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<document id="0B1FA63B034CA01F84CB4EC38BD814A3" ID-DOI="10.11646/zootaxa.3741.4.9" ID-GBIF-Dataset="6da74c9b-e13a-4008-8bfd-c0e49b249ca9" ID-ISSN="1175-5326" ID-Zenodo-Dep="223648" ID-ZooBank="E8A2187A-B3EC-4542-965D-39C9F39B177B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460704428271" checkinUser="plazi" docAuthor="Greenfield, David W. &amp; Erdmann, Mark V." docDate="2013" docId="03D7878AFFB5C3555EFDFA2BFD2C29E5" docLanguage="en" docName="zt03741p600.pdf" docOrigin="Zootaxa 3741 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eviota santanai Greenfield &amp; Erdmann, 2013, n. sp." docType="treatment" docUuid="E8BE6C87-6A59-46A9-8D05-F97A5616C4AD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="599" masterDocId="FFEEFFF2FFB4C3535E6AFF85FF822F26" masterDocTitle="Eviota santanai, a new Dwarfgoby from Timor-Leste (Teleostei: Gobiidae)" masterLastPageNumber="600" masterPageNumber="593" pageNumber="594" updateTime="1698356809481" updateUser="plazi">
<mods:mods id="6CCDC25E71D0F48995BAB8959607E1FC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="26A67FDE9FD2A7624983F7A322433AE9">
<mods:title id="92F145668FB0C370813BF11885D97503">Eviota santanai, a new Dwarfgoby from Timor-Leste (Teleostei: Gobiidae)</mods:title>
</mods:titleInfo>
<mods:name id="A89EDD176021964D56AF360123BF50D0" type="personal">
<mods:role id="C28EF1FD4283C15FF525BEB28F6167C8">
<mods:roleTerm id="F8F08CF53B66C9D42D54017525573939">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB2F76D40176EBE5A09B77A564A65C6D">Greenfield, David W.</mods:namePart>
</mods:name>
<mods:name id="EE2A722505F07BE608D6347B3E7908B3" type="personal">
<mods:role id="CFB492C911472DE5B36F8920A57ED62D">
<mods:roleTerm id="458A964EE5B232CDBF85D1DCE14C5488">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B4803DE08B7386A103970698D80D6FA">Erdmann, Mark V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="681106EE2BF7FFAD2D88A7CFB77D9F30">text</mods:typeOfResource>
<mods:relatedItem id="B1DE6C3EACC46B157F48864140ED6304" type="host">
<mods:titleInfo id="E9B6E4B849529D26671749E788810327">
<mods:title id="294AB63E7987FD36BF6CB5935D842B1B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6FA54E80D0B6BF1E2F0A9AC16E427D22">
<mods:date id="97F2C0646CCFD88191897667B452DEFE">2013</mods:date>
<mods:detail id="EA96109FBECD24AF6EB95E76DD4B76AF" type="volume">
<mods:number id="C6085A75E76DD6721C08A6105173E887">3741</mods:number>
</mods:detail>
<mods:detail id="523641E27A7B3D33AC388ED045647237" type="issue">
<mods:number id="0E5FDC81BA34514C431415AC36C890C1">4</mods:number>
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<mods:start id="6BD7E0FAB3AE39BED2B2E7D96E9FCFDC">593</mods:start>
<mods:end id="7D0E758504EA62222A2376C4FED42ED8">600</mods:end>
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<mods:classification id="F052EAC84382223097AA16BE5B51490C">journal article</mods:classification>
<mods:identifier id="3C3E8F5BBE2FAC131AD6F11437AEA482" type="DOI">10.11646/zootaxa.3741.4.9</mods:identifier>
<mods:identifier id="324A00690894F0AFC440BB4868D33343" type="GBIF-Dataset">6da74c9b-e13a-4008-8bfd-c0e49b249ca9</mods:identifier>
<mods:identifier id="2E6B0FB7FB8A04219AA4624174FE5706" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7DF1850C5ED328803DC8EEB7853C2FD8" type="Zenodo-Dep">223648</mods:identifier>
<mods:identifier id="DCFEE2A80F05BACB9FEE5392ADA9EFC9" type="ZooBank">E8A2187A-B3EC-4542-965D-39C9F39B177B</mods:identifier>
</mods:mods>
<treatment id="03D7878AFFB5C3555EFDFA2BFD2C29E5" ID-DOI="http://doi.org/10.5281/zenodo.6152335" ID-GBIF-Taxon="119583780" ID-Zenodo-Dep="6152335" LSID="urn:lsid:zoobank.org:act:E8BE6C87-6A59-46A9-8D05-F97A5616C4AD" httpUri="http://treatment.plazi.org/id/03D7878AFFB5C3555EFDFA2BFD2C29E5" lastPageId="6" lastPageNumber="599" pageId="1" pageNumber="594">
<subSubSection id="C3646517FFB5C3525EFDFA2BFDDC2975" pageId="1" pageNumber="594" type="nomenclature">
<paragraph id="8BC1369CFFB5C3525EFDFA2BFE272AEF" blockId="1.[151,421,1454,1550]" box="[151,421,1454,1481]" pageId="1" pageNumber="594">
<heading id="D08981F0FFB5C3525EFDFA2BFE272AEF" bold="true" box="[151,421,1454,1481]" fontSize="11" level="1" pageId="1" pageNumber="594" reason="1">
<emphasis id="B90AEA8EFFB5C3525EFDFA2BFE272AEF" bold="true" box="[151,421,1454,1481]" pageId="1" pageNumber="594">
<taxonomicName id="4C7E4D1FFFB5C3525EFDFA2BFED92AEE" ID-CoL="3DHNP" box="[151,347,1454,1480]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="594" phylum="Chordata" rank="species" species="santanai" status="sp. nov.">
<emphasis id="B90AEA8EFFB5C3525EFDFA2BFED92AEE" bold="true" box="[151,347,1454,1480]" italics="true" pageId="1" pageNumber="594">Eviota santanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23957F5FFB5C3525F08FA2AFE272AEF" box="[354,421,1455,1481]" pageId="1" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC1369CFFB5C3525EFDFA57FE842928" blockId="1.[151,421,1454,1550]" pageId="1" pageNumber="594">
Santana’s Dwarfgoby
<figureCitation id="13452A19FFB5C3525EFDFA70FE802928" box="[151,258,1525,1550]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1486,1508]" captionStartId-1="3.[151,250,1514,1536]" captionStartId-2="4.[151,250,1959,1981]" captionTargetBox-0="[151,1436,1060,1465]" captionTargetBox-1="[151,1436,661,1493]" captionTargetBox-2="[202,1390,210,1933]" captionTargetId-0="figure@2.[151,1436,1060,1465]" captionTargetId-1="figure@3.[151,1436,661,1493]" captionTargetId-2="figure@4.[195,1400,193,1947]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Color in preservative of holotype of E. santanai, CAS 234966, 11.8 mm. Photograph by D. W. Greenfield." captionText-1="FIGURE 2. Underwater photograph of E. santanai. Photograph by M. V. Erdmann." captionText-2="FIGURE 3. Head coloration of preserved specimens, with pigmentation differences indicated by arrows. A. Eviota santanai holotype, CAS 234966, 11.8 mm. B. Eviota latifasciata paratype, USNM 260079, 12.7 mm. Photograph by D. W. Greenfield." httpUri-0="https://zenodo.org/record/223649/files/figure.png" httpUri-1="https://zenodo.org/record/223650/files/figure.png" httpUri-2="https://zenodo.org/record/223651/files/figure.png" pageId="1" pageNumber="594">Figs. 1–3</figureCitation>
.
</paragraph>
<paragraph id="8BC1369CFFB5C3525EFDF9BAFDDC2975" blockId="1.[151,606,1597,1620]" box="[151,606,1597,1620]" pageId="1" pageNumber="594">
<taxonomicName id="4C7E4D1FFFB5C3525EFDF9BAFF5B2972" box="[151,217,1599,1620]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="594" phylum="Chordata" rank="genus">
<emphasis id="B90AEA8EFFB5C3525EFDF9BAFF5B2972" box="[151,217,1599,1620]" italics="true" pageId="1" pageNumber="594">Eviota</emphasis>
</taxonomicName>
species 1 Allen &amp; Erdmann 2013: 79.
</paragraph>
</subSubSection>
<subSubSection id="C3646517FFB5C3525EFDF905FD5128B2" pageId="1" pageNumber="594" type="materials_examined">
<paragraph id="8BC1369CFFB5C3525EFDF905FED3299B" blockId="1.[151,1436,1664,2013]" pageId="1" pageNumber="594">
<emphasis id="B90AEA8EFFB5C3525EFDF905FE8F29BF" bold="true" box="[151,269,1664,1689]" pageId="1" pageNumber="594">
<typeStatus id="54C5883EFFB5C3525EFDF905FE8A29BF" box="[151,264,1664,1689]" pageId="1" pageNumber="594" type="holotype">Holotype</typeStatus>
:
</emphasis>
CAS 234966,
<quantity id="4C869B79FFB5C3525FACF904FDAB29BE" box="[454,553,1665,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="594" unit="mm" value="11.8">11.8 mm</quantity>
male, Tutuala,
<collectingCountry id="F369760CFFB5C3525CBFF905FCE029BF" box="[725,866,1664,1689]" name="Timor-Leste" pageId="1" pageNumber="594">Timor-Leste</collectingCountry>
,
<geoCoordinate id="EE4A505BFFB5C3525D07F904FC7F29BF" box="[877,1021,1665,1689]" direction="south" orientation="latitude" pageId="1" pageNumber="594" precision="1" value="-8.36205">08°21.723’S</geoCoordinate>
,
<geoCoordinate id="EE4A505BFFB5C3525A6DF904FB2F29BF" box="[1031,1197,1665,1689]" direction="east" orientation="longitude" pageId="1" pageNumber="594" precision="1" value="127.0748">127° 04.488’E</geoCoordinate>
,
<quantity id="4C869B79FFB5C3525AD2F904FB6729BE" box="[1208,1253,1665,1689]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="594" unit="m" value="8.0">8 m</quantity>
, M.V. Erdmann,
<date id="FFC0105CFFB5C3525EFDF921FECC299B" box="[151,334,1700,1725]" pageId="1" pageNumber="594" value="2012-08-16">16 August 2012</date>
.
</paragraph>
<paragraph id="8BC1369CFFB5C3525EADF94DFAA42823" blockId="1.[151,1436,1664,2013]" pageId="1" pageNumber="594">
<emphasis id="B90AEA8EFFB5C3525EADF94DFEBD29C7" bold="true" box="[199,319,1736,1761]" pageId="1" pageNumber="594">
<typeStatus id="54C5883EFFB5C3525EADF94DFEBD29C7" box="[199,319,1736,1761]" pageId="1" pageNumber="594" type="paratype">Paratypes</typeStatus>
</emphasis>
: USNM 410587,
<quantity id="4C869B79FFB5C3525C74F94CFD0429C6" box="[542,646,1737,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="594" unit="mm" value="12.8">12.8 mm</quantity>
male, taken with
<typeStatus id="54C5883EFFB5C3525D34F94DFC4629C7" box="[862,964,1736,1761]" pageId="1" pageNumber="594" type="holotype">holotype</typeStatus>
; CAS 235688,
<quantity id="4C869B79FFB5C3525A10F94DFB6329C6" box="[1146,1249,1736,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="594" unit="mm" value="11.5">11.5 mm</quantity>
male,
<quantity id="4C869B79FFB5C3525B5EF94CFA1E29C6" box="[1332,1436,1737,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="594" unit="mm" value="10.7">10.7 mm</quantity>
female,
<collectingCountry id="F369760CFFB5C3525E9BF969FEB32823" box="[241,305,1772,1797]" name="Iceland" pageId="1" pageNumber="594">Pulau</collectingCountry>
Atauro,
<collectingCountry id="F369760CFFB5C3525FF9F969FDA22823" box="[403,544,1772,1797]" name="Timor-Leste" pageId="1" pageNumber="594">Timor-Leste</collectingCountry>
,
<geoCoordinate id="EE4A505BFFB5C3525C59F968FD402823" box="[563,706,1773,1797]" direction="south" orientation="latitude" pageId="1" pageNumber="594" precision="1" value="-8.22135">08°13.281’S</geoCoordinate>
,
<geoCoordinate id="EE4A505BFFB5C3525CA4F969FCF62823" box="[718,884,1772,1797]" direction="east" orientation="longitude" pageId="1" pageNumber="594" precision="1" value="125.61383">125° 36.830’E</geoCoordinate>
,
<quantity id="4C869B79FFB5C3525DEAF969FC2E2822" box="[896,940,1772,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="594" unit="m" value="5.0">5 m</quantity>
, M.V. Erdmann,
<date id="FFC0105CFFB5C3525A06F968FAA02823" box="[1132,1314,1773,1797]" pageId="1" pageNumber="594" value="2012-08-23">23 August 2012</date>
.
</paragraph>
<paragraph id="8BC1369CFFB5C3525EADF895FD5128B2" blockId="1.[151,1436,1664,2013]" pageId="1" pageNumber="594">
<emphasis id="B90AEA8EFFB5C3525EADF895FE75280F" bold="true" box="[199,503,1808,1833]" pageId="1" pageNumber="594">Other material examined</emphasis>
:
<taxonomicName id="4C7E4D1FFFB5C3525C7AF894FD62280E" box="[528,736,1808,1832]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="594" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB5C3525C7AF894FD62280E" box="[528,736,1808,1832]" italics="true" pageId="1" pageNumber="594">Eviota latifasciata</emphasis>
</taxonomicName>
- BPBM 28959
<typeStatus id="54C5883EFFB5C3525DFCF894FB86280F" box="[918,1028,1809,1833]" pageId="1" pageNumber="594" type="paratype">paratypes</typeStatus>
, 10.8 &amp;
<quantity id="4C869B79FFB5C3525A03F894FB4D280E" box="[1129,1231,1809,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="594" unit="mm" value="11.7">11.7 mm</quantity>
,
<collectingCountry id="F369760CFFB5C3525AB7F895FA01280F" box="[1245,1411,1808,1833]" name="Kiribati" pageId="1" pageNumber="594">Gilbert Islands</collectingCountry>
= (
<collectingCountry id="F369760CFFB5C3525EF4F8B1FF78286B" box="[158,250,1844,1869]" name="Kiribati" pageId="1" pageNumber="594">Kiribati</collectingCountry>
,
<collectingCountry id="F369760CFFB5C3525F63F8B0FEE5286B" box="[265,359,1845,1869]" name="Kiribati" pageId="1" pageNumber="594">Tungaru</collectingCountry>
Islands); CAS 52737
<typeStatus id="54C5883EFFB5C3525C04F8B0FD53286B" box="[622,721,1845,1869]" pageId="1" pageNumber="594" type="paratype">paratype</typeStatus>
,
<quantity id="4C869B79FFB5C3525C8AF8B1FCCB286A" box="[736,841,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="594" unit="mm" value="12.6">12.6 mm</quantity>
,
<collectingCountry id="F369760CFFB5C3525D32F8B1FB87286B" box="[856,1029,1844,1869]" name="Kiribati" pageId="1" pageNumber="594">Gilbert Islands</collectingCountry>
; USNM 260079
<typeStatus id="54C5883EFFB5C3525AA4F8B0FABE286B" box="[1230,1340,1845,1869]" pageId="1" pageNumber="594" type="paratype">paratypes</typeStatus>
, 10.1 &amp; 12.7,
<collectingCountry id="F369760CFFB5C3525EBCF8DDFE022857" box="[214,384,1880,1905]" name="Kiribati" pageId="1" pageNumber="594">Gilbert Islands</collectingCountry>
; CAS 226268 (4), 10.4–12.2,
<collectingCountry id="F369760CFFB5C3525CB9F8DCFCAB2856" box="[723,809,1881,1904]" name="Micronesia" pageId="1" pageNumber="594">Ponape</collectingCountry>
; CAS 226295, 11.5 &amp; 12.5,
<collectingCountry id="F369760CFFB5C3525A03F8DCFB3D2856" box="[1129,1215,1881,1904]" name="Micronesia" pageId="1" pageNumber="594">Ponape</collectingCountry>
; CAS 226296, 12.0 &amp; 12.8,
<collectingCountry id="F369760CFFB5C3525E99F8F8FECB28B2" box="[243,329,1917,1940]" name="Micronesia" pageId="1" pageNumber="594">Ponape</collectingCountry>
; CAS 226297, 9.1 &amp; 12.1,
<collectingCountry id="F369760CFFB5C3525C13F8F8FD4D28B2" box="[633,719,1917,1940]" name="Micronesia" pageId="1" pageNumber="594">Ponape</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646517FFB5C3515EADF825FD812FDE" lastPageId="2" lastPageNumber="595" pageId="1" pageNumber="594" type="diagnosis">
<paragraph id="8BC1369CFFB5C3515EADF825FD812FDE" blockId="1.[151,1436,1664,2013]" lastBlockId="2.[151,1436,151,1007]" lastPageId="2" lastPageNumber="595" pageId="1" pageNumber="594">
<emphasis id="B90AEA8EFFB5C3525EADF825FEB8289F" bold="true" box="[199,314,1952,1977]" pageId="1" pageNumber="594">Diagnosis</emphasis>
. The following combination of characters distinguishes
<taxonomicName id="4C7E4D1FFFB5C3525DA9F824FBC1289E" box="[963,1091,1953,1976]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="594" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB5C3525DA9F824FBC1289E" box="[963,1091,1953,1976]" italics="true" pageId="1" pageNumber="594">E. santanai</emphasis>
</taxonomicName>
from congeners: the cephalic sensory-canal pore system lacks the IT and PITO pores, the AITO pore is not enlarged or paired, and the POP pores are small; some pectoral-fin rays are branched; the dorsal/anal formula is 8/8; and the body has six body bars with a black midcaudal peduncle spot centered on the last bar; no occipital spot; and color in life includes pinkishmauve bars that are red dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C3646517FFB6C3505EADFE81FBA22D46" lastPageId="3" lastPageNumber="596" pageId="2" pageNumber="595" type="description">
<paragraph id="8BC1369CFFB6C3515EADFE81FD4A2C31" blockId="2.[151,1436,151,1007]" pageId="2" pageNumber="595">
<emphasis id="B90AEA8EFFB6C3515EADFE81FEDB2E3B" bold="true" box="[199,345,260,285]" pageId="2" pageNumber="595">Description.</emphasis>
Dorsal-fin rays VI-I,8; anal-fin rays I,8; dorsal and anal soft rays branched except first, the last ray branched to base; pectoral-fin rays 16, ends of many rays broken in all
<typeStatus id="54C5883EFFB6C3515D9EFEADFBB02E66" box="[1012,1074,296,320]" pageId="2" pageNumber="595">types</typeStatus>
, but some lower rays branched; pelvic fins joined by membrane only at extreme base; no pelvic frenum; pelvic-fin membrane reduced, pelvic-fin rays I,4, 4th pelvic-fin ray with 4–7 branches, 2 segments between branches; 5th pelvic-fin ray absent; 11 branched caudal-fin rays; segmented caudal-fin rays 17; lateral scale rows 24; transverse scale rows 7; no scales on head, nape, breast, or pectoral-fin base; midline of abdomen with cycloid scales; dorsal fin triangular in shape, and the dorsal-fin spines not filamentous; cephalic sensory-pore system lacking IT and PITO pores but AITO not enlarged as in Group IV; cutaneous papilla system not discernible because specimens are in poor condition; male genital papilla non-fimbriate, smooth and straight, with a blunt tip with a few short skin flaps on the end, not reaching anal-fin base; single small female with a short papilla with a few short flaps at end (difficult to observe because of condition); body slender, front of head rounded with an angle of about 70° from the horizontal axis; mouth oblique, forming an angle of about 45° to horizontal axis of body, lower jaw not projecting; maxilla extending to center of pupil; anterior tubular nares short, just extending to upper lip and light in color; gill opening extending forward to midway between edge of preoperculum and posterior end of opercular membrane; gill membranes attached anteriorly to isthmus, without a free fold.
</paragraph>
<paragraph id="8BC1369CFFB6C3515EADFCA7FE152CC9" blockId="2.[151,1436,151,1007]" pageId="2" pageNumber="595">
<emphasis id="B90AEA8EFFB6C3515EADFCA7FEF42C1D" bold="true" box="[199,374,802,827]" pageId="2" pageNumber="595">Measurements</emphasis>
(based on
<typeStatus id="54C5883EFFB6C3515F96FCA7FDDC2C1D" box="[508,606,802,827]" pageId="2" pageNumber="595" type="holotype">holotype</typeStatus>
and 2
<typeStatus id="54C5883EFFB6C3515CDEFCA6FCA02C1D" box="[692,802,803,827]" pageId="2" pageNumber="595" type="paratype">paratypes</typeStatus>
,
<quantity id="4C869B79FFB6C3515D5BFCA7FC592C1D" box="[817,987,802,827]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" metricValueMax="1.28" metricValueMin="1.15" pageId="2" pageNumber="595" unit="mm" value="12.15" valueMax="12.8" valueMin="11.5">11.5–12.8 mm</quantity>
). Head length 28.8 (28.8–29.6, 29.1); origin of first dorsal fin 37.3 (37.3–39.1, 37.9), above posterior end of pectoral-fin base; origin of second dorsal fin 57.6 (56.2–58.3, 57.4), slightly in advance of anal-fin origin; origin of anal fin 58.5 (58.5–63.3, 60.1); caudalpeduncle length 25.4 (21.3–25.4, 23.2); caudal-peduncle of moderate depth, 12.3 (11.7–13.0, 12.3); body depth slender 19.5 (19.5–21.7, 20.2); eye diameter 9.7 (9.7–10.9, 10.5); snout length 3.4 (3.4–3.9, 3.6); upper-jaw length 10.6 (10.6–12.6, 11.5).
</paragraph>
<caption id="DF016614FFB6C3515EFDFA4BFAD42AC2" box="[151,1366,1486,1508]" httpUri="https://zenodo.org/record/223649/files/figure.png" pageId="2" pageNumber="595" targetBox="[151,1436,1060,1465]" targetPageId="2">
<paragraph id="8BC1369CFFB6C3515EFDFA4BFAD42AC2" blockId="2.[151,1366,1486,1508]" box="[151,1366,1486,1508]" pageId="2" pageNumber="595">
<emphasis id="B90AEA8EFFB6C3515EFDFA4BFE912AC5" bold="true" box="[151,275,1486,1508]" pageId="2" pageNumber="595">FIGURE 1.</emphasis>
Color in preservative of holotype of
<taxonomicName id="4C7E4D1FFFB6C3515CFBFA4AFC862AC2" box="[657,772,1487,1508]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="595" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB6C3515CFBFA4AFC862AC2" box="[657,772,1487,1508]" italics="true" pageId="2" pageNumber="595">E. santanai</emphasis>
</taxonomicName>
, CAS 234966, 11.8 mm. Photograph by D.W. Greenfield.
</paragraph>
</caption>
<paragraph id="8BC1369CFFB6C3515EADF989FB802863" blockId="2.[151,1436,1548,2005]" pageId="2" pageNumber="595">
<emphasis id="B90AEA8EFFB6C3515EADF989FDCC2903" bold="true" box="[199,590,1548,1573]" pageId="2" pageNumber="595">
Color in preservative of
<typeStatus id="54C5883EFFB6C3515F8DF989FDCC2903" box="[487,590,1548,1573]" pageId="2" pageNumber="595" type="holotype">holotype</typeStatus>
</emphasis>
: (
<figureCitation id="13452A19FFB6C3515C06F989FD292902" box="[620,683,1548,1573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1486,1508]" captionTargetBox="[151,1436,1060,1465]" captionTargetId="figure@2.[151,1436,1060,1465]" captionTargetPageId="2" captionText="FIGURE 1. Color in preservative of holotype of E. santanai, CAS 234966, 11.8 mm. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223649/files/figure.png" pageId="2" pageNumber="595">Fig. 1</figureCitation>
) Background color of head and body pale yellowish. Top of head with scattered irregular brown spots behind eyes, extending back to edge of preopercle. A few similar spots extending behind eye along top of cheek and preopercle, with a concentration towards the end of the preopercle. A small dark spot on lower portion of preopercle on left side of
<typeStatus id="54C5883EFFB6C3515D08F9F2FC4629B6" box="[866,964,1655,1680]" pageId="2" pageNumber="595" type="holotype">holotype</typeStatus>
(lacking in
<typeStatus id="54C5883EFFB6C3515A3BF9FDFB4329B6" box="[1105,1217,1656,1680]" pageId="2" pageNumber="595" type="paratype">paratypes</typeStatus>
). Interorbital area, snout, and ventral surface of head lacking any dark pigment. Pectoral-fin base and sides of body lacking dark pigment except for a distinct black spot about half the size of the eye centered on the caudal peduncle. Pectoral and pelvic fins immaculate. Anal fin heavily pigmented, darker than other fins. Ventral half of caudal fin peppered with small dark spots with a few extending dorsally at center of fin. Distal portion of first dorsal-fin margin with small dark spots as is the second dorsal fin. Iris of eye black, pupil yellowish.
</paragraph>
<paragraph id="8BC1369CFFB6C3505EADF8CAFBA22D46" blockId="2.[151,1436,1548,2005]" lastBlockId="3.[151,1436,151,608]" lastPageId="3" lastPageNumber="596" pageId="2" pageNumber="595">
<emphasis id="B90AEA8EFFB6C3515EADF8CAFEC2284E" bold="true" box="[199,320,1871,1896]" pageId="2" pageNumber="595">Live color</emphasis>
: (
<figureCitation id="13452A19FFB6C3515F30F8CAFE1D284E" box="[346,415,1871,1896]" captionStart="FIGURE 2" captionStartId="3.[151,250,1514,1536]" captionTargetBox="[151,1436,661,1493]" captionTargetId="figure@3.[151,1436,661,1493]" captionTargetPageId="3" captionText="FIGURE 2. Underwater photograph of E. santanai. Photograph by M. V. Erdmann." httpUri="https://zenodo.org/record/223650/files/figure.png" pageId="2" pageNumber="595">Fig. 2</figureCitation>
) Background color of head and body translucent white. Body with six internal pinkishmauve bars extending from backbone to ventral surface, the first two bars the widest, more triangular in shape, and most pronounced: first bar behind pectoral-fin base below nape and first four dorsal-fin spines; second bar below last two spines of first dorsal fin and space between the two dorsal fins; third bar below anterior half of second dorsal fin; fourth below second half of fin; fifth behind second dorsal fin; and sixth the narrowest and on the caudal peduncle with a distinct black spot on the midline. There is a crescent shaped red bar on the caudal-fin base. Each of these six internal bars have a red area over the backbone with two or three narrow red-orange bars extending up from them to the dorsal surface of body. These narrow red bars have some yellow pigment mixed with the red on the anterior half of the body. The translucent white area between these narrow dorsal bars has a very narrow vertical silvery-white line separating them. There also is a wider silvery-white bar separating the first two mauve bars ventrally. The nape is crossed by three yellow bars that are sprinkled with red. The area on top of the head and behind the eyes is bright red, with the red extending back along the backbone to under the first dorsal fin. The red also extends down onto the cheek and preoperculum. The red area on the top of the head with scattered silverywhite irregular spots that also are present on the top of the snout. Jaws with a tinge of red and area under the eye translucent white. The pupil of the eye is black and the iris red with yellow spokes radiating out from around the pupil and many small yellow spots dorsally. The operculum is yellow as is the pectoral-fin base, except that a silvery-white stripe runs across its center. Spines and rays of dorsal fins reddish.
</paragraph>
</subSubSection>
<caption id="DF016614FFB7C3505EFDFA6FFC6B2926" box="[151,1001,1514,1536]" httpUri="https://zenodo.org/record/223650/files/figure.png" pageId="3" pageNumber="596" targetBox="[151,1436,661,1493]" targetPageId="3">
<paragraph id="8BC1369CFFB7C3505EFDFA6FFC6B2926" blockId="3.[151,1001,1514,1536]" box="[151,1001,1514,1536]" pageId="3" pageNumber="596">
<emphasis id="B90AEA8EFFB7C3505EFDFA6FFE902AD9" bold="true" box="[151,274,1514,1536]" pageId="3" pageNumber="596">FIGURE 2.</emphasis>
Underwater photograph of
<taxonomicName id="4C7E4D1FFFB7C3505C58FA6EFD272926" box="[562,677,1515,1536]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505C58FA6EFD272926" box="[562,677,1515,1536]" italics="true" pageId="3" pageNumber="596">E. santanai</emphasis>
</taxonomicName>
. Photograph by M.V. Erdmann.
</paragraph>
</caption>
<subSubSection id="C3646517FFB7C3505EADF9A2FC3C2966" box="[199,958,1575,1600]" pageId="3" pageNumber="596" type="distribution">
<paragraph id="8BC1369CFFB7C3505EADF9A2FC3C2966" blockId="3.[151,1436,1575,2033]" box="[199,958,1575,1600]" pageId="3" pageNumber="596">
<emphasis id="B90AEA8EFFB7C3505EADF9A2FEE22966" bold="true" box="[199,352,1575,1600]" pageId="3" pageNumber="596">Distribution.</emphasis>
Known only from the northern coast of
<collectingCountry id="F369760CFFB7C3505D41F9A2FC3B2966" box="[811,953,1575,1600]" name="Timor-Leste" pageId="3" pageNumber="596">Timor-Leste</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646517FFB7C3565EADF9C9FD3A29DB" lastPageId="5" lastPageNumber="598" pageId="3" pageNumber="596" type="etymology">
<paragraph id="8BC1369CFFB7C3505EADF9C9FD1829F6" blockId="3.[151,1436,1575,2033]" pageId="3" pageNumber="596">
<emphasis id="B90AEA8EFFB7C3505EADF9C9FECC2943" bold="true" box="[199,334,1612,1637]" pageId="3" pageNumber="596">Etymology.</emphasis>
The specific epithet is named in honor of Connisso Antonino (&quot;Nino Konis&quot;) Santana, a national hero in Timor-Leste's struggle for independence who was renowned for his environmental awareness. The
<typeStatus id="54C5883EFFB7C3505B06F9F5FA1E29AE" box="[1388,1436,1648,1672]" pageId="3" pageNumber="596">type</typeStatus>
locality of this beautiful goby species is in Tutuala, just offshore of Santana's birthplace, and moreover is located within the Nino Konis Santana National Park.
</paragraph>
<paragraph id="8BC1369CFFB7C3505EADF959FC2E28D6" blockId="3.[151,1436,1575,2033]" pageId="3" pageNumber="596">
<emphasis id="B90AEA8EFFB7C3505EADF959FEE729D3" bold="true" box="[199,357,1756,1781]" pageId="3" pageNumber="596">Comparisons</emphasis>
. In general live color pattern,
<taxonomicName id="4C7E4D1FFFB7C3505CA2F958FCCB29D2" box="[712,841,1757,1780]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505CA2F958FCCB29D2" box="[712,841,1757,1780]" italics="true" pageId="3" pageNumber="596">E. santanai</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C7E4D1FFFB7C3505A75F958FB3F29D2" box="[1055,1213,1756,1780]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB7C3505A75F958FB3F29D2" box="[1055,1213,1756,1780]" italics="true" pageId="3" pageNumber="596">E. latifasciata</emphasis>
</taxonomicName>
and also shares the same meristics. It does, however, differ in cephalic sensory-canal pore pattern. Whereas
<taxonomicName id="4C7E4D1FFFB7C3505AEEF884FAA3283E" box="[1156,1313,1792,1816]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB7C3505AEEF884FAA3283E" box="[1156,1313,1792,1816]" italics="true" pageId="3" pageNumber="596">E. latifasciata</emphasis>
</taxonomicName>
belongs to Group II, only lacking the IT pore,
<taxonomicName id="4C7E4D1FFFB7C3505C4DF8A0FD24281A" box="[551,678,1829,1852]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505C4DF8A0FDBA281A" box="[551,568,1829,1852]" italics="true" pageId="3" pageNumber="596">E</emphasis>
.
<emphasis id="B90AEA8EFFB7C3505C2CF8A0FD24281A" box="[582,678,1829,1852]" italics="true" pageId="3" pageNumber="596">santanai</emphasis>
</taxonomicName>
also lacks the PITO pore and the POP pores are very small. Other
<taxonomicName id="4C7E4D1FFFB7C3505EFDF8CDFF622879" box="[151,224,1864,1887]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="B90AEA8EFFB7C3505EFDF8CDFF622879" box="[151,224,1864,1887]" italics="true" pageId="3" pageNumber="596">Eviota</emphasis>
</taxonomicName>
species that lack both the PITO and IT pores have the AITO pore enlarged or paired (Group IV), but this is not the case in
<taxonomicName id="4C7E4D1FFFB7C3505F24F8E8FE5328A2" box="[334,465,1901,1924]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505F24F8E8FEDD28A2" box="[334,351,1901,1924]" italics="true" pageId="3" pageNumber="596">E</emphasis>
.
<emphasis id="B90AEA8EFFB7C3505F1BF8E8FE5328A2" box="[369,465,1901,1924]" italics="true" pageId="3" pageNumber="596">santanai</emphasis>
</taxonomicName>
. There are only three species in Group IV that also have the same meristics as
<taxonomicName id="4C7E4D1FFFB7C3505BEEF8E8FF752881" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505BEEF8E8FF752881" italics="true" pageId="3" pageNumber="596">E. santanai</emphasis>
</taxonomicName>
:
<taxonomicName id="4C7E4D1FFFB7C3505F63F814FE10288E" box="[265,402,1936,1960]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="shimadai">
<emphasis id="B90AEA8EFFB7C3505F63F814FE10288E" box="[265,402,1936,1960]" italics="true" pageId="3" pageNumber="596">E. shimadai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E4D1FFFB7C3505FC9F814FDCA288E" box="[419,584,1936,1960]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="lachdeberei">
<emphasis id="B90AEA8EFFB7C3505FC9F814FDCA288E" box="[419,584,1936,1960]" italics="true" pageId="3" pageNumber="596">E. lachdeberei</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7E4D1FFFB7C3505CE1F814FCCE288E" box="[651,844,1936,1960]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="partimaculata">
<emphasis id="B90AEA8EFFB7C3505CE1F814FCCE288E" box="[651,844,1936,1960]" italics="true" pageId="3" pageNumber="596">E. partimaculata</emphasis>
</taxonomicName>
, but all of these have unbranched pectoral-fin rays (branched in
<taxonomicName id="4C7E4D1FFFB7C3505F41F830FE2828EA" box="[299,426,1973,1996]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505F41F830FEBE28EA" box="[299,316,1973,1996]" italics="true" pageId="3" pageNumber="596">E</emphasis>
.
<emphasis id="B90AEA8EFFB7C3505F20F830FE2828EA" box="[330,426,1973,1996]" italics="true" pageId="3" pageNumber="596">santanai</emphasis>
</taxonomicName>
).
<taxonomicName id="4C7E4D1FFFB7C3505FA2F830FDFB28EA" box="[456,633,1973,1996]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB7C3505FA2F830FDFB28EA" box="[456,633,1973,1996]" italics="true" pageId="3" pageNumber="596">Eviota santanai</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C7E4D1FFFB7C3505D2BF830FC5C28EA" box="[833,990,1972,1996]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB7C3505D2BF830FC5C28EA" box="[833,990,1972,1996]" italics="true" pageId="3" pageNumber="596">E. latifasciata</emphasis>
</taxonomicName>
by lacking an occipital spot and having a pigment spot behind the eye that is lacking in
<taxonomicName id="4C7E4D1FFFB7C3505CC4F85CFCC928D6" box="[686,843,2008,2032]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="596" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB7C3505CC4F85CFCC928D6" box="[686,843,2008,2032]" italics="true" pageId="3" pageNumber="596">E. latifasciata</emphasis>
</taxonomicName>
(
<figureCitation id="13452A19FFB7C3505D30F85DFC1C28D6" box="[858,926,2008,2033]" captionStart="FIGURE 3" captionStartId="4.[151,250,1959,1981]" captionTargetBox="[202,1390,210,1933]" captionTargetId="figure@4.[195,1400,193,1947]" captionTargetPageId="4" captionText="FIGURE 3. Head coloration of preserved specimens, with pigmentation differences indicated by arrows. A. Eviota santanai holotype, CAS 234966, 11.8 mm. B. Eviota latifasciata paratype, USNM 260079, 12.7 mm. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223651/files/figure.png" pageId="3" pageNumber="596">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF016614FFB0C3575EFDF822FA1A28FA" httpUri="https://zenodo.org/record/223651/files/figure.png" pageId="4" pageNumber="597" targetBox="[202,1390,210,1933]" targetPageId="4">
<paragraph id="8BC1369CFFB0C3575EFDF822FA1A28FA" blockId="4.[151,1435,1959,2012]" pageId="4" pageNumber="597">
<emphasis id="B90AEA8EFFB0C3575EFDF822FE91289A" bold="true" box="[151,275,1959,1981]" pageId="4" pageNumber="597">FIGURE 3.</emphasis>
Head coloration of preserved specimens, with pigmentation differences indicated by arrows. A.
<taxonomicName id="4C7E4D1FFFB0C3575A96F82DFA19289B" box="[1276,1435,1960,1981]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="597" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB0C3575A96F82DFA19289B" box="[1276,1435,1960,1981]" italics="true" pageId="4" pageNumber="597">Eviota santanai</emphasis>
</taxonomicName>
holotype, CAS 234966, 11.8 mm. B.
<taxonomicName id="4C7E4D1FFFB0C3575C7CF842FD5228FA" box="[534,720,1990,2012]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="597" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB0C3575C7CF842FD5228FA" box="[534,720,1990,2012]" italics="true" pageId="4" pageNumber="597">Eviota latifasciata</emphasis>
</taxonomicName>
paratype, USNM 260079, 12.7 mm. Photograph by D.W. Greenfield.
</paragraph>
</caption>
<paragraph id="8BC1369CFFB1C3565EADFF12FBAA2DC9" blockId="5.[151,1436,151,752]" pageId="5" pageNumber="598">
The
<typeStatus id="54C5883EFFB1C3565E94FF12FEE22F96" box="[254,352,151,176]" pageId="5" pageNumber="598" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7E4D1FFFB1C3565FE5FF1DFDE02F96" box="[399,610,152,176]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565FE5FF1DFDE02F96" box="[399,610,152,176]" italics="true" pageId="5" pageNumber="598">Eviota latifasciata</emphasis>
</taxonomicName>
was originally described from Abaiang Atoll in the
<collectingCountry id="F369760CFFB1C3565ABCFF12FA022F96" box="[1238,1408,151,176]" name="Kiribati" pageId="5" pageNumber="598">Gilbert Islands</collectingCountry>
= (
<collectingCountry id="F369760CFFB1C3565EF4FF39FF7C2FF2" box="[158,254,188,212]" name="Kiribati" pageId="5" pageNumber="598">Tungaru</collectingCountry>
Islands,
<collectingCountry id="F369760CFFB1C3565F07FF39FE442FF3" box="[365,454,188,213]" name="Kiribati" pageId="5" pageNumber="598">Kiribati</collectingCountry>
) with
<typeStatus id="54C5883EFFB1C3565C72FF38FD012FF3" box="[536,643,189,213]" pageId="5" pageNumber="598" type="paratype">paratypes</typeStatus>
from both Abaiang and
<collectingCountry id="F369760CFFB1C3565DDBFF38FB862FF2" box="[945,1028,189,212]" name="Micronesia" pageId="5" pageNumber="598">Ponape</collectingCountry>
in the
<collectingCountry id="F369760CFFB1C3565A0AFF39FA9E2FF3" box="[1120,1308,188,213]" name="Micronesia" pageId="5" pageNumber="598">Caroline Islands</collectingCountry>
(Jewett &amp; Lachner, 1983). Non-type material was listed from
<collectingCountry id="F369760CFFB1C3565C9FFF5AFC342FDE" box="[757,950,223,248]" name="Christmas Island" pageId="5" pageNumber="598">Christmas Island</collectingCountry>
in the Indian Ocean and Kapingamarangi Atoll. Since that time
<taxonomicName id="4C7E4D1FFFB1C3565FF1FE80FDB82E3A" box="[411,570,260,284]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565FF1FE80FDB82E3A" box="[411,570,260,284]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
has also been reported by Allen &amp; Erdmann (2012) from “
<collectingCountry id="F369760CFFB1C3565AB1FE81FA1E2E3B" box="[1243,1436,260,285]" name="Christmas Island" pageId="5" pageNumber="598">Christmas Island</collectingCountry>
(Indian Ocean),
<collectingCountry id="F369760CFFB1C3565F3AFEA2FE3F2E66" box="[336,445,295,320]" name="Indonesia" pageId="5" pageNumber="598">Indonesia</collectingCountry>
(Lesser Sunda Islands, Banda, Halmahera, and West
<collectingCountry id="F369760CFFB1C3565A75FEADFBEA2E66" box="[1055,1128,296,320]" name="Papua New Guinea" pageId="5" pageNumber="598">Papua</collectingCountry>
),
<collectingCountry id="F369760CFFB1C3565A17FEA2FB792E66" box="[1149,1275,295,320]" name="Philippines" pageId="5" pageNumber="598">Philippines</collectingCountry>
(Palawan and Cebu),
<collectingCountry id="F369760CFFB1C3565E80FEC8FE462E43" box="[234,452,332,357]" name="Papua New Guinea" pageId="5" pageNumber="598">Papua New Guinea</collectingCountry>
(Milne Bay),
<collectingCountry id="F369760CFFB1C3565C0FFEC9FD242E43" box="[613,678,332,357]" name="Palau" pageId="5" pageNumber="598">Palau</collectingCountry>
,
<collectingCountry id="F369760CFFB1C3565CD8FEC8FC852E42" box="[690,775,333,356]" name="Micronesia" pageId="5" pageNumber="598">Ponape</collectingCountry>
, and
<collectingCountry id="F369760CFFB1C3565D2EFEC9FC682E43" box="[836,1002,332,357]" name="Kiribati" pageId="5" pageNumber="598">Gilbert Islands</collectingCountry>
(
<collectingCountry id="F369760CFFB1C3565D90FEC9FBDA2E43" box="[1018,1112,332,357]" name="Kiribati" pageId="5" pageNumber="598">Kiribati</collectingCountry>
). ” It was also reported from the Ryukyu Islands,
<collectingCountry id="F369760CFFB1C3565FE8FEF5FE402EA1" box="[386,450,368,391]" name="Japan" pageId="5" pageNumber="598">Japan</collectingCountry>
by Senou
<emphasis id="B90AEA8EFFB1C3565C55FEF4FDFB2EAE" box="[575,633,368,392]" italics="true" pageId="5" pageNumber="598">et al.</emphasis>
(2004), and
<collectingCountry id="F369760CFFB1C3565D64FEF5FCD12EAE" box="[782,851,368,392]" name="Tonga" pageId="5" pageNumber="598">Tonga</collectingCountry>
by Randall,
<emphasis id="B90AEA8EFFB1C3565D83FEF4FBA12EAE" box="[1001,1059,368,392]" italics="true" pageId="5" pageNumber="598">et al.</emphasis>
(2003). Yet photographs of live individuals from many of these locations show considerable variation (based on photographs provided by G.R. Allen), and as discussed by Allen, Brooks &amp; Erdmann (2013), “Live colour patterns in particular (including eye colouration) are highly diagnostic for members of this genus...”. D.F. Hoese provided color photographs of specimens from the
<typeStatus id="54C5883EFFB1C3565FEBFD85FE332D3E" box="[385,433,512,536]" pageId="5" pageNumber="598">type</typeStatus>
series of
<taxonomicName id="4C7E4D1FFFB1C3565C4CFD84FD442D3E" box="[550,710,512,536]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565C4CFD84FD442D3E" box="[550,710,512,536]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
from the
<collectingCountry id="F369760CFFB1C3565D56FE7AFC642D3E" box="[828,998,511,536]" name="Kiribati" pageId="5" pageNumber="598">Gilbert Islands</collectingCountry>
, the locality of the
<typeStatus id="54C5883EFFB1C3565AA1FE7AFAB22D3E" box="[1227,1328,511,536]" pageId="5" pageNumber="598" type="holotype">holotype</typeStatus>
, (
<figureCitation id="13452A19FFB1C3565B2CFE7AFA0F2D3E" box="[1350,1421,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1153,1175]" captionTargetBox="[283,1321,805,1131]" captionTargetId="figure@5.[265,1321,805,1131]" captionTargetPageId="5" captionText="FIGURE 4. Fresh specimen from the type series of E. latifasciata from Abaiang Atoll, Gilbert Islands, photograph by D. F. Hoese (DFH 73 - 36)." httpUri="https://zenodo.org/record/223652/files/figure.png" pageId="5" pageNumber="598">Fig. 4</figureCitation>
), another photograph of a specimen from the
<typeStatus id="54C5883EFFB1C3565CEEFDA0FD362D1B" box="[644,692,549,573]" pageId="5" pageNumber="598">type</typeStatus>
locality (
<figureCitation id="13452A19FFB1C3565D75FDA1FCE12D1B" box="[799,867,548,573]" captionStart="FIGURE 5" captionStartId="5.[151,250,1565,1587]" captionTargetBox="[265,1321,1255,1543]" captionTargetId="figure@5.[265,1321,1255,1543]" captionTargetPageId="5" captionText="FIGURE 5. Fresh specimen of E. latifasciata from Abaiang Atoll, Gilbert Islands, AMS I. 18048 - 026 (DFH 73 - 45), photograph by D. F. Hoese." httpUri="https://zenodo.org/record/223653/files/figure.png" pageId="5" pageNumber="598">Fig. 5</figureCitation>
), and G.R. Allen provided color photographs from
<collectingCountry id="F369760CFFB1C3565EFDFDCDFF682D79" box="[151,234,584,607]" name="Micronesia" pageId="5" pageNumber="598">Ponape</collectingCountry>
in
<collectingCountry id="F369760CFFB1C3565F7AFDC2FE0C2D46" box="[272,398,583,608]" name="Micronesia" pageId="5" pageNumber="598">Micronesia</collectingCountry>
to the northeast where some
<typeStatus id="54C5883EFFB1C3565CB4FDCDFCC82D46" box="[734,842,584,608]" pageId="5" pageNumber="598" type="paratype">paratypes</typeStatus>
were taken (
<figureCitation id="13452A19FFB1C3565DB4FDC2FBA12D46" box="[990,1059,583,608]" captionStart="FIGURE 6. C" captionStartId="6.[151,250,1029,1051]" captionTargetBox="[151,1436,373,1008]" captionTargetId="figure@6.[151,1436,373,1008]" captionTargetPageId="6" captionText="FIGURE 6. C. Underwater photograph of E. latifasciata, Ponape, photograph by G. R. Allen. Note parasite on abdomen." httpUri="https://zenodo.org/record/223654/files/figure.png" pageId="5" pageNumber="598">Fig. 6</figureCitation>
). The color of the body bars are pinkish-mauve in
<taxonomicName id="4C7E4D1FFFB1C3565F07FDE8FE712DA2" box="[365,499,621,644]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB1C3565F07FDE8FE712DA2" box="[365,499,621,644]" italics="true" pageId="5" pageNumber="598">E. santanai</emphasis>
</taxonomicName>
, whereas they are dark red in the Gilbert
<collectingCountry id="F369760CFFB1C3565D92FDE9FBBF2DA3" box="[1016,1085,620,645]" name="Iceland" pageId="5" pageNumber="598">Island</collectingCountry>
and
<collectingCountry id="F369760CFFB1C3565A15FDE8FB502DA2" box="[1151,1234,621,644]" name="Micronesia" pageId="5" pageNumber="598">Ponape</collectingCountry>
specimens of
<taxonomicName id="4C7E4D1FFFB1C3565BEEFDE8FE972D8E" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565BEEFDE8FE972D8E" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
. The
<typeStatus id="54C5883EFFB1C3565F34FD15FE4B2D8E" box="[350,457,656,680]" pageId="5" pageNumber="598" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C7E4D1FFFB1C3565F9AFD14FD0F2D8E" box="[496,653,656,680]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565F9AFD14FD0F2D8E" box="[496,653,656,680]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
examined from the
<collectingCountry id="F369760CFFB1C3565D1BFD0AFB952D8E" box="[881,1047,655,680]" name="Kiribati" pageId="5" pageNumber="598">Gilbert Islands</collectingCountry>
have a definite dark occipital spot even after having been preserved for 40 years (
<figureCitation id="13452A19FFB1C3565CA1FD31FC972DEA" box="[715,789,692,717]" captionStart="FIGURE 7" captionStartId="6.[151,250,1464,1486]" captionTargetBox="[151,1436,1101,1443]" captionTargetId="figure@6.[151,1436,1101,1443]" captionTargetPageId="6" captionText="FIGURE 7. Color in preservative of paratype of E. latifasciata, BPBM 28959, 11.7 mm, Abaiang Atoll, Gilbert Islands. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223655/files/figure.png" pageId="5" pageNumber="598">Fig. 7</figureCitation>
), as does the comparative material from
<collectingCountry id="F369760CFFB1C3565A95FD30FAD72DEA" box="[1279,1365,693,716]" name="Micronesia" pageId="5" pageNumber="598">Ponape</collectingCountry>
. The specimens of
<taxonomicName id="4C7E4D1FFFB1C3565F5BFD5CFE322DC9" box="[305,432,728,752]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB1C3565F5BFD5CFE322DC9" box="[305,432,728,752]" italics="true" pageId="5" pageNumber="598">E. santanai</emphasis>
</taxonomicName>
collected in 2012 do not have an obvious occipital spot.
</paragraph>
<caption id="DF016614FFB1C3565EFDFB04FEEA2B93" httpUri="https://zenodo.org/record/223652/files/figure.png" pageId="5" pageNumber="598" targetBox="[283,1321,805,1131]" targetPageId="5">
<paragraph id="8BC1369CFFB1C3565EFDFB04FEEA2B93" blockId="5.[151,1436,1153,1205]" pageId="5" pageNumber="598">
<emphasis id="B90AEA8EFFB1C3565EFDFB04FE962BB0" bold="true" box="[151,276,1153,1175]" pageId="5" pageNumber="598">FIGURE 4.</emphasis>
Fresh specimen from the type series of
<taxonomicName id="4C7E4D1FFFB1C3565CD4FB07FCCC2BB1" box="[702,846,1153,1175]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565CD4FB07FCCC2BB1" box="[702,846,1153,1175]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
from Abaiang Atoll, Gilbert Islands, photograph by D.F. Hoese (DFH 73-36).
</paragraph>
</caption>
<caption id="DF016614FFB1C3565EFDF998FE2A2974" httpUri="https://zenodo.org/record/223653/files/figure.png" pageId="5" pageNumber="598" targetBox="[265,1321,1255,1543]" targetPageId="5">
<paragraph id="8BC1369CFFB1C3565EFDF998FE2A2974" blockId="5.[151,1435,1565,1618]" pageId="5" pageNumber="598">
<emphasis id="B90AEA8EFFB1C3565EFDF998FE9B2914" bold="true" box="[151,281,1565,1587]" pageId="5" pageNumber="598">FIGURE 5.</emphasis>
Fresh specimen of
<taxonomicName id="4C7E4D1FFFB1C3565C6FF99BFD182915" box="[517,666,1565,1587]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565C6FF99BFD182915" box="[517,666,1565,1587]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
from Abaiang Atoll, Gilbert Islands, AMS I.18048-026 (DFH 73-45), photograph by D.F. Hoese.
</paragraph>
</caption>
<paragraph id="8BC1369CFFB1C3565EADF9FCFD3A29DB" blockId="5.[151,1436,1657,2006]" pageId="5" pageNumber="598">
Recently
<taxonomicName id="4C7E4D1FFFB1C3565F5EF9FFFE7529B7" box="[308,503,1657,1681]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="dorsimaculata">
<emphasis id="B90AEA8EFFB1C3565F5EF9FFFE7529B7" box="[308,503,1657,1681]" italics="true" pageId="5" pageNumber="598">E. dorsimaculata</emphasis>
</taxonomicName>
was described from the Marquesas Islands and was said to be most similar to
<taxonomicName id="4C7E4D1FFFB1C3565BEEF9FFFDAD2993" authority="Tornabene et al. 2013" authorityName="Tornabene et al." authorityYear="2013" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565BEEF9FFFE972993" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
(Tornabene
<emphasis id="B90AEA8EFFB1C3565FCDF91BFE632993" box="[423,481,1693,1717]" italics="true" pageId="5" pageNumber="598">et al.</emphasis>
2013)
</taxonomicName>
. This species also belongs to Group II, only lacking the IT pore as does
<taxonomicName id="4C7E4D1FFFB1C3565BEEF91BFE9729FF" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565BEEF91BFE9729FF" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
, separating it from
<taxonomicName id="4C7E4D1FFFB1C3565F9BF947FDFC29FF" box="[497,638,1730,1753]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="sanatanai">
<emphasis id="B90AEA8EFFB1C3565F9BF947FDFC29FF" box="[497,638,1730,1753]" italics="true" pageId="5" pageNumber="598">E. sanatanai</emphasis>
</taxonomicName>
. It also has a dark spot on the caudal peduncle that is above the lateral midline (centered on the midline in
<taxonomicName id="4C7E4D1FFFB1C3565C42F963FD2529DB" box="[552,679,1766,1789]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB1C3565C42F963FDBB29DB" box="[552,569,1766,1789]" italics="true" pageId="5" pageNumber="598">E</emphasis>
.
<emphasis id="B90AEA8EFFB1C3565C2DF963FD2529DB" box="[583,679,1766,1789]" italics="true" pageId="5" pageNumber="598">santanai</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646517FFB1C3555EADF88CFD2C29E5" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598" type="discussion">
<paragraph id="8BC1369CFFB1C3555EADF88CFD882E66" blockId="5.[151,1436,1657,2006]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598">
<emphasis id="B90AEA8EFFB1C3565EADF88CFEB92804" bold="true" box="[199,315,1801,1826]" pageId="5" pageNumber="598">Remarks.</emphasis>
The similarity of
<taxonomicName id="4C7E4D1FFFB1C3565C66F88FFD0F2807" box="[524,653,1802,1825]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="santanai">
<emphasis id="B90AEA8EFFB1C3565C66F88FFD9F2807" box="[524,541,1802,1825]" italics="true" pageId="5" pageNumber="598">E</emphasis>
.
<emphasis id="B90AEA8EFFB1C3565C46F88FFD0F2807" box="[556,653,1802,1825]" italics="true" pageId="5" pageNumber="598">santanai</emphasis>
</taxonomicName>
to
<taxonomicName id="4C7E4D1FFFB1C3565CDAF88FFCCF2807" box="[688,845,1801,1825]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB1C3565CDAF88FFCCF2807" box="[688,845,1801,1825]" italics="true" pageId="5" pageNumber="598">E. latifasciata</emphasis>
</taxonomicName>
presents another example of a species that is similar in color pattern to another species, but differs in the absence of a head pore. Greenfield and Randall (2010) noted that
<taxonomicName id="4C7E4D1FFFB1C3565EA2F8D7FECE284F" box="[200,332,1873,1897]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="toshiyuki">
<emphasis id="B90AEA8EFFB1C3565EA2F8D7FECE284F" box="[200,332,1873,1897]" italics="true" pageId="5" pageNumber="598">E. toshiyuki</emphasis>
</taxonomicName>
, known only from
<collectingCountry id="F369760CFFB1C3565C4FF8D7FDEA284F" box="[549,616,1874,1897]" name="Japan" pageId="5" pageNumber="598">Japan</collectingCountry>
, was very similar in color to
<taxonomicName id="4C7E4D1FFFB1C3565DC5F8D7FB9E284F" box="[943,1052,1874,1897]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="598" phylum="Chordata" rank="species" species="zonura">
<emphasis id="B90AEA8EFFB1C3565DC5F8D7FB9E284F" box="[943,1052,1874,1897]" italics="true" pageId="5" pageNumber="598">E. zonura</emphasis>
</taxonomicName>
, known from
<collectingCountry id="F369760CFFB1C3565AD6F8D4FAA1284C" box="[1212,1315,1873,1898]" name="Australia" pageId="5" pageNumber="598">Australia</collectingCountry>
east to the
<collectingCountry id="F369760CFFB1C3565EFDF8F0FEC928A8" box="[151,331,1909,1934]" name="Northern Mariana Islands" pageId="5" pageNumber="598">Mariana Islands</collectingCountry>
and
<collectingCountry id="F369760CFFB1C3565FEFF8F0FE5628AB" box="[389,468,1909,1933]" name="Samoa" pageId="5" pageNumber="598">Samoa</collectingCountry>
, but differs in the absence of a head pore, and suggested that this might indicate that they were sister species. Earlier Lachner &amp; Karnella (1980) suggested that cephalic sensory-canal pore systems might represent natural groupings of species, and continued to present new groupings in subsequent papers. These groupings have served useful in distinguishing the many different
<taxonomicName id="4C7E4D1FFFB2C3555DEEFF1DFC4F2F89" box="[900,973,152,175]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="599" phylum="Chordata" rank="genus">
<emphasis id="B90AEA8EFFB2C3555DEEFF1DFC4F2F89" box="[900,973,152,175]" italics="true" pageId="6" pageNumber="599">Eviota</emphasis>
</taxonomicName>
species from one another, particularly in dichotomous keys; however, in a recent paper Tornabene
<emphasis id="B90AEA8EFFB2C3555D52FF38FCF62FF2" box="[824,884,188,212]" italics="true" pageId="6" pageNumber="599">et al.</emphasis>
(2013) demonstrated that pore patterns did not correlate with genetic relationships in the species they studied. Unexpectedly the correlation was with branching patterns in the pectoral-fin rays. Thus, the suspected sister species mentioned above that differed in pore patterns might actually be closely related.
</paragraph>
<caption id="DF016614FFB2C3555EFDFB80FAEB2B3D" box="[151,1385,1029,1051]" httpUri="https://zenodo.org/record/223654/files/figure.png" pageId="6" pageNumber="599" targetBox="[151,1436,373,1008]" targetPageId="6">
<paragraph id="8BC1369CFFB2C3555EFDFB80FAEB2B3D" blockId="6.[151,1385,1029,1051]" box="[151,1385,1029,1051]" pageId="6" pageNumber="599">
<emphasis id="B90AEA8EFFB2C3555EFDFB80FE902B3C" bold="true" box="[151,274,1029,1051]" pageId="6" pageNumber="599">FIGURE 6.</emphasis>
C. Underwater photograph of
<taxonomicName id="4C7E4D1FFFB2C3555C25FB83FD5F2B3D" box="[591,733,1029,1051]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="599" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB2C3555C25FB83FD5F2B3D" box="[591,733,1029,1051]" italics="true" pageId="6" pageNumber="599">E. latifasciata</emphasis>
</taxonomicName>
, Ponape, photograph by G.R. Allen. Note parasite on abdomen.
</paragraph>
</caption>
<caption id="DF016614FFB2C3555EFDFA3DFE622ACB" httpUri="https://zenodo.org/record/223655/files/figure.png" pageId="6" pageNumber="599" targetBox="[151,1436,1101,1443]" targetPageId="6">
<paragraph id="8BC1369CFFB2C3555EFDFA3DFE622ACB" blockId="6.[151,1436,1464,1517]" pageId="6" pageNumber="599">
<emphasis id="B90AEA8EFFB2C3555EFDFA3DFE942AE8" bold="true" box="[151,278,1464,1486]" pageId="6" pageNumber="599">FIGURE 7.</emphasis>
Color in preservative of paratype of
<taxonomicName id="4C7E4D1FFFB2C3555CC7FA3FFCBC2AE8" box="[685,830,1464,1487]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="599" phylum="Chordata" rank="species" species="latifasciata">
<emphasis id="B90AEA8EFFB2C3555CC7FA3FFCBC2AE8" box="[685,830,1464,1487]" italics="true" pageId="6" pageNumber="599">E. latifasciata</emphasis>
</taxonomicName>
, BPBM 28959, 11.7 mm, Abaiang Atoll, Gilbert Islands. Photograph by D.W. Greenfield.
</paragraph>
</caption>
<paragraph id="8BC1369CFFB2C3555EADF99FFD2C29E5" blockId="6.[151,1436,1562,1731]" pageId="6" pageNumber="599">
Both the
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and
<typeStatus id="54C5883EFFB2C3555FA7F99EFDAF2915" box="[461,557,1563,1587]" pageId="6" pageNumber="599" type="paratype">paratype</typeStatus>
specimens were collected from similar habitats consisting of shallow, wavewashed fringing reef crest in
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depth. Both sites were exposed to clear, relatively cool (25-27
<emphasis id="B90AEA8EFFB2C3555A86F9B8FB752970" bold="true" box="[1260,1271,1597,1622]" pageId="6" pageNumber="599">°</emphasis>
C) water with reefs that quickly dropped off to over
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depth. The species was moreover observed at an additional four sites along the
<collectingCountry id="F369760CFFB2C3555F63F900FE1629B8" box="[265,404,1669,1694]" name="Timor-Leste" pageId="6" pageNumber="599">Timor-Leste</collectingCountry>
coastline (recorded at a total of 6 of the 20 survey sites) and is thus considered relatively common (though cryptic and shy) in the region.
</paragraph>
</subSubSection>
</treatment>
</document>
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