346 lines
52 KiB
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346 lines
52 KiB
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<document id="AF6DB8560B6E4ACA290A0CC3C132D876" ID-DOI="10.11646/zootaxa.3964.3.3" ID-GBIF-Dataset="d68cb8e7-9803-4052-92dc-057b53dd531a" ID-ISSN="1175-5326" ID-Zenodo-Dep="243903" ID-ZooBank="823F6194-5EAE-4EE5-93A2-FFFDE1EA8A1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461075227785" checkinUser="plazi" docAuthor="Zhang, Chao & Zhang, Feng" docDate="2015" docId="03EF5248FFC2FFEE8FABF8D4FD64FBA4" docLanguage="en" docName="zt03964p351.pdf" docOrigin="Zootaxa 3964 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paramaracandus dolabratus Zhang & Zhang, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="346" masterDocId="FFD62A30FFC4FFE58F3CFFBDFFB4FFA1" masterDocTitle="The assamiids harvestmen (Opiliones: Laniatores: Assamiidae) from Champasak Province, Laos" masterLastPageNumber="351" masterPageNumber="335" pageNumber="341" updateTime="1698586245962" updateUser="plazi">
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<mods:title id="39ECCA9CC6B94F2C4BE527D51DAB075A">The assamiids harvestmen (Opiliones: Laniatores: Assamiidae) from Champasak Province, Laos</mods:title>
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<mods:namePart id="61936D0CCCC893A61D3ACD0F9855E3C1">Zhang, Chao</mods:namePart>
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<mods:namePart id="44E88A703A3FEE4047C538E00FD18510">Zhang, Feng</mods:namePart>
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<mods:typeOfResource id="63C64356A5C3E564AC7E108B8D023DE3">text</mods:typeOfResource>
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<mods:title id="D3BC167522B25B6E1EEAEC4CF0845B6E">Zootaxa</mods:title>
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<mods:part id="EA92348A697B24AD99EDA41360512B61">
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<mods:date id="CD4E756EE6E8FD829E9DA7650B2CF1A1">2015</mods:date>
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<mods:number id="B038BC817374E3903ECE20EB8CB7EC99">3964</mods:number>
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<mods:identifier id="2FF5ADE726E25366D0DE3E18892B2C4B" type="DOI">10.11646/zootaxa.3964.3.3</mods:identifier>
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<treatment id="03EF5248FFC2FFEE8FABF8D4FD64FBA4" ID-DOI="http://doi.org/10.5281/zenodo.5615584" ID-GBIF-Taxon="119617599" ID-Zenodo-Dep="5615584" LSID="urn:lsid:plazi:treatment:03EF5248FFC2FFEE8FABF8D4FD64FBA4" httpUri="http://treatment.plazi.org/id/03EF5248FFC2FFEE8FABF8D4FD64FBA4" lastPageId="11" lastPageNumber="346" pageId="6" pageNumber="341">
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<subSubSection id="C35CB0D5FFC2FFE28FABF8D4FEB4FF74" lastPageId="7" lastPageNumber="342" pageId="6" pageNumber="341" type="nomenclature">
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<paragraph id="8BF9E35EFFC2FFE38FABF8D4FDE0F822" blockId="6.[151,596,1897,1956]" box="[151,596,1897,1923]" pageId="6" pageNumber="341">
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<heading id="D0B15432FFC2FFE38FABF8D4FDE0F822" bold="true" box="[151,596,1897,1923]" fontSize="11" level="1" pageId="6" pageNumber="341" reason="1">
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<emphasis id="B9323F4CFFC2FFE38FABF8D4FDE0F822" bold="true" box="[151,596,1897,1923]" pageId="6" pageNumber="341">
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<taxonomicName id="4C4698DDFFC2FFE38FABF8D4FE5BF822" ID-CoL="7XWMW" box="[151,495,1897,1923]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
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<emphasis id="B9323F4CFFC2FFE38FABF8D4FE5BF822" bold="true" box="[151,495,1897,1923]" italics="true" pageId="6" pageNumber="341">Paramaracandus dolabratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2018237FFC2FFE38ECAF8D4FDE0F822" box="[502,596,1897,1923]" pageId="6" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF9E35EFFC2FFE38FABF836FE9AF805" blockId="6.[151,596,1897,1956]" box="[151,302,1931,1956]" pageId="6" pageNumber="341">
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(
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<figureCitation id="137DFFDBFFC2FFE38FA3F836FE92F805" box="[159,294,1931,1956]" captionStart-0="FIGURES 26 – 31" captionStart-1="FIGURES 32 – 37" captionStart-2="FIGURES 38 – 46" captionStartId-0="8.[151,264,1952,1975]" captionStartId-1="9.[151,264,1834,1857]" captionStartId-2="10.[151,264,1906,1929]" captionTargetBox-0="[246,1342,188,1906]" captionTargetBox-1="[208,1366,203,1742]" captionTargetBox-2="[263,1329,239,1843]" captionTargetId-0="figure@8.[181,1368,157,1937]" captionTargetId-1="figure@9.[200,1387,193,1782]" captionTargetId-2="figure@10.[200,1387,193,1885]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." captionText-1="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." captionText-2="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri-0="https://zenodo.org/record/243908/files/figure.png" httpUri-1="https://zenodo.org/record/243909/files/figure.png" httpUri-2="https://zenodo.org/record/243910/files/figure.png" pageId="6" pageNumber="341">Figs. 26–46</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF9E35EFFC2FFE38FABF869FA28F84D" blockId="6.[151,1436,2004,2029]" box="[151,1436,2004,2029]" pageId="6" pageNumber="341">
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<emphasis id="B9323F4CFFC2FFE38FABF869FF57F84C" bold="true" box="[151,227,2004,2029]" pageId="6" pageNumber="341">
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<typeStatus id="54FD5DFCFFC2FFE38FABF869FF6BF84C" box="[151,223,2004,2029]" pageId="6" pageNumber="341">Types</typeStatus>
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.
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</emphasis>
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<typeStatus id="54FD5DFCFFC2FFE38FD0F869FEE0F84C" box="[236,340,2004,2029]" pageId="6" pageNumber="341" type="holotype">Holotype</typeStatus>
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male,
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<collectingCountry id="F351A3CEFFC2FFE38E9FF868FE6AF84D" box="[419,478,2005,2028]" name="Laos" pageId="6" pageNumber="341">Laos</collectingCountry>
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: Champasak, Phou Salao [
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<geoCoordinate id="EE728599FFC2FFE38C31F868FCCFF84C" box="[781,891,2004,2029]" direction="north" orientation="latitude" pageId="6" pageNumber="341" precision="925" value="15.083333">N 15°05´</geoCoordinate>
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,
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<geoCoordinate id="EE728599FFC2FFE38CB4F868FBB5F84C" box="[904,1025,2004,2029]" direction="east" orientation="longitude" pageId="6" pageNumber="341" precision="925" value="105.8">E 105°48´</geoCoordinate>
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], alt.
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<quantity id="4CBE4EBBFFC2FFE38B7DF869FB30F84C" box="[1089,1156,2004,2029]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="341" unit="m" value="150.0">150m</quantity>
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,
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<date id="FFF8C59EFFC2FFE38BADF869FAC2F84D" box="[1169,1398,2004,2029]" pageId="6" pageNumber="341" value="2011-11-15">November 15, 2011</date>
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, C.
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</paragraph>
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<paragraph id="8BF9E35EFFC3FFE28FABFF2AFEB4FF74" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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Zhang leg. (MHBU-Opi-11ZC1129).
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<typeStatus id="54FD5DFCFFC3FFE28D6BFF25FD08FF11" box="[599,700,152,176]" pageId="7" pageNumber="342" type="paratype">Paratype</typeStatus>
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, one male (MHBU-Opi-11ZC1130), same collecting data as
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<typeStatus id="54FD5DFCFFC3FFE28FABFF01FF49FF74" box="[151,253,188,213]" pageId="7" pageNumber="342" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CB0D5FFC3FFE28FFBFF62FB9CFEE1" pageId="7" pageNumber="342" type="diagnosis">
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<paragraph id="8BF9E35EFFC3FFE28FFBFF62FB9CFEE1" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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<emphasis id="B9323F4CFFC3FFE28FFBFF62FEF5FF59" bold="true" box="[199,321,223,248]" pageId="7" pageNumber="342">Diagnosis.</emphasis>
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Coxa IV prolaterally with many conspicuous hair-tipped granules. Femur of pedipalpus with a setiferous tubercle on the medial distal side. Ventral plate of penis somewhat triangular, and the tip of ventral plate nearly axe-shaped. Stylus long, cylindrical and gradually thickened towards base.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CB0D5FFC3FFE28FFBFEF1FCEFFE29" pageId="7" pageNumber="342" type="etymology">
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<paragraph id="8BF9E35EFFC3FFE28FFBFEF1FCEFFE29" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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<emphasis id="B9323F4CFFC3FFE28FFBFEF1FEFAFEC4" bold="true" box="[199,334,332,357]" pageId="7" pageNumber="342">Etymology.</emphasis>
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The specific name is derived from the Latin word “
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<taxonomicName id="4C4698DDFFC3FFE28C91FEF1FB92FEC5" box="[941,1062,332,356]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="342" phylum="Arthropoda" rank="species" species="dolabratus">
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<emphasis id="B9323F4CFFC3FFE28C91FEF1FB92FEC5" box="[941,1062,332,356]" italics="true" pageId="7" pageNumber="342">dolabratus</emphasis>
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</taxonomicName>
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” meaning “shaped like an axe”, referring to the shape of the tip of the ventral plate in the penis.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CB0D5FFC3FFEE8FFBFE29FBB1FDF4" lastPageId="11" lastPageNumber="346" pageId="7" pageNumber="342" type="description">
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<paragraph id="8BF9E35EFFC3FFE28FFBFE29FAEAFD9C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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<emphasis id="B9323F4CFFC3FFE28FFBFE29FEEDFE0C" bold="true" box="[199,345,404,429]" pageId="7" pageNumber="342">Description.</emphasis>
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Male habitus as in
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<figureCitation id="137DFFDBFFC3FFE28D70FE29FD6EFE0C" box="[588,730,404,429]" captionStart="FIGURES 26 – 31" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[246,1342,188,1906]" captionTargetId="figure@8.[181,1368,157,1937]" captionTargetPageId="8" captionText="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="7" pageNumber="342">Figs. 26–27</figureCitation>
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. Coloration: entire body yellowish brown, with dark brown patches on the dorsum; median area of carapace with dark brown reticulations; both lateral ridges of the carapace and scutum with blackish brown stripes; free tergites I–III each with a blackish brown band; coxae with dark brown reticulations; free sternites with transverse band of blackish brown; chelicerae and pedipalps reticulated above; trochanters of all legs pale yellow, femora, patellae, tibiae and metatarsi with black reticulations, tarsi lighter.
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</paragraph>
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<paragraph id="8BF9E35EFFC3FFE28FFBFDF5FBA3FC96" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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Dorsum (
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<figureCitation id="137DFFDBFFC3FFE28E0DFDFAFE32FDC1" box="[305,390,583,608]" captionStart="FIGURES 26 – 31" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[246,1342,188,1906]" captionTargetId="figure@8.[181,1368,157,1937]" captionTargetPageId="8" captionText="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="7" pageNumber="342">Fig. 27</figureCitation>
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). Dorsal scutum trapezoid in shape, widest portion of body at scutal area V. Anterior margin of carapace with two spines at the lateral portion and a single median spine: the lateral spines of similar length, the median a little smaller; front margin of the ocularium dorsally with a median spine, a transverse row of seven small tubercles on each side of this median spine. Ocularium small, oval, with a pair of conspicuous spines and two tubercles. Opisthosomal region of scutum with five areas, area II wider than others. Area I without a median groove. Area I with two median hair-tipped granules, areas II–III each with two median spines, the rear pair larger. Free tergites unarmed, with a transverse row of tubercles, similar to areas IV–V.
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</paragraph>
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<paragraph id="8BF9E35EFFC3FFE28FFBFCF8FCDCFC4C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">Venter. Surface of all coxae tuberculated. Coxa I with a row of four tubercles prolaterally, and two rows of tubercles on the surface. Coxa II with a row of marginal tubercles on the prolateral surface, and disto-dorsally with an enlarged tubercle. Coxa III with prolateral and retrolateral rows of tubercles. Coxa IV larger than others, prolaterally with many scattered tubercles, and disto-retrolaterally with two teeth. Genital operculum with a few hair-tipped granules. Free sternites smooth. Spiracles concealed.</paragraph>
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<paragraph id="8BF9E35EFFC3FFE28FFBFC4AFE6BFBDC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
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Chelicera (
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<figureCitation id="137DFFDBFFC3FFE28E7EFC4AFE79FBB1" box="[322,461,1015,1040]" captionStart="FIGURES 26 – 31" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[246,1342,188,1906]" captionTargetId="figure@8.[181,1368,157,1937]" captionTargetPageId="8" captionText="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="7" pageNumber="342">Figs. 28–30</figureCitation>
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). Basichelicerite with distinct bulla, no prominent armaments except a few small hairtipped tubercles. Cheliceral hand unarmed, with sparse hairs only. Fingers relatively short, inner edges toothed as illustrated (
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<figureCitation id="137DFFDBFFC3FFE28E24FB82FEDAFBF9" box="[280,366,1087,1112]" captionStart="FIGURES 26 – 31" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[246,1342,188,1906]" captionTargetId="figure@8.[181,1368,157,1937]" captionTargetPageId="8" captionText="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="7" pageNumber="342">Fig. 30</figureCitation>
|
|||
|
|
): moveable finger with 13 teeth, most square except for the proximal and distal ones crested; fixed finger with seven teeth.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBFB3AFC3BFA61" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
|
|||
|
|
Pedipalpus (
|
|||
|
|
<figureCitation id="137DFFDBFFC3FFE28E6DFB3AFE6EFB01" box="[337,474,1159,1184]" captionStart-0="FIGURES 26 – 31" captionStart-1="FIGURES 32 – 37" captionStartId-0="8.[151,264,1952,1975]" captionStartId-1="9.[151,264,1834,1857]" captionTargetBox-0="[246,1342,188,1906]" captionTargetBox-1="[208,1366,203,1742]" captionTargetId-0="figure@8.[181,1368,157,1937]" captionTargetId-1="figure@9.[200,1387,193,1782]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." captionText-1="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri-0="https://zenodo.org/record/243908/files/figure.png" httpUri-1="https://zenodo.org/record/243909/files/figure.png" pageId="7" pageNumber="342">Figs. 31–33</figureCitation>
|
|||
|
|
). Coxa dorsally with two setiferous tubercles. Trochanter dorsally with two setiferous tubercles, and ventrally with one small and two distal setiferous tubercles. Femur compressed laterally, widest at the middle of its length, ventrally with a row of 20 homogeneous setiferous tubercles; dorsally with many low conical tubercles along the entire length; on the medial distal side with a setiferous tubercle. Patella distoventromesally with a conspicuous bifurcated and disto-ventroectally with a non-bifurcated setiferous tubercle. Tibia ventromesally with two enlarged setiferous tubercles and three small ones, and ventroectally with a fairly enlarged setiferous tubercle and two small ones on its distal side. Tarsus ventromesally with a row of two slightly enlarged and four small setiferous tubercles; and ventroectally expanded, with two slightly enlarged and five small setiferous tubercles. Tarsal claw slightly curved, shorter than tarsus.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBFA70FCEDF9F1" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
|
|||
|
|
Legs. Short. Trochanters I–IV with hair-tipped granules on the ventral surface, the granules of trochanter I conspicuous. All femora with hair-tipped granules, femora III and IV curved. Tarsi III–IV without a pseudonychium, but with two bare claws. Tarsal formula (I–IV): 6/11–
|
|||
|
|
<date id="FFF8C59EFFC3FFE28C97F9A9FC73F98C" box="[939,967,1556,1581]" pageId="7" pageNumber="342" value="2012-08-07">12</date>
|
|||
|
|
/7/8. Distitarsus I two-jointed and II threejointed. The remaining leg-segments with hair-tipped granules.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBF9E1FD8EF889" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
|
|||
|
|
Penis (
|
|||
|
|
<figureCitation id="137DFFDBFFC3FFE28E2BF9E1FE11F9D4" box="[279,421,1628,1653]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="7" pageNumber="342">Figs. 38–46</figureCitation>
|
|||
|
|
). Shaft slender, gradually distended from base to apical portion (pars distalis), slightly undulate (best seen from lateral). Ventral plate somewhat triangular, arched ventrally and concave dorsally, distal end axe-shaped (seen from dorsal and ventral). Glans partly sunken into the truncus. Capsula externa extremely compressed after eversion (
|
|||
|
|
<figureCitation id="137DFFDBFFC3FFE28EF5F97AFDA9F941" box="[457,541,1735,1760]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="7" pageNumber="342">Fig. 45</figureCitation>
|
|||
|
|
). Spiny funnel with many dense, small spines dorsally, and ventral small portion (capsula interna) lacking spines (
|
|||
|
|
<figureCitation id="137DFFDBFFC3FFE28D39F951FDEDF8A4" box="[517,601,1772,1797]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="7" pageNumber="342">Fig. 46</figureCitation>
|
|||
|
|
). Stylus long, columnar, distally blunt, gradually thickened to base. Ventral plate with many short setae (
|
|||
|
|
<figureCitation id="137DFFDBFFC3FFE28EE4F8B2FD99F889" box="[472,557,1807,1832]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="7" pageNumber="342">Fig. 43</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBF889FE2AF8EC" blockId="7.[151,1437,151,2013]" box="[199,414,1844,1869]" pageId="7" pageNumber="342">Female. Unknown.</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBF8EAFA31F835" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
|
|||
|
|
<emphasis id="B9323F4CFFC3FFE28FFBF8EAFEC2F8D1" bold="true" box="[199,374,1879,1904]" pageId="7" pageNumber="342">Measurements</emphasis>
|
|||
|
|
. Male: body 6.26 long, 4.12 wide at the widest portion, scutum 5.26 long. Ocularium 0.38 long, 0.69 wide. Pedipalpus claw 0.59 long. Penis 3.94 long. Measurements of pedipalpus and legs as in
|
|||
|
|
<tableCitation id="C6C4D6E5FFC3FFE28A10F8C1FA35F835" box="[1324,1409,1916,1941]" captionStart="TABLE 2" captionStartId="11.[151,239,152,175]" captionTargetBox="[159,1341,202,431]" captionTargetPageId="11" captionText="TABLE 2. Pedipalpus and leg measurements of the male of Paramaracandus dolabratus sp. nov." httpUri="http://table.plazi.org/id/DF39B3D6FFCFFFEE8FABFF25FB05FF0E" pageId="7" pageNumber="342" tableUuid="DF39B3D6FFCFFFEE8FABFF25FB05FF0E">Table 2</tableCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFC3FFE28FFBF81DFF5AF87C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="342">
|
|||
|
|
<emphasis id="B9323F4CFFC3FFE28FFBF81DFE97F818" bold="true" box="[199,291,1952,1977]" pageId="7" pageNumber="342">Habitat</emphasis>
|
|||
|
|
. The specimens were collected by leaf litter sieving in dark moist places of the forest, under dense canopy.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF39B3D6FFCCFFED8FABF81DFBC9F855" httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="8" pageNumber="343" targetBox="[246,1342,188,1906]" targetPageId="8">
|
|||
|
|
<paragraph id="8BF9E35EFFCCFFED8FABF81DFBC9F855" blockId="8.[151,1436,1952,2036]" pageId="8" pageNumber="343">
|
|||
|
|
<emphasis id="B9323F4CFFCCFFED8FABF81DFEE2F814" bold="true" box="[151,342,1952,1975]" pageId="8" pageNumber="343">FIGURES 26–31.</emphasis>
|
|||
|
|
<taxonomicName id="4C4698DDFFCCFFED8E62F81DFD31F817" box="[350,645,1952,1974]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="343" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCCFFED8E62F81DFD31F817" box="[350,645,1952,1974]" italics="true" pageId="8" pageNumber="343">Paramaracandus dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9323F4CFFCCFFED8DB1F81CFD54F817" bold="true" box="[653,736,1953,1974]" pageId="8" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCCFFED8DB1F81CFD54F817" box="[653,736,1953,1974]" pageId="8" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5mm (26–27); 1mm (31); 0.75mm (28–30).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF39B3D6FFCDFFEC8FABF897FEE5F83D" httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="9" pageNumber="344" targetBox="[208,1366,203,1742]" targetPageId="9">
|
|||
|
|
<paragraph id="8BF9E35EFFCDFFEC8FABF897FEE5F83D" blockId="9.[151,1436,1834,1948]" pageId="9" pageNumber="344">
|
|||
|
|
<emphasis id="B9323F4CFFCDFFEC8FABF897FEE7F89E" bold="true" box="[151,339,1834,1857]" pageId="9" pageNumber="344">FIGURES 32–37.</emphasis>
|
|||
|
|
<taxonomicName id="4C4698DDFFCDFFEC8E66F897FDCBF8E1" box="[346,639,1834,1856]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCDFFEC8E66F897FDCBF8E1" box="[346,639,1834,1856]" italics="true" pageId="9" pageNumber="344">Paramaracandus dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9323F4CFFCDFFEC8DB9F896FD62F8E1" bold="true" box="[645,726,1835,1856]" pageId="9" pageNumber="344">
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCDFFEC8DB9F896FD62F8E1" box="[645,726,1835,1856]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
32–33. Holotype. 34–37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5mm (34); 1mm (32–33, 36– 37); 0.75mm (35).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF39B3D6FFCEFFEF8FABF8CFFCF9F845" httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="10" pageNumber="345" targetBox="[263,1329,239,1843]" targetPageId="10">
|
|||
|
|
<paragraph id="8BF9E35EFFCEFFEF8FABF8CFFCF9F845" blockId="10.[151,1436,1906,2020]" pageId="10" pageNumber="345">
|
|||
|
|
<emphasis id="B9323F4CFFCEFFEF8FABF8CFFEE1F826" bold="true" box="[151,341,1906,1929]" pageId="10" pageNumber="345">FIGURES 38–46.</emphasis>
|
|||
|
|
<taxonomicName id="4C4698DDFFCEFFEF8E60F8CFFD36F829" box="[348,642,1906,1928]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="345" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCEFFEF8E60F8CFFD36F829" box="[348,642,1906,1928]" italics="true" pageId="10" pageNumber="345">Paramaracandus dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9323F4CFFCEFFEF8DB5F8CEFD6EF829" bold="true" box="[649,730,1907,1928]" pageId="10" pageNumber="345">
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCEFFEF8DB5F8CEFD6EF829" box="[649,730,1907,1928]" pageId="10" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3mm (38–40); 0.75mm (44–46); 0.5mm (41–43).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF39B3D6FFCFFFEE8FABFF25FB05FF0E" ID-Table-UUID="DF39B3D6FFCFFFEE8FABFF25FB05FF0E" box="[151,1201,151,175]" httpUri="http://table.plazi.org/id/DF39B3D6FFCFFFEE8FABFF25FB05FF0E" pageId="11" pageNumber="346" targetBox="[159,1341,202,431]" targetIsTable="true" targetPageId="11">
|
|||
|
|
<paragraph id="8BF9E35EFFCFFFEE8FABFF25FB05FF0E" blockId="11.[151,1201,151,175]" box="[151,1201,151,175]" pageId="11" pageNumber="346">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8FABFF25FEBDFF0E" bold="true" box="[151,265,152,175]" pageId="11" pageNumber="346">TABLE 2.</emphasis>
|
|||
|
|
Pedipalpus and leg measurements of the male of
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8C27FF2AFBE0FF0E" box="[795,1108,151,175]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8C27FF2AFBE0FF0E" bold="true" box="[795,1108,151,175]" italics="true" pageId="11" pageNumber="346">Paramaracandus dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCFFFEE8B67FF25FB05FF0E" box="[1115,1201,152,175]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BF9E35EFFCFFFEE8E7EFF77FA88FE0E" pageId="11" pageNumber="346">
|
|||
|
|
<table id="F94611FEFFCF001A8FA3FF77FA89FE0E" box="[159,1341,202,431]" gridcols="5" gridrows="6" pageId="11" pageNumber="346">
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FF77FA89FF41" box="[159,1341,202,224]" gridrow="0" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FF77FD92FF41" box="[159,550,202,224]" gridcol="0" gridrow="0" pageId="11" pageNumber="346">Trochanter Femur</th>
|
|||
|
|
<th id="76A78860FFCF001A8DBFFF77FD7CFF41" box="[643,712,202,224]" gridcol="1" gridrow="0" pageId="11" pageNumber="346">Patella</th>
|
|||
|
|
<th id="76A78860FFCF001A8C1FFF77FCE3FF41" box="[803,855,202,224]" gridcol="2" gridrow="0" pageId="11" pageNumber="346">Tibia</th>
|
|||
|
|
<th id="76A78860FFCF001A8CFEFF77FB13FF41" box="[962,1191,202,224]" gridcol="3" gridrow="0" pageId="11" pageNumber="346">Metatarsus Tarsus</th>
|
|||
|
|
<th id="76A78860FFCF001A8A38FF77FA89FF41" box="[1284,1341,202,224]" gridcol="4" gridrow="0" pageId="11" pageNumber="346">Total</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FF49FA89FEAB" box="[159,1341,244,266]" gridrow="1" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FF49FD92FEAB" box="[159,550,244,266]" gridcol="0" gridrow="1" pageId="11" pageNumber="346">Pedipalpus 0.75 1.66</th>
|
|||
|
|
<td id="76A78860FFCF001A8DBFFF49FD7CFEAB" box="[643,712,244,266]" gridcol="1" gridrow="1" pageId="11" pageNumber="346">1.19</td>
|
|||
|
|
<td id="76A78860FFCF001A8C1FFF49FCE3FEAB" box="[803,855,244,266]" gridcol="2" gridrow="1" pageId="11" pageNumber="346">0.84</td>
|
|||
|
|
<td id="76A78860FFCF001A8CFEFF49FB13FEAB" box="[962,1191,244,266]" gridcol="3" gridrow="1" pageId="11" pageNumber="346">0.89</td>
|
|||
|
|
<td id="76A78860FFCF001A8A38FF49FA89FEAB" box="[1284,1341,244,266]" gridcol="4" gridrow="1" pageId="11" pageNumber="346">5.33</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FEA0FA89FE92" box="[159,1341,285,307]" gridrow="2" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FEA0FD92FE92" box="[159,550,285,307]" gridcol="0" gridrow="2" pageId="11" pageNumber="346">Leg I 0.63 3.03</th>
|
|||
|
|
<td id="76A78860FFCF001A8DBFFEA0FD7CFE92" box="[643,712,285,307]" gridcol="1" gridrow="2" pageId="11" pageNumber="346">0.99</td>
|
|||
|
|
<td id="76A78860FFCF001A8C1FFEA0FCE3FE92" box="[803,855,285,307]" gridcol="2" gridrow="2" pageId="11" pageNumber="346">1.91</td>
|
|||
|
|
<td id="76A78860FFCF001A8CFEFEA0FB13FE92" box="[962,1191,285,307]" gridcol="3" gridrow="2" pageId="11" pageNumber="346">3.21 1.68</td>
|
|||
|
|
<td id="76A78860FFCF001A8A38FEA0FA89FE92" box="[1284,1341,285,307]" gridcol="4" gridrow="2" pageId="11" pageNumber="346">11.45</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FEFBFA89FEFD" box="[159,1341,326,348]" gridrow="3" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FEFBFD92FEFD" box="[159,550,326,348]" gridcol="0" gridrow="3" pageId="11" pageNumber="346">Leg II 0.71 5.57</th>
|
|||
|
|
<td id="76A78860FFCF001A8DBFFEFBFD7CFEFD" box="[643,712,326,348]" gridcol="1" gridrow="3" pageId="11" pageNumber="346">1.33</td>
|
|||
|
|
<td id="76A78860FFCF001A8C1FFEFBFCE3FEFD" box="[803,855,326,348]" gridcol="2" gridrow="3" pageId="11" pageNumber="346">4.57</td>
|
|||
|
|
<td id="76A78860FFCF001A8CFEFEFBFB13FEFD" box="[962,1191,326,348]" gridcol="3" gridrow="3" pageId="11" pageNumber="346">5.42 3.07</td>
|
|||
|
|
<td id="76A78860FFCF001A8A38FEFBFA89FEFD" box="[1284,1341,326,348]" gridcol="4" gridrow="3" pageId="11" pageNumber="346">20.67</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FECDFA89FE27" box="[159,1341,368,390]" gridrow="4" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FECDFD92FE27" box="[159,550,368,390]" gridcol="0" gridrow="4" pageId="11" pageNumber="346">Leg III 0.83 4.01</th>
|
|||
|
|
<td id="76A78860FFCF001A8DBFFECDFD7CFE27" box="[643,712,368,390]" gridcol="1" gridrow="4" pageId="11" pageNumber="346">1.26</td>
|
|||
|
|
<td id="76A78860FFCF001A8C1FFECDFCE3FE27" box="[803,855,368,390]" gridcol="2" gridrow="4" pageId="11" pageNumber="346">2.34</td>
|
|||
|
|
<td id="76A78860FFCF001A8CFEFECDFB13FE27" box="[962,1191,368,390]" gridcol="3" gridrow="4" pageId="11" pageNumber="346">4.04 1.78</td>
|
|||
|
|
<td id="76A78860FFCF001A8A38FECDFA89FE27" box="[1284,1341,368,390]" gridcol="4" gridrow="4" pageId="11" pageNumber="346">14.26</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3576E11CFFCF001A8FA3FE24FA89FE0E" box="[159,1341,409,431]" gridrow="5" pageId="11" pageNumber="346">
|
|||
|
|
<th id="76A78860FFCF001A8FA3FE24FD92FE0E" box="[159,550,409,431]" gridcol="0" gridrow="5" pageId="11" pageNumber="346">Leg IV 0.89 5.51</th>
|
|||
|
|
<td id="76A78860FFCF001A8DBFFE24FD7CFE0E" box="[643,712,409,431]" gridcol="1" gridrow="5" pageId="11" pageNumber="346">1.33</td>
|
|||
|
|
<td id="76A78860FFCF001A8C1FFE24FCE3FE0E" box="[803,855,409,431]" gridcol="2" gridrow="5" pageId="11" pageNumber="346">3.26</td>
|
|||
|
|
<td id="76A78860FFCF001A8CFEFE24FB13FE0E" box="[962,1191,409,431]" gridcol="3" gridrow="5" pageId="11" pageNumber="346">5.99 2.13</td>
|
|||
|
|
<td id="76A78860FFCF001A8A38FE24FA89FE0E" box="[1284,1341,409,431]" gridcol="4" gridrow="5" pageId="11" pageNumber="346">19.11</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF9E35EFFCFFFEE8FFBFE49FBB1FDF4" blockId="11.[151,1436,500,1029]" pageId="11" pageNumber="346">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8FFBFE49FE81FDAC" bold="true" box="[199,309,500,525]" pageId="11" pageNumber="346">Variation</emphasis>
|
|||
|
|
(male
|
|||
|
|
<typeStatus id="54FD5DFCFFCFFFEE8EB0FE49FE46FDAD" box="[396,498,500,524]" pageId="11" pageNumber="346" type="paratype">paratype</typeStatus>
|
|||
|
|
). Body 7.07 long, 4.31 wide at the widest portion, scutum 5.22 long. The single
|
|||
|
|
<typeStatus id="54FD5DFCFFCFFFEE8FABFDA5FF4EFD91" box="[151,250,536,560]" pageId="11" pageNumber="346" type="paratype">paratype</typeStatus>
|
|||
|
|
, with larger body and rounded abdomen posteriorly (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8C64FDAAFC18FD91" box="[856,940,535,560]" captionStart="FIGURES 32 – 37" captionStartId="9.[151,264,1834,1857]" captionTargetBox="[208,1366,203,1742]" captionTargetId="figure@9.[200,1387,193,1782]" captionTargetPageId="9" captionText="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="11" pageNumber="346">Fig. 34</figureCitation>
|
|||
|
|
), and its finger teeth of chelicerae (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8A7DFDAAFA20FD91" box="[1345,1428,535,560]" captionStart="FIGURES 32 – 37" captionStartId="9.[151,264,1834,1857]" captionTargetBox="[208,1366,203,1742]" captionTargetId="figure@9.[200,1387,193,1782]" captionTargetPageId="9" captionText="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="11" pageNumber="346">Fig. 35</figureCitation>
|
|||
|
|
) and pedipalpus (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8E73FD81FE6CFDF4" box="[335,472,572,597]" captionStart="FIGURES 32 – 37" captionStartId="9.[151,264,1834,1857]" captionTargetBox="[208,1366,203,1742]" captionTargetId="figure@9.[200,1387,193,1782]" captionTargetPageId="9" captionText="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="11" pageNumber="346">Figs. 36–37</figureCitation>
|
|||
|
|
), shows only slight differences with the
|
|||
|
|
<typeStatus id="54FD5DFCFFCFFFEE8CA7FD81FBB5FDF4" box="[923,1025,572,597]" pageId="11" pageNumber="346" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C35CB0D5FFCFFFEE8FFBFDE2FD45FDD9" box="[199,753,607,632]" pageId="11" pageNumber="346" type="distribution">
|
|||
|
|
<paragraph id="8BF9E35EFFCFFFEE8FFBFDE2FD45FDD9" blockId="11.[151,1436,500,1029]" box="[199,753,607,632]" pageId="11" pageNumber="346">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8FFBFDE2FEEDFDD9" bold="true" box="[199,345,607,632]" pageId="11" pageNumber="346">Distribution</emphasis>
|
|||
|
|
. Known only from the
|
|||
|
|
<typeStatus id="54FD5DFCFFCFFFEE8D5CFDDDFD24FDD9" box="[608,656,608,632]" pageId="11" pageNumber="346">type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C35CB0D5FFCFFFEE8FFBFD39FD64FBA4" pageId="11" pageNumber="346" type="discussion">
|
|||
|
|
<paragraph id="8BF9E35EFFCFFFEE8FFBFD39FBF2FC39" blockId="11.[151,1436,500,1029]" pageId="11" pageNumber="346">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8FFBFD39FE80FD3C" bold="true" box="[199,308,644,669]" pageId="11" pageNumber="346">Remarks</emphasis>
|
|||
|
|
. The genus
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8EEAFD39FD2CFD3D" box="[470,664,644,668]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8EEAFD39FD2CFD3D" box="[470,664,644,668]" italics="true" pageId="11" pageNumber="346">Paramaracandus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
encompasses three species,
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8CC4FD38FBEAFD3D" box="[1016,1118,644,668]" italics="true" pageId="11" pageNumber="346">P. f u s c u s</emphasis>
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8B4DFD39FAB9FD3C" author="Suzuki" box="[1137,1293,644,669]" pageId="11" pageNumber="346" refString="Suzuki, S. (1976) Report on a collection of opilionids from Pasoh Forest Reserve, West Malaysia. Nature and Life in Southeast Asia, 7, 9 - 38." type="journal article" year="1976">Suzuki, 1976</bibRefCitation>
|
|||
|
|
(
|
|||
|
|
<typeStatus id="54FD5DFCFFCFFFEE8A18FD39FAE2FD3D" box="[1316,1366,644,668]" pageId="11" pageNumber="346">type</typeStatus>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="F351A3CEFFCFFFEE8FABFD1AFEB6FD61" box="[151,258,679,704]" name="Malaysia" pageId="11" pageNumber="346">Malaysia</collectingCountry>
|
|||
|
|
),
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8E24FD14FDE8FD61" authority="Suzuki, 1985" authorityName="Suzuki" authorityYear="1985" box="[280,604,679,704]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="sexdentatus">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8E24FD14FE0EFD61" box="[280,442,680,704]" italics="true" pageId="11" pageNumber="346">P. sexdentatus</emphasis>
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8EF8FD1AFDE8FD61" author="Suzuki" box="[452,604,679,704]" pageId="11" pageNumber="346" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11 (3), 69 - 110." type="journal article" year="1985">Suzuki, 1985</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(from
|
|||
|
|
<collectingCountry id="F351A3CEFFCFFFEE8D8DFD1AFCA2FD61" box="[689,790,679,704]" name="Thailand" pageId="11" pageNumber="346">Thailand</collectingCountry>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8C60FD14FC40FD61" box="[860,1012,680,704]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8C60FD14FC40FD61" box="[860,1012,680,704]" italics="true" pageId="11" pageNumber="346">P. dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8CC3FD1AFBE8FD61" bold="true" box="[1023,1116,679,704]" pageId="11" pageNumber="346">
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCFFFEE8CC3FD1AFBE8FD61" box="[1023,1116,679,704]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(from
|
|||
|
|
<collectingCountry id="F351A3CEFFCFFFEE8B8DFD1AFB5EFD61" box="[1201,1258,679,704]" name="Laos" pageId="11" pageNumber="346">Laos</collectingCountry>
|
|||
|
|
).The latter two species seem more similar to each other than
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8DA4FD70FD42FD45" box="[664,758,716,740]" italics="true" pageId="11" pageNumber="346">P. f u s c us</emphasis>
|
|||
|
|
, e.g., dorsa are more swollen, ocular tubercles are provided with a pair of conspicuous spines, coxae IV possess many tubercles prolaterally (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8B1AFD52FBCDFCA9" box="[1062,1145,751,776]" captionStart="FIGURES 26 – 31" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[246,1342,188,1906]" captionTargetId="figure@8.[181,1368,157,1937]" captionTargetPageId="8" captionText="FIGURES 26 – 31. Paramaracandus dolabratus sp. nov., Holotype. 26. Male body, lateral view. 27. Same, dorsal view. 28. Left chelicera of male, medial view. 29. Same, ectal view. 30. Cheliceral fingers of male, frontal view. 31. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (26 – 27); 1 mm (31); 0.75 mm (28 – 30)." httpUri="https://zenodo.org/record/243908/files/figure.png" pageId="11" pageNumber="346">Fig. 26</figureCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8BB9FD52FAA2FCA9" author="Suzuki" box="[1157,1302,751,776]" pageId="11" pageNumber="346" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11 (3), 69 - 110." type="journal article" year="1985">Suzuki 1985</bibRefCitation>
|
|||
|
|
: 98, fig. B), femora of pedipalps have a setiferous tubercle on the medial distal side (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8CD5FCA9FBF4FC8D" box="[1001,1088,788,813]" captionStart="FIGURES 32 – 37" captionStartId="9.[151,264,1834,1857]" captionTargetBox="[208,1366,203,1742]" captionTargetId="figure@9.[200,1387,193,1782]" captionTargetPageId="9" captionText="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="11" pageNumber="346">Fig. 33</figureCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8B73FCA9FB57FC8C" author="Suzuki" box="[1103,1251,788,813]" pageId="11" pageNumber="346" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11 (3), 69 - 110." type="journal article" year="1985">Suzuki 1985</bibRefCitation>
|
|||
|
|
: 98, fig. E) and ventral plates of penis are extended noticeably and covered with many short setae (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8B0FFC8AFB32FCF1" box="[1075,1158,823,848]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="11" pageNumber="346">Fig. 43</figureCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8BAEFC8AFA96FCF1" author="Suzuki" box="[1170,1314,823,848]" pageId="11" pageNumber="346" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11 (3), 69 - 110." type="journal article" year="1985">Suzuki 1985</bibRefCitation>
|
|||
|
|
: 98, fig. I). However,
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8E31FCE0FE10FCD5" box="[269,420,860,884]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="dolabratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8E31FCE0FE10FCD5" box="[269,420,860,884]" italics="true" pageId="11" pageNumber="346">P. dolabratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8E92FCE1FDBFFCD4" bold="true" box="[430,523,860,885]" pageId="11" pageNumber="346">
|
|||
|
|
<taxonomicNameLabel id="A2018237FFCFFFEE8E92FCE1FDBFFCD4" box="[430,523,860,885]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
can be easily distinguished from
|
|||
|
|
<taxonomicName id="4C4698DDFFCFFFEE8CAFFCE0FB80FCD5" box="[915,1076,860,884]" class="Arachnida" family="Assamiidae" genus="Paramaracandus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="sexdentatus">
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8CAFFCE0FB80FCD5" box="[915,1076,860,884]" italics="true" pageId="11" pageNumber="346">P. sexdentatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<emphasis id="B9323F4CFFCFFFEE8B53FCE0FB64FCD5" box="[1135,1232,860,884]" italics="true" pageId="11" pageNumber="346">P. f u s c u s</emphasis>
|
|||
|
|
by the triangular ventral plate in the penis (
|
|||
|
|
<figureCitation id="137DFFDBFFCFFFEE8E85FCC2FDB9FC39" box="[441,525,895,920]" captionStart="FIGURES 38 – 46" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[263,1329,239,1843]" captionTargetId="figure@10.[200,1387,193,1885]" captionTargetPageId="10" captionText="FIGURES 38 – 46. Paramaracandus dolabratus sp. nov., Holotype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Distal part of penis, dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Same (expanded), dorsal view. 45. Same, lateral view. 46. Same, ventral view. CI, capsula interna; CE, capsula externa; Fo, follis; S, stylus; SF, spiny funnel. Scale bars: 3 mm (38 – 40); 0.75 mm (44 – 46); 0.5 mm (41 – 43)." httpUri="https://zenodo.org/record/243910/files/figure.png" pageId="11" pageNumber="346">Fig. 38</figureCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFD79EAFFFCFFFEE8D25FCC2FD1DFC39" author="Suzuki" box="[537,681,895,920]" pageId="11" pageNumber="346" refString="Suzuki, S. (1985) A synopsis of the Opiliones of Thailand (Arachnida) I. Cyphophthalmi and Laniatores. Steenstrupia, 11 (3), 69 - 110." type="journal article" year="1985">Suzuki 1985</bibRefCitation>
|
|||
|
|
: 98, fig. G;
|
|||
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<bibRefCitation id="EFD79EAFFFCFFFEE8C12FCC2FC0BFC39" author="Suzuki" box="[814,959,895,920]" pageId="11" pageNumber="346" refString="Suzuki, S. (1976) Report on a collection of opilionids from Pasoh Forest Reserve, West Malaysia. Nature and Life in Southeast Asia, 7, 9 - 38." type="journal article" year="1976">Suzuki 1976</bibRefCitation>
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: 25, fig. H).
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</paragraph>
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<paragraph id="8BF9E35EFFCFFFEE8FFBFC19FD64FBA4" blockId="11.[151,1436,500,1029]" pageId="11" pageNumber="346">
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The appearance of the only
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<typeStatus id="54FD5DFCFFCFFFEE8D1CFC18FD37FC1C" box="[544,643,933,957]" pageId="11" pageNumber="346" type="paratype">paratype</typeStatus>
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, having larger body size and especially the blunt round abdomen posteriorly, initially led us to consider it a female (
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<figureCitation id="137DFFDBFFCFFFEE8DEEFC7AFC93FC41" box="[722,807,967,992]" captionStart="FIGURES 32 – 37" captionStartId="9.[151,264,1834,1857]" captionTargetBox="[208,1366,203,1742]" captionTargetId="figure@9.[200,1387,193,1782]" captionTargetPageId="9" captionText="FIGURES 32 – 37. Paramaracandus dolabratus sp. nov. 32 – 33. Holotype. 34 – 37. Paratype. 32. Left pedipalpus of male, ectal view. 33. Same, medial view. 34. Male body, dorsal view. 35. Left cheliceral fingers of male, frontal view. 36. Left pedipalpus of male, ectal view. 37. Patella, tibia and tarsus of male left pedipalpus, dorsal view. Scale bars: 5 mm (34); 1 mm (32 – 33, 36 – 37); 0.75 mm (35)." httpUri="https://zenodo.org/record/243909/files/figure.png" pageId="11" pageNumber="346">Fig. 34</figureCitation>
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). After we dissected the genitalia, we recognized it as a male. This is another example of species variation.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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