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Genus 
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, 420; 1866: 279.—Crotch 1874: 158; 
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<emphasis id="B91A2C66149D46F2BE37327FFBFFFB81" box="[895,1074,1026,1050]" italics="true" pageId="197" pageNumber="198">Oenopia cinctella</emphasis>
Mulsant
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, by subsequent designation of 
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c: 134. (
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wrongly designated 
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<emphasis id="B91A2C66149D46F2BE06325DFC68FBA3" box="[846,933,1057,1080]" italics="true" pageId="197" pageNumber="198">Oenopia</emphasis>
(
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) 
<emphasis id="B91A2C66149D46F2BEA1325DFBE9FBA2" box="[1001,1060,1057,1081]" italics="true" pageId="197" pageNumber="198">kirbyi</emphasis>
<bibRefCitation id="EFFF8D85149D46F2B962325DFB70FBA2" author="Mulsant, E." box="[1066,1213,1057,1081]" pageId="197" pageNumber="198" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
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<emphasis id="B91A2C66149D46F2BDDF323CFED3FBC3" box="[151,286,1088,1112]" italics="true" pageId="197" pageNumber="198">Synharmonia</emphasis>
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[as 
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<emphasis id="B91A2C66149D46F2BF71323CFD6AFBC3" box="[569,679,1088,1112]" italics="true" pageId="197" pageNumber="198">Coccinella</emphasis>
(
<emphasis id="B91A2C66149D46F2BFFE323CFCF0FBC3" box="[694,829,1088,1112]" italics="true" pageId="197" pageNumber="198">Synharmonia</emphasis>
)
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]. 
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species: 
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<emphasis id="B91A2C66149D46F2BEA3323CFB1FFBC3" box="[1003,1234,1088,1112]" italics="true" pageId="197" pageNumber="198">Coccinella conglobata</emphasis>
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<treatmentCitation id="0ACFD665149D46F2BDDF3202FE21FB0E" author="Timberlake, P. H." box="[151,492,1149,1173]" page="28" pageId="197" pageNumber="198" year="1943">
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<emphasis id="B91A2C66149D46F2BDDF3202FEC8FB0E" box="[151,261,1150,1173]" italics="true" pageId="197" pageNumber="198">Protocaria</emphasis>
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. 
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species: 
<taxonomicName id="4C6E8BF7149D46F2BFC03202FBC1FB0E" authority="Timberlake, 1943" authorityName="Timberlake" authorityYear="1943" box="[648,1036,1149,1173]" class="Insecta" family="Coccinellidae" genus="Protocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="scalaris">
<emphasis id="B91A2C66149D46F2BFC03202FC83FB0E" box="[648,846,1149,1173]" italics="true" pageId="197" pageNumber="198">Protocaria scalaris</emphasis>
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, by original designation. Synonymized by 
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</paragraph>
<paragraph id="8BD1F074149D46F2BDDF32C7FCFFFB6A" blockId="197.[151,1436,995,1388]" pageId="197" pageNumber="198">
<treatmentCitationGroup id="AB7ED75A149D46F2BDDF32C7FCFFFB6A" pageId="197" pageNumber="198">
<treatmentCitation id="0ACFD665149D46F2BDDF32C7FE23FB49" author="Timberlake, P. H." box="[151,494,1210,1234]" page="39" pageId="197" pageNumber="198" year="1943">
<taxonomicName id="4C6E8BF7149D46F2BDDF32C7FE23FB49" authority="Timberlake, 1943: 39" authorityName="Timberlake" authorityPageNumber="39" authorityYear="1943" box="[151,494,1210,1234]" class="Insecta" family="Coccinellidae" genus="Gyrocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66149D46F2BDDF32C7FECDFB49" box="[151,256,1211,1234]" italics="true" pageId="197" pageNumber="198">Gyrocaria</emphasis>
<bibRefCitation id="EFFF8D85149D46F2BC4332C6FE23FB49" author="Timberlake, P. H." box="[267,494,1210,1234]" pageId="197" pageNumber="198" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake, 1943: 39</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.—Bielawski 1964b: 83. 
<typeStatus id="54D54ED6149D46F2BFB232C7FCE1FB48" box="[762,812,1211,1235]" pageId="197" pageNumber="198">Type</typeStatus>
species: 
<taxonomicName id="4C6E8BF7149D46F2BEDB32C6FAD7FB49" authority="Blackburn, 1892" authorityName="Blackburn" authorityYear="1892" box="[915,1306,1210,1234]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="B91A2C66149D46F2BEDB32C6FB92FB49" box="[915,1119,1210,1234]" italics="true" pageId="197" pageNumber="198">Coelophora guttata</emphasis>
Blackburn, 1892
</taxonomicName>
, by original designation. Synonymized by 
<treatmentCitation id="0ACFD665149D46F2BCB232A5FCE0FB6A" author="Iablokoff-Khnzorian, S. M." box="[506,813,1241,1265]" page="69" pageId="197" pageNumber="198" year="1979">
<bibRefCitation id="EFFF8D85149D46F2BCB232A5FCE0FB6A" author="Iablokoff-Khnzorian, S. M." box="[506,813,1241,1265]" pageId="197" pageNumber="198" pagination="37 - 75" refId="ref114003" refString="Iablokoff-Khnzorian, S. M. (1979) Genera der Palaarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75." type="journal article" year="1979">Iablokoff-Khnzorian 1979: 69</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F074149D46F2BDDF3284FCEFFAB5" blockId="197.[151,1436,995,1388]" pageId="197" pageNumber="198">
<emphasis id="B91A2C66149D46F2BDDF3284FE89FA8B" box="[151,324,1272,1296]" italics="true" pageId="197" pageNumber="198">Pseudoharmonia</emphasis>
Savojskaja, 1963: 37.— 
<bibRefCitation id="EFFF8D85149D46F2BF743284FCD1FA94" author="Kuznetsov, V. N." box="[572,796,1271,1296]" pageId="197" pageNumber="198" refId="ref116665" refString="Kuznetsov, V. N. (1997) Lady Beetles of the Russian Far East. Memoir No. 1. Centre for Systematic Entomology, The Sandhill Crane Press, Gainesville, Florida, 248 pp." type="book" year="1997">Kuznetsov 1997: 186</bibRefCitation>
. 
<typeStatus id="54D54ED6149D46F2BE6D3284FC9AFA8B" box="[805,855,1272,1296]" pageId="197" pageNumber="198">Type</typeStatus>
species: 
<taxonomicName id="4C6E8BF7149D46F2BEFD3284FAB5FA94" authority="Savojskaja, 1963" authorityName="Savojskaja" authorityYear="1963" box="[949,1400,1271,1296]" class="Insecta" family="Coccinellidae" genus="Pseudoharmonia" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="B91A2C66149D46F2BEFD3284FB0FFA94" box="[949,1218,1272,1296]" italics="true" pageId="197" pageNumber="198">Pseudoharmonia montana</emphasis>
Savojskaja, 1963
</taxonomicName>
, by monotypy. Synonymized by 
<bibRefCitation id="EFFF8D85149D46F2BCA2336AFCD3FAB5" author="Iablokoff-Khnzorian, S. M." box="[490,798,1302,1326]" pageId="197" pageNumber="198" pagination="37 - 75" refId="ref114003" refString="Iablokoff-Khnzorian, S. M. (1979) Genera der Palaarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75." type="journal article" year="1979">Iablokoff-Khnzorian 1979: 69</bibRefCitation>
.
</paragraph>
<paragraph id="8BD1F074149D46F2BDDF3349FD50FAF0" blockId="197.[151,1436,995,1388]" pageId="197" pageNumber="198">
<treatmentCitationGroup id="AB7ED75A149D46F2BDDF3349FD50FAF0" pageId="197" pageNumber="198">
<treatmentCitation id="0ACFD665149D46F2BDDF3349FE20FAD6" author="Hoang, D. N." box="[151,493,1333,1357]" page="10" pageId="197" pageNumber="198" year="1982">
<taxonomicName id="4C6E8BF7149D46F2BDDF3349FE20FAD6" authority="Hoang, 1982: 10" authorityName="Hoang" authorityPageNumber="10" authorityYear="1982" box="[151,493,1333,1357]" class="Insecta" family="Coccinellidae" genus="Paramulsantina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C66149D46F2BDDF3349FEF7FAD6" box="[151,314,1333,1357]" italics="true" pageId="197" pageNumber="198">Paramulsantina</emphasis>
<bibRefCitation id="EFFF8D85149D46F2BC093349FE20FAD6" author="Hoang, D. N." box="[321,493,1333,1357]" pageId="197" pageNumber="198" pagination="10 - 15" refId="ref113469" refString="Hoang, D. N. (1982) A new genus and some new species of subfamily Coccinellinae, (Coleoptera: Coccinellidae) from Vietnam. Tap chi Sinh Vat Hoc, 4 (3), 10 - 15." type="journal article" year="1982">Hoang, 1982: 10</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="54D54ED6149D46F2BCBF3349FDE4FAD6" box="[503,553,1333,1357]" pageId="197" pageNumber="198">Type</typeStatus>
species: 
<taxonomicName id="4C6E8BF7149D46F2BFCE3349FBDDFAD6" authority="Hoang, 1982" authorityName="Hoang" authorityYear="1982" box="[646,1040,1333,1357]" class="Insecta" family="Coccinellidae" genus="Paramulsantina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B91A2C66149D46F2BFCE3349FC49FAD6" box="[646,900,1333,1357]" italics="true" pageId="197" pageNumber="198">Paramulsantina gratiosa</emphasis>
<bibRefCitation id="EFFF8D85149D46F2BEC33349FBDDFAD6" author="Hoang, D. N." box="[907,1040,1333,1357]" pageId="197" pageNumber="198" pagination="10 - 15" refId="ref113469" refString="Hoang, D. N. (1982) A new genus and some new species of subfamily Coccinellinae, (Coleoptera: Coccinellidae) from Vietnam. Tap chi Sinh Vat Hoc, 4 (3), 10 - 15." type="journal article" year="1982">Hoang, 1982</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by 
<treatmentCitation id="0ACFD665149D46F2BDAF3328FD54FAF0" author="Jadwiszczak, A. &amp; Pokojowczyk, A." box="[231,665,1363,1388]" page="54" pageId="197" pageNumber="198" year="1990">
<bibRefCitation id="EFFF8D85149D46F2BDAF3328FD54FAF0" author="Jadwiszczak, A. &amp; Pokojowczyk, A." box="[231,665,1363,1388]" pageId="197" pageNumber="198" pagination="54 - 55" refId="ref114505" refString="Jadwiszczak, A. &amp; Pokojowczyk, A. (1990) Paramulsantina Hoang (Coleoptera: Coccinellidae), a new synonym of the genus Oenopia Mulsant. Coccinella, 2 (2), 54 - 55." type="journal article" year="1990">Jadwiszczak &amp; Pokojowczyk 1990: 54–55</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF149D46F2BDDF33E5FCCDF887" pageId="197" pageNumber="198" type="diagnosis">
<paragraph id="8BD1F074149D46F2BDDF33E5FCCDF887" blockId="197.[151,1437,1433,1964]" pageId="197" pageNumber="198">
<emphasis id="B91A2C66149D46F2BDDF33E5FEDFFA28" bold="true" box="[151,274,1433,1459]" pageId="197" pageNumber="198">Diagnosis.</emphasis>
Body outline variable from rounded to broad oval to distinctly elongate, dorsum convex to flattened, glabrous. Head with anterior clypeal margin straight between lateral projections, inner margins of compound eyes almost straight. Antenna 11-segmented, slightly shorter than head capsule with the apical three antennomeres forming a compact club. Scutellar shield broader than long, triangular. Prothoracic hypomeron with anterior inner corners shallowly foveate or not. Prosternal carinae present. Anterior margin of mesoventrite distinctly to very shallowly emarginate. Middle and hind tibiae with a pair of spurs. Elytral epipleura foveolate to receive the tips of mid-and / or hind femora. Abdominal postcoxal line incomplete, with an oblique dividing lateral line. Spermatheca characteristic with a highly elongate, apically swollen ramus and a distinct nodulus, a short, flexible sperm duct and a distinct, sclerotized, diagnostically shaped infundibulum (with the sole exception of 
<taxonomicName id="4C6E8BF7149D46F2B91D30C6FB28F94F" box="[1109,1253,1722,1748]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="197" pageNumber="198" phylum="Arthropoda" rank="species" species="signatella">
<emphasis id="B91A2C66149D46F2B91D30C6FB28F94F" box="[1109,1253,1722,1748]" italics="true" pageId="197" pageNumber="198">O. signatella</emphasis>
</taxonomicName>
). Male genitalia with penis guide apically upturned and medially notched or more or less subtruncate, parameres arched. Penis with a well-developed capsule and variously modified apex.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF149D46F2BD8F3159FBACF8FF" pageId="197" pageNumber="198" type="distribution">
<paragraph id="8BD1F074149D46F2BD8F3159FBACF8FF" blockId="197.[151,1437,1433,1964]" pageId="197" pageNumber="198">
<emphasis id="B91A2C66149D46F2BD8F3159FEACF8A4" bold="true" box="[199,353,1829,1855]" pageId="197" pageNumber="198">Distribution.</emphasis>
A moderately large genus, with ~30 species distributed in Europe, Asiatic 
<collectingCountry id="F379B0E4149D46F2B9E6315AFB31F8DB" box="[1198,1276,1830,1856]" name="Russia" pageId="197" pageNumber="198">Russia</collectingCountry>
, Africa, 
<collectingCountry id="F379B0E4149D46F2B814315AFA55F8DB" box="[1372,1432,1830,1856]" name="India" pageId="197" pageNumber="198">India</collectingCountry>
, 
<collectingCountry id="F379B0E4149D46F2BDDF3136FE9FF8FF" box="[151,338,1866,1892]" name="Sri Lanka China" pageId="197" pageNumber="198">Sri Lanka China</collectingCountry>
, 
<collectingCountry id="F379B0E4149D46F2BC153136FE6DF8FF" box="[349,416,1866,1892]" name="Japan" pageId="197" pageNumber="198">Japan</collectingCountry>
, 
<collectingCountry id="F379B0E4149D46F2BCE23136FDE6F8FF" box="[426,555,1866,1892]" name="Philippines" pageId="197" pageNumber="198">Philippines</collectingCountry>
, New 
<collectingCountry id="F379B0E4149D46F2BF3A3136FD0EF8FF" box="[626,707,1866,1892]" name="Guinea-Bissau" pageId="197" pageNumber="198">Guinea</collectingCountry>
and 
<collectingCountry id="F379B0E4149D46F2BFB03136FC92F8FF" box="[760,863,1866,1892]" name="Australia" pageId="197" pageNumber="198">Australia</collectingCountry>
(
<bibRefCitation id="EFFF8D85149D46F2BE253137FB9BF8FF" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[877,1110,1866,1894]" pageId="197" pageNumber="198" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński 
<emphasis id="B91A2C66149D46F2BE923137FBC6F8FF" box="[986,1035,1866,1892]" italics="true" pageId="197" pageNumber="198">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF149D46F2BD8F3111FD7FF837" pageId="197" pageNumber="198" type="discussion">
<paragraph id="8BD1F074149D46F2BD8F3111FD7FF837" blockId="197.[151,1437,1433,1964]" pageId="197" pageNumber="198">
<emphasis id="B91A2C66149D46F2BD8F3111FEF0F81C" bold="true" box="[199,317,1901,1927]" pageId="197" pageNumber="198">Affinities.</emphasis>
<bibRefCitation id="EFFF8D85149D46F2BC0A3112FDA3F813" author="Tomaszewska, W. &amp; Escalona, H. E. &amp; Hartley, D. &amp; Li, J. &amp; Wang, X. &amp; Li, H. S. &amp; Pang, H. &amp; Slipinski, A. &amp; Zwick, A." box="[322,622,1902,1928]" pageId="197" pageNumber="198" pagination="611 - 631" refId="ref123461" refString="Tomaszewska, W., Escalona, H. E., Hartley, D., Li, J., Wang, X., Li, H. S., Pang, H., Slipinski, A. &amp; Zwick, A. (2021) Phylogeny of true ladybird beetles (Coccinellidae: Coccinellini) reveals pervasive convergent evolution and a rapid Cenozoic radiation. Systematic Entomology, 46, 611 - 631. https: // doi. org / 10.1111 / syen. 12479" type="journal article" year="2021">
Tomaszewska 
<emphasis id="B91A2C66149D46F2BCAD3113FDD0F813" box="[485,541,1902,1928]" italics="true" pageId="197" pageNumber="198">et al.</emphasis>
(2021)
</bibRefCitation>
included it in the 
<emphasis id="B91A2C66149D46F2BE7D3112FC75F813" box="[821,952,1902,1928]" italics="true" pageId="197" pageNumber="198">Coccinella-</emphasis>
group of genera and found it to be a distinct genus strongly supported in molecular analyses.
</paragraph>
</subSubSection>
<caption id="DF11A0FC149E46F1BDDF31EBFB2DF848" ID-DOI="http://doi.org/10.5281/zenodo.8262003" ID-Zenodo-Dep="8262003" httpUri="https://zenodo.org/record/8262003/files/figure.png" pageId="198" pageNumber="199" startId="198.[151,250,1943,1967]" targetBox="[167,1429,181,1916]" targetPageId="198" targetType="figure">
<paragraph id="8BD1F074149E46F1BDDF31EBFB2DF848" blockId="198.[151,1436,1943,2004]" pageId="198" pageNumber="199">
<emphasis id="B91A2C66149E46F1BDDF31EBFEE3F834" bold="true" box="[151,302,1943,1967]" pageId="198" pageNumber="199">FIGURE 140.</emphasis>
<taxonomicName id="4C6E8BF7149E46F1BC7E31E4FDA3F82B" authority="(Swartz)" baseAuthorityName="Swartz" box="[310,622,1944,1968]" class="Insecta" family="Coccinellidae" genus="Microcaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="198" pageNumber="199" phylum="Arthropoda" rank="species" species="pupillata">
<emphasis id="B91A2C66149E46F1BC7E31E4FDC1F82B" box="[310,524,1944,1968]" italics="true" pageId="198" pageNumber="199">Microcaria pupillata</emphasis>
(Swartz)
</taxonomicName>
: a, b. adult, dorsal view of nominate form; c. dorsal view, variant (BMNH); d. abdomen, male; e–h. male genitalia: e. tegmen, lateral view; f. tegmen, ventral view; g. penis; h. penis apex.
</paragraph>
</caption>
<subSubSection id="C374A3FF149F46F0BD8F36EBFDCEFF62" pageId="199" pageNumber="200" type="taxon_list">
<paragraph id="8BD1F074149F46F0BD8F36EBFDCEFF62" blockId="199.[151,1436,150,249]" pageId="199" pageNumber="200">
<emphasis id="B91A2C66149F46F0BD8F36EBFE5FFF2B" bold="true" box="[199,402,150,177]" pageId="199" pageNumber="200">Included species.</emphasis>
It is one of the largest genera of 
<taxonomicName id="4C6E8BF7149F46F0BE4E36EBFC42FF2A" box="[774,911,151,177]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="tribe" tribe="Coccinellini">Coccinellini</taxonomicName>
of the Indian region and at present, 15 species are known, most of them from the northeastern region. Only 
<taxonomicName id="4C6E8BF7149F46F0BE0C36C7FBE3FF4E" authority="(Mulsant)" baseAuthorityName="Mulsant" box="[836,1070,187,213]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="species" species="guerini">
<emphasis id="B91A2C66149F46F0BE0C36C7FC75FF4F" box="[836,952,187,213]" italics="true" pageId="199" pageNumber="200">O. guerini</emphasis>
(Mulsant)
</taxonomicName>
(= 
<taxonomicName id="4C6E8BF7149F46F0B90636C7FABDFF4E" authority="(Sicard))" authorityName=": Iablokoff-Khnzorian" authorityYear="1979" baseAuthorityName="Sicard" box="[1102,1392,187,213]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="species" species="walteri">
<emphasis id="B91A2C66149F46F0B90636C7FAC8FF4E" box="[1102,1285,187,213]" italics="true" pageId="199" pageNumber="200">Oenopia walteri</emphasis>
(Sicard))
</taxonomicName>
has been recorded from South 
<collectingCountry id="F379B0E4149F46F0BC8A36A3FE32FF62" box="[450,511,223,249]" name="India" pageId="199" pageNumber="200">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>