1.43), femur 1.61–1.62/0.19–0.20 (8.05–8.53), patella 0.64–0.67/0.23–0.24 (2.67–2.91), chela 2.27–2.29/0.26–0.27 (8.41–8.81), hand 0.78–0.83/0.26– 0.27 (3.00–3.07), movable chelal finger length 1.39–1.43. Chelicera 0.89–0.91/0.36–0.38 (2.39–2.47), movable finger length 0.48–0.49. Carapace 0.74–0.76/0.74–0.76 (0.97–1.03). Leg I: trochanter 0.23–0.24/0.18–0.19 (1.21– 1.33), femur 0.89–0.92/0.09–0.10 (9.20–9.89), patella 0.47/0.09 (5.22), tibia 0.39–0.41/0.07–0.08 (5.13–5.57), tarsus 0.95–0.98/0.07 (13.57–14.00). Leg IV: trochanter 0.35/0.19 (1.84), femoropatella 1.23–1.25/0.27–0.28 (4.39– 4.63), tibia 0.82–0.86/0.12 (6.83–7.17), metatarsus 0.41/0.09 (4.56), tarsus 1.02–1.06/0.07–0.08 (13.25–14.57).</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFA3FFEFFF705B4C3BFEFADE" pageId="53" pageNumber="54" type="discussion">
<paragraph id="756236A7FFA3FFEFFF705B4C3F6AFB53" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">
<emphasis id="47A9EAB5FFA3FFEFFF705B4C3B2BFBC5" bold="true" box="[199,315,1116,1142]" pageId="53" pageNumber="54">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FFA3FFEFFEF55B4C3879FBC5" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[322,617,1116,1142]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA3FFEFFEF55B4C3879FBC5" box="[322,617,1116,1142]" italics="true" pageId="53" pageNumber="54">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA3FFEFFDD85B4C38D9FBC5" bold="true" box="[623,713,1116,1142]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="5C9A57CEFFA3FFEFFDD85B4C38D9FBC5" box="[623,713,1116,1142]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="B2DD4D24FFA3FFEFFCEC5B4D3E00FBC5" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[859,1040,1116,1142]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="parcidentatus" status="sp. nov.">
<emphasis id="47A9EAB5FFA3FFEFFCEC5B4D3E00FBC5" box="[859,1040,1116,1142]" italics="true" pageId="53" pageNumber="54">T. parcidentatus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA3FFEFFBA05B4C3E60FBC5" bold="true" box="[1047,1136,1116,1142]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="5C9A57CEFFA3FFEFFBA05B4C3E60FBC5" box="[1047,1136,1116,1142]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having intercalary teeth on the fixed chelal finger only and a pointed, triangular epistome, but differs by a smaller body size but longer appendages (body length 1.96–2.06 (♂), 2.12–2.58 (
<specimenCount id="63DBFD2EFFA3FFEFFD7E5BB338CBFB0E" box="[713,731,1187,1213]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
)
<emphasis id="47A9EAB5FFA3FFEFFD5A5BB3391CFB0F" box="[749,780,1187,1212]" italics="true" pageId="53" pageNumber="54">vs.</emphasis>
2.29–2.62 (♂), 2.64–2.66 (
<specimenCount id="63DBFD2EFFA3FFEFFBF75BB33E42FB0E" box="[1088,1106,1187,1213]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
) mm; chela length 2.08–2.11 (♂), 2.27–2.29 (
<specimenCount id="63DBFD2EFFA3FFEFFEF85BD63B71FB53" box="[335,353,1222,1248]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
)
<emphasis id="47A9EAB5FFA3FFEFFEC55BD73B81FB53" box="[370,401,1223,1248]" italics="true" pageId="53" pageNumber="54">vs.</emphasis>
1.89–1.93 (♂), 2.02–2.05 (
<specimenCount id="63DBFD2EFFA3FFEFFD715BD638C8FB53" box="[710,728,1222,1248]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
) mm) and more fixed chelal finger teeth (42‒45
<emphasis id="47A9EAB5FFA3FFEFFB495BD73F0CFB53" box="[1278,1308,1223,1248]" italics="true" pageId="53" pageNumber="54">vs.</emphasis>
30‒31).
</paragraph>
<paragraph id="756236A7FFA3FFEFFF705BF93BFEFADE" blockId="53.[151,1437,151,1425]" pageId="53" pageNumber="54">
<taxonomicName id="B2DD4D24FFA3FFEFFF705BF93BFDFAB0" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[199,493,1257,1283]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="47A9EAB5FFA3FFEFFF705BF93BFDFAB0" box="[199,493,1257,1283]" italics="true" pageId="53" pageNumber="54">Tyrannochthonius gracilis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA3FFEFFE445BF9385CFAB0" bold="true" box="[499,588,1257,1283]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="5C9A57CEFFA3FFEFFE445BF9385CFAB0" box="[499,588,1257,1283]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="B2DD4D24FFA3FFEFFCC05BFA39F8FAB0" authorityName="Mahnert" authorityYear="2009" box="[887,1000,1257,1283]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="akaelus">
<emphasis id="47A9EAB5FFA3FFEFFCC05BFA39F8FAB0" box="[887,1000,1257,1283]" italics="true" pageId="53" pageNumber="54">T. akaelus</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FFA3FFEFFC425BFA3EB7FAB0" authorityName="Mahnert" authorityYear="2009" box="[1013,1191,1257,1283]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="antridraconis">
<emphasis id="47A9EAB5FFA3FFEFFC425BFA3EB7FAB0" box="[1013,1191,1257,1283]" italics="true" pageId="53" pageNumber="54">T. antridraconis</emphasis>
</taxonomicName>
and
<taxonomicName id="B2DD4D24FFA3FFEFFB6A5BFA3F8CFAB0" authorityName="Mahnert" authorityYear="2009" box="[1245,1436,1257,1283]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ganshuanensis">
<emphasis id="47A9EAB5FFA3FFEFFB6A5BFA3F8CFAB0" box="[1245,1436,1257,1283]" italics="true" pageId="53" pageNumber="54">T. ganshuanensis</emphasis>
</taxonomicName>
by the presence of intercalary teeth on the fixed chelal finger only, from
<taxonomicName id="B2DD4D24FFA3FFEFFC645A1D3E54FA95" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[979,1092,1292,1318]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="47A9EAB5FFA3FFEFFC645A1D3E54FA95" box="[979,1092,1292,1318]" italics="true" pageId="53" pageNumber="54">T. harveyi</emphasis>
</taxonomicName>
by the presence of intercalary teeth on the fixed chelal finger, from
<taxonomicName id="B2DD4D24FFA3FFEFFD985A20388BFAFA" authorityName="Gao, Wynne &amp; Fe.Zhang" authorityYear="2018" box="[559,667,1327,1353]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="chixing">
<emphasis id="47A9EAB5FFA3FFEFFD985A20388BFAFA" box="[559,667,1327,1353]" italics="true" pageId="53" pageNumber="54">T. chixing</emphasis>
</taxonomicName>
and
<taxonomicName id="B2DD4D24FFA3FFEFFD785A203908FAFA" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[719,792,1327,1353]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="zhai">
<emphasis id="47A9EAB5FFA3FFEFFD785A203908FAFA" box="[719,792,1327,1353]" italics="true" pageId="53" pageNumber="54">T. zhai</emphasis>
</taxonomicName>
by the number of setae on tergites I–IV (2
<emphasis id="47A9EAB5FFA3FFEFFB595A203F1CFAFA" box="[1262,1292,1328,1353]" italics="true" pageId="53" pageNumber="54">vs.</emphasis>
4) (
<bibRefCitation id="114C4B56FFA3FFEFFA805A3F3AC5FADE" author="Mahnert, V." pageId="53" pageNumber="54" pagination="185 - 201" refId="ref77141" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert 2009</bibRefCitation>
;
<bibRefCitation id="114C4B56FFA3FFEFFF565A433B87FADE" author="Gao, Z. Z. &amp; Wynne, J. J. &amp; Zhang, F." box="[225,407,1363,1389]" pageId="53" pageNumber="54" pagination="345 - 354" refId="ref76312" refString="Gao, Z. Z., Wynne, J. J. &amp; Zhang, F. (2018) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. Journal of Arachnology, 46, 345 - 354. https: // doi. org / 10.1636 / JoA-S- 17 - 063.1" type="journal article" year="2018">
Gao
<emphasis id="47A9EAB5FFA3FFEFFEAF5A433B43FADE" box="[280,339,1363,1389]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2018
</bibRefCitation>
,
<bibRefCitation id="114C4B56FFA3FFEFFE145A433BF0FADE" author="Gao, Z. Z. &amp; Zhang, F. &amp; Chen, H. M." box="[419,480,1363,1389]" pageId="53" pageNumber="54" pagination="572 - 580" refId="ref76374" refString="Gao, Z. Z., Zhang, F. &amp; Chen, H. M. (2020) Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa, 4853 (4), 572 - 580. https: // doi. org / 10.11646 / zootaxa. 4853.4.6" type="journal article" year="2020">2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFA3FFEFFF705A6638E1FA22" box="[199,753,1398,1425]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="756236A7FFA3FFEFFF705A6638E1FA22" blockId="53.[151,1437,151,1425]" box="[199,753,1398,1425]" pageId="53" pageNumber="54">
<emphasis id="47A9EAB5FFA3FFEFFF705A663B71FA23" bold="true" box="[199,353,1398,1424]" pageId="53" pageNumber="54">Distribution.</emphasis>
Known only from the
<typeStatus id="AA668805FFA3FFEFFDD75A673880FA22" box="[608,656,1399,1425]" pageId="53" pageNumber="54">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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