<documentid="BF340091A74A0DBCAA01441BB2535EE8"ID-CLB-Dataset="27575"ID-DOI="10.11646/zootaxa.4554.1.9"ID-GBIF-Dataset="ff7707f0-058f-4f40-8eb8-2b6c9c75a173"ID-ISSN="1175-5326"ID-Zenodo-Dep="2623565"ID-ZooBank="A804E949-109A-468D-B58B-CF7C8BCB3059"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="diego"checkinTime="1554271225461"checkinUser="plazi"docAuthor="Vandenberg, Natalia J. & Hanson, Paul E."docDate="2019"docId="DF18F971FFD70B3DFF4BFD236E61BE8C"docLanguage="en"docName="zootaxa.4554.1.9.pdf"docOrigin="Zootaxa 4554 (1)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Moiradiomus atropos Vandenberg & Hanson, 2019, sp. nov."docType="treatment"docVersion="10"lastPageNumber="277"masterDocId="23218109FFC20B2BFFDCFFF96C54BC27"masterDocTitle="Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)"masterLastPageNumber="285"masterPageNumber="255"pageNumber="276"updateTime="1728643685052"updateUser="ExternalLinkService">
<mods:titleid="B5BE7548957F31E3379E29A6CD5583B1">Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)</mods:title>
<taxonomicNameid="90B133E4FFD70B3EFF4BFD236DCFBED3"authorityName="Vandenberg & Hanson"authorityYear="2019"box="[151,411,730,756]"class="Insecta"family="Coccinellidae"genus="Moiradiomus"higherTaxonomySource="CoL"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="276"phylum="Arthropoda"rank="species"species="atropos"status="sp. nov.">
<figureCitationid="CF8A54E2FFD70B3EFEEEFD076D05BF31"box="[306,337,766,790]"captionStart="FIGURES 25–28"captionStartId="15.[151,264,1975,1998]"captionTargetBox="[151,1436,1466,1918]"captionTargetId="figure@15.[151,1436,1461,1924]"captionTargetPageId="15"captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623578"httpUri="https://zenodo.org/record/2623578/files/figure.png"pageId="21"pageNumber="276">27</figureCitation>
<figureCitationid="CF8A54E2FFD70B3EFE81FD076D2FBF31"box="[349,379,766,790]"captionStart="FIGURES 29–32"captionStartId="16.[151,264,1700,1723]"captionTargetBox="[154,1436,190,1666]"captionTargetId="figure@16.[151,1436,189,1667]"captionTargetPageId="16"captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623580"httpUri="https://zenodo.org/record/2623580/files/figure.png"pageId="21"pageNumber="276">31</figureCitation>
<figureCitationid="CF8A54E2FFD70B3EFE5AFD076DF7BF31"box="[390,419,766,790]"captionStart="FIGURES 40–42"captionStartId="20.[151,264,1957,1980]"captionTargetBox="[151,1436,193,1936]"captionTargetId="figure@20.[151,1436,193,1936]"captionTargetPageId="20"captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii."figureDoi="http://doi.org/10.5281/zenodo.2623584"httpUri="https://zenodo.org/record/2623584/files/figure.png"pageId="21"pageNumber="276">42</figureCitation>
This species is readily distinguished from its congeners by the male genitalia which have the basal lobe distinctly humped in lateral view and apex of parameres obliquely truncate. In addition, the adult form is more slender and apically tapered and the prosternal carinae are weakly convergent, framing an elongate ovoid depression with broadly rounded apex. It is also distinguished by its larval host plant:
<taxonomicNameid="90B133E4FFD70B3EFB84FC4B6D10BFC9"authority="Casimir de Condolle, 1869"authorityName="Casimir de Condolle"authorityYear="1869"class="Magnoliopsida"family="Piperaceae"genus="Piper"kingdom="Plantae"order="Piperales"pageId="21"pageNumber="276"phylum="Tracheophyta"rank="species"species="friedrichsthalii">
<figureCitationid="CF8A54E2FFD70B3EFF43F9146CA6BB21"box="[159,242,1773,1798]"captionStart="FIGURES 3–12"captionStartId="12.[151,264,1784,1807]"captionTargetBox="[151,1433,212,1735]"captionTargetId="figure@12.[151,1436,193,1763]"captionTargetPageId="12"captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon & Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)"figureDoi="http://doi.org/10.5281/zenodo.2623569"httpUri="https://zenodo.org/record/2623569/files/figure.png"pageId="21"pageNumber="276">Fig. 12</figureCitation>
) with carinae of intercoxal process extending nearly to apex, weakly convergent, framing elongate ovoid depression with broadly rounded apex. Suture between abdominal ventrites 1 and 2 distinct, linear; 5
<figureCitationid="CF8A54E2FFD70B3EFE5CF87A6D80BBBB"box="[384,468,1923,1948]"captionStart="FIGURES 29–32"captionStartId="16.[151,264,1700,1723]"captionTargetBox="[154,1436,190,1666]"captionTargetId="figure@16.[151,1436,189,1667]"captionTargetPageId="16"captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623580"httpUri="https://zenodo.org/record/2623580/files/figure.png"pageId="21"pageNumber="276">Fig. 31</figureCitation>
<figureCitationid="CF8A54E2FFD70B3EFCC2F87A6F29BBBB"box="[798,893,1923,1948]"captionStart="FIGURES 29–32"captionStartId="16.[151,264,1700,1723]"captionTargetBox="[154,1436,190,1666]"captionTargetId="figure@16.[151,1436,189,1667]"captionTargetPageId="16"captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623580"httpUri="https://zenodo.org/record/2623580/files/figure.png"pageId="21"pageNumber="276">Fig. 31b</figureCitation>
) weakly, unevenly tapered toward oblique apex, with small thorn-like process at distal most point, distinctly shorter than paramere; in lateral view (
<figureCitationid="CF8A54E2FFD70B3EFB27F85E690DBB98"box="[1275,1369,1959,1984]"captionStart="FIGURES 29–32"captionStartId="16.[151,264,1700,1723]"captionTargetBox="[154,1436,190,1666]"captionTargetId="figure@16.[151,1436,189,1667]"captionTargetPageId="16"captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623580"httpUri="https://zenodo.org/record/2623580/files/figure.png"pageId="21"pageNumber="276">Fig. 31a</figureCitation>
) with strong hump, apex flat; paramere elongate, strongly expanded toward obliquely truncated apex, about 2.5× as long as wide, about 2× longer than longest setae. Penis (
<figureCitationid="CF8A54E2FFD40B3DFD07FF6E6F6EBC97"box="[731,826,151,176]"captionStart="FIGURES 29–32"captionStartId="16.[151,264,1700,1723]"captionTargetBox="[154,1436,190,1666]"captionTargetId="figure@16.[151,1436,189,1667]"captionTargetPageId="16"captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita."figureDoi="http://doi.org/10.5281/zenodo.2623580"httpUri="https://zenodo.org/record/2623580/files/figure.png"pageId="22"pageNumber="277">Fig. 31c</figureCitation>
<taxonomicNameid="90B133E4FFD40B3DFCFDFE616FB1BD88"authorityName="Casimir de Condolle"authorityYear="1869"box="[801,997,406,431]"class="Magnoliopsida"family="Piperaceae"genus="Piper"kingdom="Plantae"order="Piperales"pageId="22"pageNumber="277"phylum="Tracheophyta"rank="species"species="friedrichsthalii">
<figureCitationid="CF8A54E2FFD40B3DFC28FE6F6812BD88"box="[1012,1094,406,431]"captionStart="FIGURES 40–42"captionStartId="20.[151,264,1957,1980]"captionTargetBox="[151,1436,193,1936]"captionTargetId="figure@20.[151,1436,193,1936]"captionTargetPageId="20"captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii."figureDoi="http://doi.org/10.5281/zenodo.2623584"httpUri="https://zenodo.org/record/2623584/files/figure.png"pageId="22"pageNumber="277">Fig. 42</figureCitation>
Atropos, proper noun in apposition. Ancient Greek Ἄτρoπoς (Atropos), literally meaning inflexible or not turning, from the Greek a-, (not) + tropos, (to turn). In Greek mythology, one of the three Fates, or Moirae; the cutter of the thread of life, depicted as an old woman.
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