treatments-xml/data/1D/18/DD/1D18DDCCD18C5B3E8A2EEF56D702C345.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis &amp; Ribeiro, Guilherme Cunha" docDate="2022" docId="1D18DDCCD18C5B3E8A2EEF56D702C345" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) immaculatus Santos &amp; Santos &amp; Ribeiro 2022, sp. nov." docType="treatment" docUuid="7211DAE6-7F6B-439D-ADDD-5C0AA2561FD7" docUuidSource="ZooBank" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Daubian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>439</mods:start>
<mods:end>494</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:7211DAE6-7F6B-439D-ADDD-5C0AA2561FD7" httpUri="http://treatment.plazi.org/id/1D18DDCCD18C5B3E8A2EEF56D702C345" lastPageNumber="439" pageId="0" pageNumber="439">
<subSubSection pageId="0" pageNumber="439" type="nomenclature">
<paragraph pageId="0" pageNumber="439">
3.3.14. 
<taxonomicName LSID="http://zoobank.org/7211DAE6-7F6B-439D-ADDD-5C0AA2561FD7" authority="Santos &amp; Santos &amp; Ribeiro, 2022" authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) immaculatus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="immaculatus" status="sp. nov." subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) immaculatus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="439">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Holotype</typeStatus>
</emphasis>
: 
<specimenCount>♂</specimenCount>
, Chile, Osorno, Pucotrihue [
<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32′S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42 W</geoCoordinate>
], 10/ 
<date value="1968-04-12">12-IV-1968</date>
, 
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)*. 
<materialsCitation collectingDate="1968-04-12" county="Pena" latitude="-40.583332" location="Chile" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Paratypes</typeStatus>
</emphasis>
: 
<specimenCount type="male">1 ♂</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:1D18DDCCD18C5B3E8A2EEF56D702C345:A02E71E7701E16AACB4861C7A36EBB96" county="Pena" latitude="-40.583332" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" name="Chile">Chile</location>
, 
<location LSID="urn:lsid:plazi:treatment:1D18DDCCD18C5B3E8A2EEF56D702C345:B0FAE46514EB247EFD99E447D2D63914" county="Pena" latitude="-40.583332" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" name="Osorno">Osorno</location>
, 
<location LSID="urn:lsid:plazi:treatment:1D18DDCCD18C5B3E8A2EEF56D702C345:B2A22D9BA0EB4533A5330804242D1A39" county="Pena" latitude="-40.583332" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" name="Bahia Mansa">Bahia Mansa</location>
[
<geoCoordinate degrees="40" direction="south" minutes="35" orientation="latitude" precision="925" value="-40.583332">40°35′S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="44" orientation="longitude" precision="925" value="-73.73333">73°44′W</geoCoordinate>
], 
<collectingMunicipality>South of Pucotrihue</collectingMunicipality>
, 
<collectingDate value="1968-04-12">12-IV-1968</collectingDate>
, 
<normalizedToken originalValue="Peña">
<collectingCounty>Pena</collectingCounty>
</normalizedToken>
(USNM)
</materialsCitation>
; 
<materialsCitation collectingDate="1955-03-20" elevation="810" latitude="-39.933334" location="Chile" longLatPrecision="1165" longitude="-73.566666" municipality="Chaihuin" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount type="male">2 ♂</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:1D18DDCCD18C5B3E8A2EEF56D702C345:80D6997DCB47FB34D56BC08BE205261D" latitude="-39.933334" longLatPrecision="1165" longitude="-73.566666" municipality="Chaihuin" name="Chile">Chile</location>
, 
<collectingMunicipality>Chaihuin</collectingMunicipality>
[
<geoCoordinate degrees="39" direction="south" minutes="56" orientation="latitude" precision="925" value="-39.933334">39°56′S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="34" orientation="longitude" precision="925" value="-73.566666">73°34′W</geoCoordinate>
], 
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.1" unit="m" value="810.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.1" unit="m" value="810.0">810 m</elevation>
</quantity>
, 
<collectingDate value="1955-03-20">20-III-1955</collectingDate>
, 
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)
</materialsCitation>
. - 
<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
: 
<materialsCitation collectingDate="1952-01-15" collectingDateMax="1954-02-08" collectingDateMin="1952-01-15" collectorName="Chiloe Is., Pena, Aulen, L. P. Guzman, Chaiten, Aucar" country="CHILE" latitude="-42.15" location="CHILE" longLatPrecision="1152" longitude="-73.5" specimenCount="2" specimenCount-female="2">
<emphasis bold="true" pageId="0" pageNumber="439">
<collectingCountry name="Chile">CHILE</collectingCountry>
.
</emphasis>
<specimenCount type="female">2 ♀</specimenCount>
and 1 [sex unknown], 
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
, Chepu [
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
], 1/ 
<collectingDate value="1952-02-03">3-II-1952</collectingDate>
, 
<normalizedToken originalValue="Peña">
<collectorName>Pena</collectorName>
</normalizedToken>
(USNM); 2 [sex unknown], 
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
, 
<collectorName>Aulen</collectorName>
[
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
], 
<collectingDate value="1952-02-08">8-II-1952</collectingDate>
, 
<collectorName>L.P. Guzman</collectorName>
(USNM); 1 [sex unknown], 
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
, 
<collectorName>Chaiten</collectorName>
[
<geoCoordinate degrees="42" direction="south" minutes="58" orientation="latitude" precision="925" value="-42.966667">42°58′S</geoCoordinate>
<geoCoordinate degrees="72" direction="west" minutes="32" orientation="longitude" precision="925" value="-72.53333">72°32′W</geoCoordinate>
], 5/ 
<collectingDate value="1954-02-08">8-II-1954</collectingDate>
, 
<normalizedToken originalValue="Peña">
<collectorName>Pena</collectorName>
</normalizedToken>
(USNM); 1 [sex unknown], 
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
, 
<collectorName>Aucar</collectorName>
[
<geoCoordinate degrees="42" direction="south" minutes="9" orientation="latitude" precision="925" value="-42.15">42°9′S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="30" orientation="longitude" precision="925" value="-73.5">73°30′W</geoCoordinate>
], 
<collectingDate value="1952-01-15">15-I-1952</collectingDate>
, 
<collectorName>L.P. Guzman</collectorName>
(USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="etymology">
<paragraph pageId="0" pageNumber="439">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="439">
The word 
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus immaculatus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="immaculatus">
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus immaculatus</emphasis>
</taxonomicName>
is Latin for 
<normalizedToken originalValue="“unstained”">&quot;unstained&quot;</normalizedToken>
. The term refers to the fact that the wing has no pterostigma or markings.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="439">This species is characterized by having sinuous cross-vein m-cu and no markings near R2. Furthermore, the species is characterized by male terminalia with a curved sheath of aedeagus and right mesal lobe similar in shape, but longer than left one, and scutellum rounded in lateral view.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="description">
<paragraph pageId="0" pageNumber="439">Description.</paragraph>
<paragraph pageId="0" pageNumber="439">
Wing length 6.22 mm, width 1.74 mm. - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
: General coloration brownish-yellow. Rostrum and palpus dark brown. First segments of antenna yellow, remainding segments dark yellow. Head dark brown. Eye black. Thorax brownish yellow, pleura pale. Scutum without stripes. Halter brownish-yellow. Coxae brownish yellow and legs brown. Abdomen blackish brown. - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
(dorsal view Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19B</figureCitation>
, ventral view Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19A</figureCitation>
): Rostrum bifurcated with short appendices; first palpal segment as long as terminal segment; scape as thick as pedicel; terminal flagellomere as long as penultimate. - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
(lateral view Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19E</figureCitation>
, dorsal view Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19F</figureCitation>
): Short setae on prescutum. Division line of laterotergite almost vertical; anatergite taller than katatergite. Halter with dilated knob. Wing (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19D</figureCitation>
) without markings; veins near fork of bM (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19C</figureCitation>
) visible; M1+2 as long as M1; R2+3+4 longer than R2+3; wing very light on pigmentation, almost translucent. - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Female terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19H</figureCitation>
): Female tergite IX shorter and as wide as tergite X; cercus slightly curved dorsad. - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Amphineurus (Rhamphoneurus) immaculatus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439">19G</figureCitation>
): Male tergite IX slightly darkened along posterior margin, with deep V-shaped median notch; posterolateral lobes of triangular shape with rounded tips. Ventral branch of gonocoxite expanded in distal tip. Dorsal branch of gonocoxite half-length of ventral branch, curved ventrally. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, wider than stem. Clasper of gonostylus with similarly shaped branches: lateral branch rounded, narrow; medial branch wider, oblong. Mesal lobes of gonocoxite symmetrical; both straight with slightly bent pointed tip. Sheath of aedeagus curved halfway and darkened only near tip.
</paragraph>
<caption doi="10.3897/asp.80.e83035.figure19" httpUri="https://binary.pensoft.net/fig/740572" pageId="0" pageNumber="439" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">Figure 19.</emphasis>
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) immaculatus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="immaculatus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) immaculatus</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
head (ventral view); 
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
head (dorsal view); 
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
detail of wing; 
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
wing; 
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
thorax (lateral view); 
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
thorax (dorsal view); 
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
male tergite IX (dorsal view) and remainder of male terminalia (lateral view); 
<emphasis bold="true" pageId="0" pageNumber="439">H</emphasis>
female terminalia (dorsal view). - Abbreviations: 
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
, sheath of aedeagus; 
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
, anatergite; 
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
, anepimeron; 
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
, anepisternum; 
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
, anteropronotum; 
<emphasis bold="true" pageId="0" pageNumber="439">cer</emphasis>
, cercus; 
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
, clasper of gonostylus; 
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
, coxa; 
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
, gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
, halter; 
<emphasis bold="true" pageId="0" pageNumber="439">hyp val</emphasis>
, hypogynial valve; 
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
, knob of halter; 
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
, katepimeron; 
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
, katepisternum; 
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
, katatergite; 
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
, left mesal lobe of gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
; lobe of gonostylus; 
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
, meron; 
<emphasis bold="true" pageId="0" pageNumber="439">ms lobe</emphasis>
, mesal lobe of gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
, metanepisternum; 
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
, metepimeron; 
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
, mediotergite; 
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
, metakatepisternum; 
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
, metanotum; 
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
, posterior basalare; 
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
, paratergite; 
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
, postpronotum; 
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
, prescutum; 
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
, right mesal lobe of gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
, scutum; 
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
, scutellum; 
<emphasis bold="true" pageId="0" pageNumber="439">st</emphasis>
, sternite; 
<emphasis bold="true" pageId="0" pageNumber="439">tg</emphasis>
, tergite; 
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
, male tergite IX.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="remarks">
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="439">
Some specimens of this new species were previously identified by C.P. Alexander as 
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) sanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="sanus" subGenus="R.">A. (R.) sanus</taxonomicName>
Alexander, some as 
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) nullus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="nullus" subGenus="R.">A. (R.) nullus</taxonomicName>
Alexander and some as 
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) nonnullus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="nonnullus" subGenus="R.">A. (R.) nonnullus</taxonomicName>
Alexander. This species resembles 
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) billighami" order="Diptera" pageId="0" pageNumber="439" rank="species" species="billighami" subGenus="R.">A. (R.) billighami</taxonomicName>
sp. nov. but differs mainly in attachment of m-cu, setae concentration in gonocoxite, shape of male tergite IX and right mesal lobe.
</paragraph>
</subSubSection>
</treatment>
</document>