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<document id="A96352E96178F829A4A12C91489947B1" ID-DOI="10.5281/zenodo.5741842" ID-ISSN="1536-9307" ID-Zenodo-Dep="5741842" ID-ZooBank="DCAC2354-0168-4A66-AC36-87F1BB19EAA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638266231204" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Stockmann, Mark &amp; Šťáhlavský, František" docDate="2020" docId="038C87D0FF985010FEC3FC21FBB02924" docLanguage="en" docName="Euscorpius.325.1-140.pdf" docOrigin="Euscorpius 325" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Scorpiops scheibeae Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský 2020, sp. n." docType="treatment" docUuid="11191FD9-ADA9-4FF9-94D5-86127C76D660" docUuidSource="ZooBank" docVersion="3" lastPageNumber="96" masterDocId="FFB5FFA8FFF95071FFDFFFEDFFD72E46" masterDocTitle="Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida Scorpiones)" masterLastPageNumber="140" masterPageNumber="1" pageNumber="96" updateTime="1699436806083" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6A40E0D8F57C490EA64BEA8C185E9206">Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida Scorpiones)</mods:title>
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<figureCitation id="131E2A43FF985010FEDBFC03FE6B2A40" box="[260,444,1006,1030]" captionStart-0="Figures 634–638" captionStart-1="Figures 639–642" captionStart-2="Figures 643–646" captionStart-3="Figures 647–652" captionStartId-0="98.[115,192,1984,2006]" captionStartId-1="99.[118,195,1977,1999]" captionStartId-2="100.[117,194,2001,2023]" captionStartId-3="101.[116,193,1958,1980]" captionTargetBox-0="[121,1464,255,1950]" captionTargetBox-1="[121,1470,280,1870]" captionTargetBox-2="[120,1465,260,1965]" captionTargetBox-3="[113,1474,255,1921]" captionTargetId-0="figure-13@98.[113,1474,234,1958]" captionTargetId-1="figure-8@99.[113,1474,263,1903]" captionTargetId-2="figure-14@100.[113,1474,234,1979]" captionTargetId-3="figure-9@101.[113,1474,255,1923]" captionTargetPageId-0="98" captionTargetPageId-1="99" captionTargetPageId-2="100" captionTargetPageId-3="101" captionText-0="Figures 634–638: Scorpiops scheibeae sp. n., male holotype. Figures 634–635. Dorsal (634) and ventral (635) views. Figures 636–638. Metasoma and telson lateral (636), ventral (637), and dorsal (638) views. Scale bars: 10 mm (634–635, 636–638)." captionText-1="Figures 639–642: Scorpiops scheibeae sp. n., male holotype, pedipalp segments. Chela and patell dorsal (639), external (640) and ventral (641) views. Movable finger dentition (642). Trichobothrial pattern is indicated by white circles (639–641)." captionText-2="Figures 643–646. Scorpiops scheibeae sp. n., male holotype, left legs I–IV, prolateral/ ventral aspect." captionText-3="Figures 647–652: Scorpiops scheibeae sp. n., male holotype. Figures 647–648. Carapace and tergites I–III (647), coxosternal area and sternites III–IV (648), left chelicera in dorsal (649) and ventral (650) views. Figure 651. Telson lateral view. Figure 652. Male holotype in vivo habitus." figureDoi-0="http://doi.org/10.5281/zenodo.5741991" figureDoi-1="http://doi.org/10.5281/zenodo.5741993" figureDoi-2="http://doi.org/10.5281/zenodo.5741995" figureDoi-3="http://doi.org/10.5281/zenodo.5741997" httpUri-0="https://zenodo.org/record/5741991/files/figure.png" httpUri-1="https://zenodo.org/record/5741993/files/figure.png" httpUri-2="https://zenodo.org/record/5741995/files/figure.png" httpUri-3="https://zenodo.org/record/5741997/files/figure.png" pageId="97" pageNumber="96">Figures 634–652</figureCitation>
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, 9)
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<paragraph id="8B9A36C6FF985010FFAEFA5CFDAA284F" blockId="97.[113,772,1457,1545]" pageId="97" pageNumber="96">ETYMOLOGY. The species epithet is a patronym honoring Renate Scheibe from University Osnadrück who assisted one of authors (MS) with a venom extraction project.</paragraph>
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DIAGNOSIS (
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). Total length
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. Base color uniformly reddish black to black. Telson and tarsomere II of legs reddish brown. Pectine teeth number
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male; fulcra reduced to absent; 3 marginal and 5 middle lamellae. Patella of pedipalp with 58 external and 24 ventral trichobothria. Chela of pedipalp with 14
<emphasis id="B951EAD4FF985010FF12F93FFF0A28AF" box="[205,221,1746,1769]" italics="true" pageId="97" pageNumber="96">V</emphasis>
series trichobothria, of which the 3 most proximal are located on external surface near
<emphasis id="B951EAD4FF985010FE35F91CFDD0294F" box="[490,519,1777,1801]" italics="true" pageId="97" pageNumber="96">Eb</emphasis>
<subScript id="17A13483FF985010FDD8F8ECFDD82949" attach="left" box="[519,527,1793,1807]" fontSize="6" pageId="97" pageNumber="96">1</subScript>
. Chelal trichobothrium
<emphasis id="B951EAD4FF985010FFAEF8FCFF412969" box="[113,150,1809,1839]" italics="true" pageId="97" pageNumber="96">
Eb
<subScript id="17A13483FF985010FF51F8CCFF412969" attach="left" box="[142,150,1825,1839]" fontSize="6" pageId="97" pageNumber="96">3</subScript>
</emphasis>
located in distal half of manus between trichobothria
<emphasis id="B951EAD4FF985010FD36F8FFFCD4296F" box="[745,771,1810,1833]" italics="true" pageId="97" pageNumber="96">Dt</emphasis>
and
<emphasis id="B951EAD4FF985010FF7EF8DFFF15290F" box="[161,194,1842,1865]" italics="true" pageId="97" pageNumber="96">Est</emphasis>
. Fingers of pedipalps undulate in male. Chela length to width ratio 4. Pedipalp movable finger with 68 IAD, which have the same size as MD (&gt;
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number) and create a second row; there are also 5 ID and 15 OD present. Tarsomere II of leg III with four stout median ventral spinules and two pairs of flanking setae. Metasoma I with 10 carinae and metasoma II–IV with 8 carinae. Telson elongate and granulate, length to depth ratio
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male; annular ring developed.
</paragraph>
</subSubSection>
<subSubSection id="C33F654DFF985010FCEFFEA6FBB02924" pageId="97" pageNumber="96" type="description">
<paragraph id="8B9A36C6FF985010FCEFFEA6FAA72F85" blockId="97.[816,1475,331,1891]" pageId="97" pageNumber="96">
DESCRIPTION (
<specimenCount id="9D23FD4FFF985010FC0CFEA6FC302F25" box="[979,999,331,355]" pageId="97" pageNumber="96" type="male">♂</specimenCount>
<typeStatus id="549E8864FF985010FC29FEA6FB822F25" box="[1014,1109,331,355]" pageId="97" pageNumber="96" type="holotype">holotype</typeStatus>
). Total length
<quantity id="4CDD9B23FF985010FAD5FEA6FAB62F24" box="[1290,1377,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="97" pageNumber="96" unit="mm" value="70.0">70 mm</quantity>
. Female unknown. Habitus as shown in
<figureCitation id="131E2A43FF985010FB4FFE86FAE72FC4" box="[1168,1328,362,387]" captionStart="Figures 634–638" captionStartId="98.[115,192,1984,2006]" captionTargetBox="[121,1464,255,1950]" captionTargetId="figure-13@98.[113,1474,234,1958]" captionTargetPageId="98" captionText="Figures 634–638: Scorpiops scheibeae sp. n., male holotype. Figures 634–635. Dorsal (634) and ventral (635) views. Figures 636–638. Metasoma and telson lateral (636), ventral (637), and dorsal (638) views. Scale bars: 10 mm (634–635, 636–638)." figureDoi="http://doi.org/10.5281/zenodo.5741991" httpUri="https://zenodo.org/record/5741991/files/figure.png" pageId="97" pageNumber="96">Figs. 634–635</figureCitation>
. For position and distribution of trichobothria on pedipalps, see
<figureCitation id="131E2A43FF985010FA8FFE66FC8C2F85" captionStart="Figures 639–642" captionStartId="99.[118,195,1977,1999]" captionTargetBox="[121,1470,280,1870]" captionTargetId="figure-8@99.[113,1474,263,1903]" captionTargetPageId="99" captionText="Figures 639–642: Scorpiops scheibeae sp. n., male holotype, pedipalp segments. Chela and patell dorsal (639), external (640) and ventral (641) views. Movable finger dentition (642). Trichobothrial pattern is indicated by white circles (639–641)." figureDoi="http://doi.org/10.5281/zenodo.5741993" httpUri="https://zenodo.org/record/5741993/files/figure.png" pageId="97" pageNumber="96">Figs. 639– 641</figureCitation>
. Fingers of pedipalps undulate in male (
<figureCitation id="131E2A43FF985010FAD8FE46FAB32F85" box="[1287,1380,427,451]" captionStart="Figures 639–642" captionStartId="99.[118,195,1977,1999]" captionTargetBox="[121,1470,280,1870]" captionTargetId="figure-8@99.[113,1474,263,1903]" captionTargetPageId="99" captionText="Figures 639–642: Scorpiops scheibeae sp. n., male holotype, pedipalp segments. Chela and patell dorsal (639), external (640) and ventral (641) views. Movable finger dentition (642). Trichobothrial pattern is indicated by white circles (639–641)." figureDoi="http://doi.org/10.5281/zenodo.5741993" httpUri="https://zenodo.org/record/5741993/files/figure.png" pageId="97" pageNumber="96">Fig. 640</figureCitation>
).
</paragraph>
<paragraph id="8B9A36C6FF985010FCEFFE27FBEE2D25" blockId="97.[816,1475,331,1891]" pageId="97" pageNumber="96">
<emphasis id="B951EAD4FF985010FCEFFE27FC7C2FA4" bold="true" box="[816,939,458,482]" pageId="97" pageNumber="96">Coloration</emphasis>
(
<figureCitation id="131E2A43FF985010FC1AFE26FBBC2FA4" box="[965,1131,458,483]" captionStart="Figures 634–638" captionStartId="98.[115,192,1984,2006]" captionTargetBox="[121,1464,255,1950]" captionTargetId="figure-13@98.[113,1474,234,1958]" captionTargetPageId="98" captionText="Figures 634–638: Scorpiops scheibeae sp. n., male holotype. Figures 634–635. Dorsal (634) and ventral (635) views. Figures 636–638. Metasoma and telson lateral (636), ventral (637), and dorsal (638) views. Scale bars: 10 mm (634–635, 636–638)." figureDoi="http://doi.org/10.5281/zenodo.5741991" httpUri="https://zenodo.org/record/5741991/files/figure.png" pageId="97" pageNumber="96">Figs. 634–635</figureCitation>
,
<figureCitation id="131E2A43FF985010FB5EFE27FB7A2FA4" box="[1153,1197,458,482]" captionStart="Figures 647–652" captionStartId="101.[116,193,1958,1980]" captionTargetBox="[113,1474,255,1921]" captionTargetId="figure-9@101.[113,1474,255,1923]" captionTargetPageId="101" captionText="Figures 647–652: Scorpiops scheibeae sp. n., male holotype. Figures 647–648. Carapace and tergites I–III (647), coxosternal area and sternites III–IV (648), left chelicera in dorsal (649) and ventral (650) views. Figure 651. Telson lateral view. Figure 652. Male holotype in vivo habitus." figureDoi="http://doi.org/10.5281/zenodo.5741997" httpUri="https://zenodo.org/record/5741997/files/figure.png" pageId="97" pageNumber="96">652</figureCitation>
). Base color uniformly reddish black to black. Telson, legs, and sternites are reddish brown. Chelicerae reddish brown and reticulate, fingers black.
<emphasis id="B951EAD4FF985010FCEFFDC7FB982C05" bold="true" box="[816,1103,554,579]" pageId="97" pageNumber="96">Carapace and mesosoma</emphasis>
(
<figureCitation id="131E2A43FF985010FBBFFDC6FB292C04" box="[1120,1278,554,579]" captionStart="Figures 634–638" captionStartId="98.[115,192,1984,2006]" captionTargetBox="[121,1464,255,1950]" captionTargetId="figure-13@98.[113,1474,234,1958]" captionTargetPageId="98" captionText="Figures 634–638: Scorpiops scheibeae sp. n., male holotype. Figures 634–635. Dorsal (634) and ventral (635) views. Figures 636–638. Metasoma and telson lateral (636), ventral (637), and dorsal (638) views. Scale bars: 10 mm (634–635, 636–638)." figureDoi="http://doi.org/10.5281/zenodo.5741991" httpUri="https://zenodo.org/record/5741991/files/figure.png" pageId="97" pageNumber="96">Figs. 634–635</figureCitation>
,
<figureCitation id="131E2A43FF985010FAD4FDC6FAB92C05" box="[1291,1390,555,579]" captionStart="Figures 647–652" captionStartId="101.[116,193,1958,1980]" captionTargetBox="[113,1474,255,1921]" captionTargetId="figure-9@101.[113,1474,255,1923]" captionTargetPageId="101" captionText="Figures 647–652: Scorpiops scheibeae sp. n., male holotype. Figures 647–648. Carapace and tergites I–III (647), coxosternal area and sternites III–IV (648), left chelicera in dorsal (649) and ventral (650) views. Figure 651. Telson lateral view. Figure 652. Male holotype in vivo habitus." figureDoi="http://doi.org/10.5281/zenodo.5741997" httpUri="https://zenodo.org/record/5741997/files/figure.png" pageId="97" pageNumber="96">647–648</figureCitation>
). Entire carapace covered with large granules; carinae absent. Anterior margin of carapace with a deep median notch. Carapace with 3 pairs of lateral eyes of which two are normal and one is reduced. Mesosoma granulated, with one median carina. Tergite VII is pentacarinate. Sternites smooth to very finely granulated with two parallel furrows except sternite VII which has four granulate carinae. Pectine teeth number
<quantity id="4CDD9B23FF985010FA93FCE6FA522D65" box="[1356,1413,779,803]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="97" pageNumber="96" unit="in" value="12.0">12 in</quantity>
male
<typeStatus id="549E8864FF985010FCEFFCC6FC462D05" box="[816,913,811,835]" pageId="97" pageNumber="96" type="holotype">holotype</typeStatus>
. Pectines with 3 marginal and 4 middle lamellae, fulcra reduced or absent.
</paragraph>
<paragraph id="8B9A36C6FF985010FCEFFC86FB192AC5" blockId="97.[816,1475,331,1891]" pageId="97" pageNumber="96">
<emphasis id="B951EAD4FF985010FCEFFC86FBE02DC5" bold="true" box="[816,1079,875,899]" pageId="97" pageNumber="96">Metasoma and telson</emphasis>
(
<figureCitation id="131E2A43FF985010FB90FC86FB222DC5" box="[1103,1269,875,899]" captionStart="Figures 634–638" captionStartId="98.[115,192,1984,2006]" captionTargetBox="[121,1464,255,1950]" captionTargetId="figure-13@98.[113,1474,234,1958]" captionTargetPageId="98" captionText="Figures 634–638: Scorpiops scheibeae sp. n., male holotype. Figures 634–635. Dorsal (634) and ventral (635) views. Figures 636–638. Metasoma and telson lateral (636), ventral (637), and dorsal (638) views. Scale bars: 10 mm (634–635, 636–638)." figureDoi="http://doi.org/10.5281/zenodo.5741991" httpUri="https://zenodo.org/record/5741991/files/figure.png" pageId="97" pageNumber="96">Figs. 636–638</figureCitation>
,
<figureCitation id="131E2A43FF985010FAD4FC87FAE02DC4" box="[1291,1335,874,898]" captionStart="Figures 647–652" captionStartId="101.[116,193,1958,1980]" captionTargetBox="[113,1474,255,1921]" captionTargetId="figure-9@101.[113,1474,255,1923]" captionTargetPageId="101" captionText="Figures 647–652: Scorpiops scheibeae sp. n., male holotype. Figures 647–648. Carapace and tergites I–III (647), coxosternal area and sternites III–IV (648), left chelicera in dorsal (649) and ventral (650) views. Figure 651. Telson lateral view. Figure 652. Male holotype in vivo habitus." figureDoi="http://doi.org/10.5281/zenodo.5741997" httpUri="https://zenodo.org/record/5741997/files/figure.png" pageId="97" pageNumber="96">651</figureCitation>
). Metasoma sparsely hirsute and granulated, with sparse, relatively large granules. Metasomal segment I with 10 carinae, II–IV with 8 carinae, and V with 5 carinae. Median lateral carinae of metasoma II and V indicated by isolated granules that may coalesce into carinae. Dorsolateral carinae of segments II– IV terminate posteriorly in a pronounced tooth which is on metasoma IV wide with three peaks. Telson elongate and granulate with annular ring developed.
</paragraph>
<paragraph id="8B9A36C6FF985010FCEFFB67FAD32804" blockId="97.[816,1475,331,1891]" pageId="97" pageNumber="96">
<emphasis id="B951EAD4FF985010FCEFFB67FC482AE4" bold="true" box="[816,927,1162,1186]" pageId="97" pageNumber="96">Pedipalps</emphasis>
(
<figureCitation id="131E2A43FF985010FC6EFB66FB842AE5" box="[945,1107,1163,1187]" captionStart="Figures 639–642" captionStartId="99.[118,195,1977,1999]" captionTargetBox="[121,1470,280,1870]" captionTargetId="figure-8@99.[113,1474,263,1903]" captionTargetPageId="99" captionText="Figures 639–642: Scorpiops scheibeae sp. n., male holotype, pedipalp segments. Chela and patell dorsal (639), external (640) and ventral (641) views. Movable finger dentition (642). Trichobothrial pattern is indicated by white circles (639–641)." figureDoi="http://doi.org/10.5281/zenodo.5741993" httpUri="https://zenodo.org/record/5741993/files/figure.png" pageId="97" pageNumber="96">Figs. 639–642</figureCitation>
). Pedipalps very sparsely hirsute. Patella with 58 external trichobothria (which cannot be assigned to standard
<emphasis id="B951EAD4FF985010FBD0FB26FBFF2AA5" box="[1039,1064,1227,1251]" italics="true" pageId="97" pageNumber="96">eb</emphasis>
,
<emphasis id="B951EAD4FF985010FBEAFB26FB8E2AA5" box="[1077,1113,1227,1251]" italics="true" pageId="97" pageNumber="96">esb</emphasis>
,
<emphasis id="B951EAD4FF985010FBB9FB26FB522AA4" box="[1126,1157,1227,1250]" italics="true" pageId="97" pageNumber="96">em</emphasis>
,
<emphasis id="B951EAD4FF985010FB4DFB26FB672AA4" box="[1170,1200,1227,1250]" italics="true" pageId="97" pageNumber="96">est</emphasis>
, and
<emphasis id="B951EAD4FF985010FB35FB26FB2A2AA4" box="[1258,1277,1227,1250]" italics="true" pageId="97" pageNumber="96">et</emphasis>
territories) and 24 ventral trichobothria. Chela with 14
<emphasis id="B951EAD4FF985010FB67FB06FB1F2B44" box="[1208,1224,1259,1282]" italics="true" pageId="97" pageNumber="96">V</emphasis>
series trichobothria, of which the 3 most proximal are located on external surface near
<emphasis id="B951EAD4FF985010FCEFFAC6FC992B05" box="[816,846,1323,1347]" italics="true" pageId="97" pageNumber="96">Eb</emphasis>
<subScript id="17A13483FF985010FC91FAD6FC812B0F" attach="left" box="[846,854,1339,1353]" fontSize="6" pageId="97" pageNumber="96">1</subScript>
. Femur and patella granulated. Femur with 5 granulose carinae, and patella with 5 carinae, with dorsal and ventral patellar spurs pronounced. Manus dorsally with fine, rounded granules, which in the central area form an irregular dorsal secondary carina. External surface of chela densely covered by minute granules, with an irregular external secondary carina. Movable fingers bear 68 IAD, which have the same size as MD (more than
<quantity id="4CDD9B23FF985010FBE8F9E6FBAA2865" box="[1079,1149,1547,1571]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="97" pageNumber="96" unit="in" value="100.0">100 in</quantity>
number) and create a second row; there are also 5 ID and 15 OD present.
</paragraph>
<paragraph id="8B9A36C6FF985010FCEFF9A6FB2C2905" blockId="97.[816,1475,331,1891]" pageId="97" pageNumber="96">
<emphasis id="B951EAD4FF985010FCEFF9A6FCB22825" bold="true" box="[816,869,1611,1635]" pageId="97" pageNumber="96">Legs</emphasis>
(
<figureCitation id="131E2A43FF985010FCA3F9A6FBF62825" box="[892,1057,1611,1635]" captionStart="Figures 643–646" captionStartId="100.[117,194,2001,2023]" captionTargetBox="[120,1465,260,1965]" captionTargetId="figure-14@100.[113,1474,234,1979]" captionTargetPageId="100" captionText="Figures 643–646. Scorpiops scheibeae sp. n., male holotype, left legs I–IV, prolateral/ ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.5741995" httpUri="https://zenodo.org/record/5741995/files/figure.png" pageId="97" pageNumber="96">Figs. 643–646</figureCitation>
). Tibia and tarsomeres of legs with several setae not arranged into bristle combs on dorsal surfaces but with rows of spinules on dorsolateral surfaces and on legs I
<emphasis id="B951EAD4FF985010FC53F946FC4E2884" box="[908,921,1707,1730]" italics="true" pageId="97" pageNumber="96">–</emphasis>
II also on ventrolateral surface. Tarsomere II of legs I–II, IV with 5 stout median ventral spinules and two pairs of flanking setae, leg III with 4 stout median ventral spinules and two pairs of flanking setae. Femur with 3–4 and patella 4–5 carinae; both femur and patella granulated.
</paragraph>
<paragraph id="8B9A36C6FF985010FCEFF8A6FBB02924" blockId="97.[816,1475,331,1891]" box="[816,1127,1867,1891]" pageId="97" pageNumber="96">
<emphasis id="B951EAD4FF985010FCEFF8A6FC002925" bold="true" box="[816,983,1867,1891]" pageId="97" pageNumber="96">Measurements</emphasis>
. See
<tableCitation id="C6A7037DFF985010FBCEF8A6FBB42924" box="[1041,1123,1867,1891]" captionStart="Table 7" captionStartId="102.[116,172,991,1012]" captionText="Table 7. Comparative measurements of adults of Scorpiops scheibeae sp. n., S. sherwoodae sp. n., and S. solegladi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="97" pageNumber="96">Table 7</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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