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<document id="C0FFED3E911F4335C2982B2194C48664" ID-CLB-Dataset="57388" ID-DOI="10.31610/zsr/2023.32.1.16" ID-GBIF-Dataset="939ab263-2d82-4840-881a-3b6500e0a68f" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124261" ID-ZooBank="DB58E140-9441-4C52-8B18-E3505F3D9E93" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688481640215" checkinUser="felipe" docAuthor="Evstigneev, D. A." docDate="2023" docId="03FA4A08FFBCFF9F6EEE472FFD15FDD9" docLanguage="en" docName="ZoosystRoss.32.1.16-22.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 32 (1)" docSource="http://dx.doi.org/10.31610/zsr/2023.32.1.16" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Urophora sevanensis Evstigneev 2023, sp. nov." docType="treatment" docUuid="394921CD-A77A-419F-A0EF-22FBD87CE070" docUuidSource="ZooBank" docVersion="8" lastPageNumber="21" masterDocId="FFC33270FFBDFF9A6E714276FF9CFFEE" masterDocTitle="A second species of the genus Urophora (Diptera: Tephritidae) attacking capitula of the genus Psephellus (Asteraceae)" masterLastPageNumber="22" masterPageNumber="16" pageNumber="17" updateTime="1700050579405" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="14A00FCD46ED3528FD9136404059431C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F2E4F7EFCAEA10B833F6DA974520D3AE">A second species of the genus Urophora (Diptera: Tephritidae) attacking capitula of the genus Psephellus (Asteraceae)</mods:title>
</mods:titleInfo>
<mods:name id="7768C1231BF8F0FD2D54C001D197B9DA" type="personal">
<mods:role id="55810958DBB469B3697DC92FFD208B61">
<mods:roleTerm id="BE49A51FAF9B4CFD79851CC0D3AB1F47">Author</mods:roleTerm>
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<mods:namePart id="579F1CF4ED5BA77BCF30F42865DE896F">Evstigneev, D. A.</mods:namePart>
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<mods:typeOfResource id="5089915CDC1CBEFE516E013A8490606A">text</mods:typeOfResource>
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<mods:title id="A88829D8281436CE0D028F910CF09F68">Zoosystematica Rossica</mods:title>
</mods:titleInfo>
<mods:part id="AAD09DC61FCFCE57F956746AA4384098">
<mods:date id="7B657FAA5ADC967445272BD61D4CDE0A">2023</mods:date>
<mods:detail id="165D9B137DF21CC539BE8952CE573327" type="series">
<mods:title id="83EAA38EE13AB5B467696A2ED573BDD8">Zoosyst. Rossica</mods:title>
</mods:detail>
<mods:detail id="163A9DC4B2A9222A2EB8DDB7E5C08548" type="pubDate">
<mods:number id="F704CB8B00BA810AD26AC86314E51870">2023-04-14</mods:number>
</mods:detail>
<mods:detail id="6D5C66449EC1F84E40AC68F21D84C4F3" type="volume">
<mods:number id="127FC42CEC7C54F934914BB4E73ACE1E">32</mods:number>
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<mods:number id="F4CE4ED0AEC8ABAF2795051BE871EA38">1</mods:number>
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<mods:url id="9B766A34E24E3E8BC04D6040D011803C">http://dx.doi.org/10.31610/zsr/2023.32.1.16</mods:url>
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<mods:classification id="DF3C0C50BFB49C3670797BF26C21AE83">journal article</mods:classification>
<mods:identifier id="911D695C3BA1DAD07BF86CD53C881F08" type="CLB-Dataset">57388</mods:identifier>
<mods:identifier id="00E56C05DC4F4397D4F8F8B9E84A5450" type="DOI">10.31610/zsr/2023.32.1.16</mods:identifier>
<mods:identifier id="AF596571771C166CB92C574581F3304E" type="GBIF-Dataset">939ab263-2d82-4840-881a-3b6500e0a68f</mods:identifier>
<mods:identifier id="D0FA7B666C22A16A15A1881FFBFD235A" type="ISSN">2410-0226</mods:identifier>
<mods:identifier id="9056AEE6583BB42E43034BEF3E8FEC51" type="Zenodo-Dep">10124261</mods:identifier>
<mods:identifier id="5636B16EA276A37842B3FDFC42DB7F18" type="ZooBank">DB58E140-9441-4C52-8B18-E3505F3D9E93</mods:identifier>
</mods:mods>
<treatment id="03FA4A08FFBCFF9F6EEE472FFD15FDD9" ID-DOI="http://doi.org/10.5281/zenodo.8117839" ID-GBIF-Taxon="210812433" ID-Zenodo-Dep="8117839" LSID="urn:lsid:zoobank.org:act:394921CD-A77A-419F-A0EF-22FBD87CE070" httpUri="http://treatment.plazi.org/id/03FA4A08FFBCFF9F6EEE472FFD15FDD9" lastPageId="5" lastPageNumber="21" pageId="1" pageNumber="17">
<subSubSection id="C349A895FFBCFF9B6EEE472FFDB8FA9D" box="[159,548,1369,1396]" pageId="1" pageNumber="17" type="nomenclature">
<paragraph id="8BECFB1EFFBCFF9B6EEE472FFDB8FA9D" blockId="1.[159,548,1369,1396]" box="[159,548,1369,1396]" pageId="1" pageNumber="17">
<heading id="D0A44C72FFBCFF9B6EEE472FFDB8FA9D" box="[159,548,1369,1396]" fontSize="11" level="2" pageId="1" pageNumber="17" reason="2">
<emphasis id="B927270CFFBCFF9B6EEE472FFDB8FA9D" bold="true" box="[159,548,1369,1396]" pageId="1" pageNumber="17">
<taxonomicName id="4C53809DFFBCFF9B6EEE472FFE25FA9D" ID-CoL="BDPJK" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[159,441,1369,1396]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFBCFF9B6EEE472FFE25FA9D" bold="true" box="[159,441,1369,1396]" italics="true" pageId="1" pageNumber="17">Urophora sevanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2149A77FFBCFF9B6FB1472FFDB8FA9D" box="[448,548,1369,1395]" pageId="1" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFBCFF9B6EEE470AFEB9FA79" box="[159,293,1404,1431]" pageId="1" pageNumber="17" type="description">
<paragraph id="8BECFB1EFFBCFF9B6EEE470AFEB9FA79" blockId="1.[159,293,1404,1431]" box="[159,293,1404,1431]" pageId="1" pageNumber="17">
(
<figureCitation id="1368E79BFFBCFF9B6ED9470AFE86FA79" box="[168,282,1404,1431]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Figs 1–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFBCFF9B6EB447C3FEB4F961" pageId="1" pageNumber="17" type="materials_examined">
<paragraph id="8BECFB1EFFBCFF9B6EB447C3FEF3F9A0" blockId="1.[156,779,1461,1927]" pageId="1" pageNumber="17">
<materialsCitation id="3B3BF143FFBCFF9B6EB447C3FEF3F9A0" ID-GBIF-Occurrence="4145107301" collectingDate="2021-07-25" collectingDateMax="2022-04-11" collectingDateMin="2021-07-25" collectorName="D. Evstigneev" country="Armenia" location="Sevan Lake" municipality="Female" pageId="1" pageNumber="17" specimenCount="1" stateProvince="Geghark'unik'" typeStatus="holotype">
<emphasis id="B927270CFFBCFF9B6EB447C3FEB7FA20" box="[197,299,1461,1486]" italics="true" pageId="1" pageNumber="17">
<typeStatus id="54E845BCFFBCFF9B6EB447C3FEB7FA20" box="[197,299,1461,1486]" pageId="1" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="6B886164FFBCFF9B6F3047C3FE0AFA20" box="[321,406,1461,1486]" pageId="1" pageNumber="17">Female</collectingMunicipality>
,
<emphasis id="B927270CFFBCFF9B6FD847C3FD8DFA23" bold="true" box="[425,529,1461,1485]" pageId="1" pageNumber="17">
<collectingCountry id="F344BB8EFFBCFF9B6FD847C3FD8DFA23" box="[425,529,1461,1485]" name="Armenia" pageId="1" pageNumber="17">Armenia</collectingCountry>
</emphasis>
,
<emphasis id="B927270CFFBCFF9B6C5647C3FC99FA23" box="[551,773,1461,1486]" italics="true" pageId="1" pageNumber="17">
<collectingRegion id="499735FCFFBCFF9B6C5647C3FC99FA23" box="[551,773,1461,1486]" country="Armenia" name="Geghark'unik'" pageId="1" pageNumber="17">Gegharkunik Prov.</collectingRegion>
</emphasis>
, western coast of
<location id="8E8CADC5FFBCFF9B6F2647A0FE41FA00" LSID="urn:lsid:plazi:treatment:03FA4A08FFBCFF9F6EEE472FFD15FDD9:8E8CADC5FFBCFF9B6F2647A0FE41FA00" box="[343,477,1493,1518]" country="Armenia" municipality="Female" name="Sevan Lake" pageId="1" pageNumber="17" stateProvince="Geghark'unik'">Sevan Lake</location>
,
<locationDeviation id="9A5EE23CFFBCFF9B6F9447A3FEB6F9E0" firstLocation="Noratus Vill." pageId="1" pageNumber="17" secondLocation="Eranos Vill.">between Noratus Vill. and Eranos Vill.</locationDeviation>
, among rocks, reared on
<date id="FFEDDDDEFFBCFF9B6C3D4780FD56F9E0" box="[588,714,1526,1550]" pageId="1" pageNumber="17" value="2022-04-11">
<collectingDate id="EFA92436FFBCFF9B6C3D4780FD56F9E0" box="[588,714,1526,1550]" pageId="1" pageNumber="17" value="2022-04-11">11.IV.2022</collectingDate>
</date>
from capitulum of
<taxonomicName id="4C53809DFFBCFF9B6F4A4463FE46F9C3" box="[315,474,1557,1582]" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="17" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B927270CFFBCFF9B6F4A4463FE2DF9C0" box="[315,433,1557,1582]" italics="true" pageId="1" pageNumber="17">Psephellus</emphasis>
sp.
</taxonomicName>
collected on
<date id="FFEDDDDEFFBCFF9B6C0B4463FC9AF9C0" box="[634,774,1557,1582]" pageId="1" pageNumber="17" value="2021-07-25">
<collectingDate id="EFA92436FFBCFF9B6C0B4463FC9AF9C0" box="[634,774,1557,1582]" pageId="1" pageNumber="17" value="2021-07-25">25.VII.2021</collectingDate>
</date>
,
<collectorName id="26A69EC8FFBCFF9B6EEE4440FEDDF9A0" box="[159,321,1589,1614]" pageId="1" pageNumber="17">D. Evstigneev</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BECFB1EFFBCFF9B6EB44421FEB4F961" blockId="1.[156,779,1461,1927]" pageId="1" pageNumber="17">
<materialsCitation id="3B3BF143FFBCFF9B6EB44421FEB9F961" ID-GBIF-Occurrence="4145107302" pageId="1" pageNumber="17" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B927270CFFBCFF9B6EB44421FEABF980" box="[197,311,1623,1646]" italics="true" pageId="1" pageNumber="17">
<typeStatus id="54E845BCFFBCFF9B6EB44421FEABF980" box="[197,311,1623,1646]" pageId="1" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="9D553097FFBCFF9B6F354420FE06F980" box="[324,410,1621,1646]" pageId="1" pageNumber="17" type="male">3 males</specimenCount>
and
<specimenCount id="9D553097FFBCFF9B6FA34420FDA8F980" box="[466,564,1621,1646]" pageId="1" pageNumber="17" type="female">1 female</specimenCount>
, with same data as in holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFBCFF9B6EB444EFFC6FF962" pageId="1" pageNumber="17" type="description">
<paragraph id="8BECFB1EFFBCFF9B6EB444EFFAB0FEF4" blockId="1.[156,779,1461,1927]" lastBlockId="1.[822,1447,185,1933]" pageId="1" pageNumber="17">
<emphasis id="B927270CFFBCFF9B6EB444EFFEC9F95C" box="[197,341,1689,1714]" italics="true" pageId="1" pageNumber="17">Description</emphasis>
. Medium-sized fly: female body length (including oviscape) 7.0–
<quantity id="4CAB56FBFFBCFF9B6C2044CDFD25F938" box="[593,697,1723,1750]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="17" unit="mm" value="7.8">7.8 mm</quantity>
, male body length
<quantity id="4CAB56FBFFBCFF9B6F4644A8FE53F917" box="[311,463,1758,1786]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.3" metricValueMin="4.6" pageId="1" pageNumber="17" unit="mm" value="4.949999999999999" valueMax="5.3" valueMin="4.6">4.6–5.3 mm</quantity>
, wing length 3.7–4.0 mm. General body colour black. All setae black. Ratios of head length to its height and width 1: 1.4: 1.5. Frons predominantly orange, with some areas pale brown. Palpus whitish, darkening to orange apically. Proboscis yellowish or whitish, with yellowish areas. First flagellomere orange. Pedicel yellow or orange. Scape yellow. Lunule whitish.
</paragraph>
<paragraph id="8BECFB1EFFBCFF9B6D2F4352FBE2FAC8" blockId="1.[822,1447,185,1933]" pageId="1" pageNumber="17">
Thorax predominantly black; its length
<quantity id="4CAB56FBFFBCFF9B6B1A4355FC05FE8C" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="1" pageNumber="17" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6– 1.8 mm</quantity>
. Postpronotal lobe largely yellow or whitish. Upper part of anepisternum yellowish or whitish. Postpronotal seta based in yellowish area. Upper anepisternal seta based in yellowish area (
<figureCitation id="1368E79BFFBCFF9B6D3043A0FC10FE1F" box="[833,908,470,497]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 1</figureCitation>
). Anterior notopleural seta based in black area (on margin of black and yellow areas). Posterior notopleural seta based in black area. Scutellum yellow centrally, black laterally, with two pairs of setae (
<figureCitation id="1368E79BFFBCFF9B6DFA4013FC44FD6E" box="[907,984,613,640]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 7</figureCitation>
); basal scutellar setae based in black lateral area; apical scutellar setae based in yellow central area. Wing with distinct black subbasal, discal, preapical and apical crossbands (crossbands listed from wing base to wing apex). Subbasal and discal crossbands separate (
<figureCitation id="1368E79BFFBCFF9B6AE0416EFA9AFCDC" box="[1169,1286,792,818]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Figs 1–4</figureCitation>
). Discal and preapical crossbands separate. Preapical and apical crossbands joined anteriorly between veins
<emphasis id="B927270CFFBCFF9B6B2A4116FAF3FC94" box="[1371,1391,864,890]" italics="true" pageId="1" pageNumber="17">C</emphasis>
and
<emphasis id="B927270CFFBCFF9B6D4B41F2FCF6FC4A" box="[826,874,900,932]" italics="true" pageId="1" pageNumber="17">
R
<subScript id="17D7F95BFFBCFF9B6D3C41E0FCF6FC4A" attach="left" box="[845,874,918,932]" fontSize="6" pageId="1" pageNumber="17">4+5</subScript>
</emphasis>
[black area extends from one-third (
<figureCitation id="1368E79BFFBCFF9B6B3B41F5FA05FC70" box="[1354,1433,899,926]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 3</figureCitation>
) to three-fourths (
<figureCitation id="1368E79BFFBCFF9B6A7E41D1FBC7FC2F" box="[1039,1115,935,961]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 4</figureCitation>
) of width of cell
<emphasis id="B927270CFFBCFF9B6B5B41DEFACCFC26" box="[1322,1360,936,968]" italics="true" pageId="1" pageNumber="17">
r
<subScript id="17D7F95BFFBCFF9B6B4541CCFACCFC26" attach="left" box="[1332,1360,954,968]" fontSize="6" pageId="1" pageNumber="17">2+3</subScript>
</emphasis>
]. Subbasal crossband narrowed posteriorly, forming a process (
<figureCitation id="1368E79BFFBCFF9B6DB44199FBA2FBE7" box="[965,1086,1007,1033]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Figs 1–4</figureCitation>
). Posterior process is more prominent in males. Scutum with dense microtrichial pattern, similar in colour to pollen, with or without stripes and marks (
<figureCitation id="1368E79BFFBCFF9B6AEB462CFB77FB9A" box="[1178,1259,1114,1140]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 7</figureCitation>
). Male femora usually entirely orange (
<figureCitation id="1368E79BFFBCFF9B6A1F460BFB26FB76" box="[1134,1210,1149,1176]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 1</figureCitation>
). Female fore and hind femora orange, with black longitudinal stripe; middle femora predominantly orange, with small mark on basal half (
<figureCitation id="1368E79BFFBCFF9B6A58469FFBE9FAED" box="[1065,1141,1257,1283]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 2</figureCitation>
). Two apical segments of tarsi often brown or black.
</paragraph>
<paragraph id="8BECFB1EFFBCFF9B6D2F4746FC6FF962" blockId="1.[822,1447,185,1933]" pageId="1" pageNumber="17">
Abdomen black. Male abdomen length
<quantity id="4CAB56FBFFBCFF9B6B254746FA3CFAA5" box="[1364,1440,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="17" unit="mm" value="2.0">2 mm</quantity>
. Female abdomen (including oviscape) length
<quantity id="4CAB56FBFFBCFF9B6B184722FC0BFA7C" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="1" pageNumber="17" unit="mm" value="5.05" valueMax="5.3" valueMin="4.8">4.8– 5.3 mm</quantity>
. Oviscape (= syntergosternite 7, = seventh tergosternite, = tergosternum 7) shiny black, 1.6–1.7 times as long as all tergites combined. Oviscape length 3.0–
<quantity id="4CAB56FBFFBCFF9B6A3C4795FB2EFA13" box="[1101,1202,1507,1533]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="17" unit="mm" value="3.3">3.3 mm</quantity>
. Aculeus with two pairs of preapical steps including strong primary (proximal) and weaker secondary (distal) steps (
<figureCitation id="1368E79BFFBCFF9B6D304438FC13F987" box="[833,911,1613,1641]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="1" pageNumber="17">Fig. 6</figureCitation>
). Aculeus with weak apical notch. Aculeus length
<quantity id="4CAB56FBFFBCFF9B6DE14404FC73F962" box="[912,1007,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="17" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFBCFF996D2F44E0FB4DF868" lastPageId="3" lastPageNumber="19" pageId="1" pageNumber="17" type="diagnosis">
<paragraph id="8BECFB1EFFBCFF996D2F44E0FB4DF868" blockId="1.[822,1447,185,1933]" lastBlockId="3.[823,1445,1535,1926]" lastPageId="3" lastPageNumber="19" pageId="1" pageNumber="17">
<emphasis id="B927270CFFBCFF9B6D2F44E0FC68F95E" box="[862,1012,1686,1712]" italics="true" pageId="1" pageNumber="17">Comparison</emphasis>
. The only other
<taxonomicName id="4C53809DFFBCFF9B6ABC44E0FA39F95E" box="[1229,1445,1686,1712]" pageId="1" pageNumber="17">
<emphasis id="B927270CFFBCFF9B6ABC44E0FADFF95E" box="[1229,1347,1686,1712]" italics="true" pageId="1" pageNumber="17">Urophora</emphasis>
species
</taxonomicName>
known to attack the genus
<taxonomicName id="4C53809DFFBCFF9B6AD444CFFABAF93D" box="[1189,1318,1721,1747]" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B927270CFFBCFF9B6AD444CFFABAF93D" box="[1189,1318,1721,1747]" italics="true" pageId="1" pageNumber="17">Psephellus</emphasis>
</taxonomicName>
is
<taxonomicName id="4C53809DFFBCFF9B6B2544CCFB7EF916" authority="Korneyev et White, 1996" authorityName="Korneyev et White" authorityYear="1996" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="trinervii">
<emphasis id="B927270CFFBCFF9B6B2544CCFC1DF919" italics="true" pageId="1" pageNumber="17">U. trinervii</emphasis>
<bibRefCitation id="EFC286EFFFBCFF9B6DFF44A8FB7EF916" author="Korneyev V. A. &amp; White I. M." box="[910,1250,1756,1784]" pageId="1" pageNumber="17" pagination="499 - 513" refId="ref3424" refString="Korneyev V. A. &amp; White I. M. 1996. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. II. Review of species of the subgenus Urophora s. str. Communication 3. Entomologicheskoe Obozrenie, 75 (2): 463 - 477. (In Russian; English translation: Entomological Review, 1996, 76 (4): 499 - 513)." type="journal article" year="1996">Korneyev et White, 1996</bibRefCitation>
</taxonomicName>
, which has an entirely different wing pattern characterised by reduced, faint crossbands, with the subbasal crossband absent or rarely represented by only a grey spot located between veins
<emphasis id="B927270CFFBCFF9B6A81451BFA98F868" box="[1264,1284,1901,1926]" italics="true" pageId="1" pageNumber="17">R</emphasis>
<subScript id="17D7F95BFFBCFF9B6B754509FA91F863" attach="left" box="[1284,1293,1919,1933]" fontSize="6" pageId="1" pageNumber="17">1</subScript>
and
<emphasis id="B927270CFFBCFF9B6B3F451AFAFEF868" box="[1358,1378,1900,1926]" italics="true" pageId="1" pageNumber="17">A</emphasis>
<subScript id="17D7F95BFFBCFF9B6B134509FAF7F863" attach="left" box="[1378,1387,1919,1933]" fontSize="6" pageId="1" pageNumber="17">1</subScript>
, the discal crossband obscure, represented in females by a weak, diffuse spot bordering
<emphasis id="B927270CFFBEFF996C214452FDE1F9D3" box="[592,637,1572,1597]" italics="true" pageId="3" pageNumber="19">r-m</emphasis>
(=
<emphasis id="B927270CFFBEFF996CDA4452FD5CF9D3" box="[683,704,1572,1597]" italics="true" pageId="3" pageNumber="19">ta</emphasis>
), and usually more or less expressed between
<emphasis id="B927270CFFBEFF996CF7443FFD3FF987" box="[646,675,1609,1641]" italics="true" pageId="3" pageNumber="19">
R
<subScript id="17D7F95BFFBEFF996CEB442DFD3FF987" attach="left" box="[666,675,1627,1641]" fontSize="6" pageId="3" pageNumber="19">1</subScript>
</emphasis>
and
<emphasis id="B927270CFFBEFF996CAF443FFC97F98C" box="[734,779,1609,1634]" italics="true" pageId="3" pageNumber="19">r-m</emphasis>
in males, and the preapical and apical crossbands separated in females, and joined in 65% males. The basal scutellar seta in
<taxonomicName id="4C53809DFFBEFF996C7144C0FD11F921" authorityName="Korneyev et White" authorityYear="1996" box="[512,653,1718,1743]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="trinervii">
<emphasis id="B927270CFFBEFF996C7144C0FD11F921" box="[512,653,1718,1743]" italics="true" pageId="3" pageNumber="19">U. trinervii</emphasis>
</taxonomicName>
is on the margin of the yellow and black areas (
<bibRefCitation id="EFC286EFFFBEFF996CFD44ACFEFAF8F4" author="Korneyev V. A. &amp; White I. M." pageId="3" pageNumber="19" pagination="499 - 513" refId="ref3424" refString="Korneyev V. A. &amp; White I. M. 1996. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. II. Review of species of the subgenus Urophora s. str. Communication 3. Entomologicheskoe Obozrenie, 75 (2): 463 - 477. (In Russian; English translation: Entomological Review, 1996, 76 (4): 499 - 513)." type="journal article" year="1996">Korneyev &amp; White, 1996</bibRefCitation>
) versus within the black area in
<taxonomicName id="4C53809DFFBEFF996EEE4552FED9F8D3" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[159,325,1828,1853]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFBEFF996EEE4552FED9F8D3" box="[159,325,1828,1853]" italics="true" pageId="3" pageNumber="19">U. sevanensis</emphasis>
</taxonomicName>
<emphasis id="B927270CFFBEFF996F3E4555FE28F8D3" bold="true" box="[335,436,1827,1853]" pageId="3" pageNumber="19">
<taxonomicNameLabel id="A2149A77FFBEFF996F3E4555FE28F8D3" box="[335,436,1827,1853]" pageId="3" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<bibRefCitation id="EFC286EFFFBEFF996FCA4552FC96F8D0" author="Korneyev V. A. &amp; White I. M." box="[443,778,1826,1854]" pageId="3" pageNumber="19" pagination="499 - 513" refId="ref3424" refString="Korneyev V. A. &amp; White I. M. 1996. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. II. Review of species of the subgenus Urophora s. str. Communication 3. Entomologicheskoe Obozrenie, 75 (2): 463 - 477. (In Russian; English translation: Entomological Review, 1996, 76 (4): 499 - 513)." type="journal article" year="1996">Korneyev &amp; White (1996)</bibRefCitation>
reared
<taxonomicName id="4C53809DFFBEFF996E8A453EFE14F88F" authorityName="Korneyev et White" authorityYear="1996" box="[251,392,1864,1889]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="trinervii">
<emphasis id="B927270CFFBEFF996E8A453EFE14F88F" box="[251,392,1864,1889]" italics="true" pageId="3" pageNumber="19">U. trinervii</emphasis>
</taxonomicName>
from capitulum galls on
<taxonomicName id="4C53809DFFBEFF996CA4453EFDF0F868" authority="(Willd.) Wagenitz" authorityName="Wagenitz" baseAuthorityName="Willd." class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="19" phylum="Tracheophyta" rank="species" species="trinervius">
<emphasis id="B927270CFFBEFF996CA4453EFEE5F868" italics="true" pageId="3" pageNumber="19">Psephellus trinervius</emphasis>
(Willd.) Wagenitz
</taxonomicName>
[as
<taxonomicName id="4C53809DFFBEFF996CD3451AFB8CF9F7" box="[674,1040,1535,1926]" class="Magnoliopsida" family="Asteraceae" genus="Odontolophus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="19" phylum="Tracheophyta" rank="species" species="trinervius">
<emphasis id="B927270CFFBEFF996CD3451AFB8CF9F7" box="[674,1040,1535,1926]" italics="true" pageId="3" pageNumber="19">Odontolophus trinervius</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC286EFFFBEFF996A354476FA38F9F5" author="Korneyev V. A. &amp; White I. M." box="[1092,1444,1535,1563]" pageId="3" pageNumber="19" pagination="499 - 513" refId="ref3424" refString="Korneyev V. A. &amp; White I. M. 1996. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. II. Review of species of the subgenus Urophora s. str. Communication 3. Entomologicheskoe Obozrenie, 75 (2): 463 - 477. (In Russian; English translation: Entomological Review, 1996, 76 (4): 499 - 513)." type="journal article" year="1996">Korneyev &amp; White (1996)</bibRefCitation>
and as
<taxonomicName id="4C53809DFFBEFF996DEF4452FAE7F9D0" box="[926,1403,1572,1598]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="19" phylum="Tracheophyta" rank="species" species="trinervius" subGenus="Odontolophus">
<emphasis id="B927270CFFBEFF996DEF4452FBBDF9D0" box="[926,1057,1572,1598]" italics="true" pageId="3" pageNumber="19">Centaurea</emphasis>
(
<emphasis id="B927270CFFBEFF996A4B4452FB79F9D0" box="[1082,1253,1572,1598]" italics="true" pageId="3" pageNumber="19">Odontolophus</emphasis>
)
<emphasis id="B927270CFFBEFF996B704453FAE7F9D0" box="[1281,1403,1573,1598]" italics="true" pageId="3" pageNumber="19">trinervius</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC286EFFFBEFF996D49443FFB11F98D" author="Korneyev V. A. &amp; White I. M." box="[824,1165,1607,1635]" pageId="3" pageNumber="19" pagination="296 - 309" refId="ref3506" refString="Korneyev V. A. &amp; White I. M. 1999. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. III. Key to palaearctic species. Entomologicheskoe Obozrenie, 78 (2): 464 - 482. (In Russian; English translation: Entomological Review, 1999, 79 (3): 296 - 309)." type="journal article" year="1999">Korneyev &amp; White (1999)</bibRefCitation>
].
<taxonomicName id="4C53809DFFBEFF996AD2443EFA39F98C" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[1187,1445,1608,1634]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFBEFF996AD2443EFA39F98C" box="[1187,1445,1608,1634]" italics="true" pageId="3" pageNumber="19">Urophora sevanensis</emphasis>
</taxonomicName>
<emphasis id="B927270CFFBEFF996D49441BFC00F969" bold="true" box="[824,924,1645,1671]" pageId="3" pageNumber="19">
<taxonomicNameLabel id="A2149A77FFBEFF996D49441BFC00F969" box="[824,924,1645,1671]" pageId="3" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C53809DFFBEFF996A534418FAD1F969" authority="(Fallen, 1820)" baseAuthorityName="Fallen" baseAuthorityYear="1820" box="[1058,1357,1645,1671]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="aprica">
<emphasis id="B927270CFFBEFF996A534418FB0FF969" box="[1058,1171,1646,1671]" italics="true" pageId="3" pageNumber="19">U. aprica</emphasis>
(Fallén, 1820)
</taxonomicName>
, which is associated with
<taxonomicName id="4C53809DFFBEFF996A6944E4FB6BF945" box="[1048,1271,1682,1708]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="19" phylum="Tracheophyta" rank="species" species="cyanus">
<emphasis id="B927270CFFBEFF996A6944E4FB6BF945" box="[1048,1271,1682,1708]" italics="true" pageId="3" pageNumber="19">Centaurea cyanus</emphasis>
</taxonomicName>
L. A latter fly species located in couplet
<quantity id="4CAB56FBFFBEFF996A0A44C3FB27F93E" box="[1147,1211,1717,1745]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="3" pageNumber="19" unit="in" value="54.0">54 in</quantity>
a key for the identification of the eastern Palaearctic
<taxonomicName id="4C53809DFFBEFF996A8544ACFCFAF8F7" pageId="3" pageNumber="19">
<emphasis id="B927270CFFBEFF996A8544ACFAF6F91A" box="[1268,1386,1754,1780]" italics="true" pageId="3" pageNumber="19">Urophora</emphasis>
species
</taxonomicName>
provided by
<bibRefCitation id="EFC286EFFFBEFF996A764489FAC8F8F7" author="Korneyev V. A. &amp; White I. M." box="[1031,1364,1791,1817]" pageId="3" pageNumber="19" pagination="296 - 309" refId="ref3506" refString="Korneyev V. A. &amp; White I. M. 1999. Fruit-flies of the genus Urophora R. - D. (Diptera, Tephritidae) of Eastern Palaearctic. III. Key to palaearctic species. Entomologicheskoe Obozrenie, 78 (2): 464 - 482. (In Russian; English translation: Entomological Review, 1999, 79 (3): 296 - 309)." type="journal article" year="1999">Korneyev &amp; White (1999)</bibRefCitation>
. Here, the morphological details
<taxonomicName id="4C53809DFFBEFF996A0A4552FB71F8D3" baseAuthorityName="Fallen" baseAuthorityYear="1820" box="[1147,1261,1828,1853]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="aprica">
<emphasis id="B927270CFFBEFF996A0A4552FB71F8D3" box="[1147,1261,1828,1853]" italics="true" pageId="3" pageNumber="19">U. aprica</emphasis>
</taxonomicName>
are illustrated based on the original material from the
<collectingRegion id="499735FCFFBEFF996B6D4531FC35F868" country="Russia" name="Ul'yanovsk" pageId="3" pageNumber="19">Ulyanovsk Province</collectingRegion>
of
<collectingCountry id="F344BB8EFFBEFF996DBF451AFBBDF868" box="[974,1057,1900,1926]" name="Russia" pageId="3" pageNumber="19">Russia</collectingCountry>
(
<figureCitation id="1368E79BFFBEFF996A41451AFB5FF868" box="[1072,1219,1900,1926]" captionStart="Figs 10–18" captionStartId="3.[159,211,1321,1345]" captionTargetBox="[161,1442,183,1252]" captionTargetId="figure-277@3.[159,1443,181,1254]" captionTargetPageId="3" captionText="Figs 10–18. Urophora aprica (Fallén, 1820) from European Russia (Ulyanovsk Province; reared from Centaurea cyanus), females (10, 12, 15–18) and males (11, 13, 14). 10, female habitus (in lateral view); 11, male habitus (in lateral view); 12, female abdomen (in dorsal view); 13, male wing; 14, glans of phallus; 15, aculeus; 16, distal part of aculeus; 17, 18, apex of aculeus." figureDoi="http://doi.org/10.5281/zenodo.10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="3" pageNumber="19">Figs 10–18</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2CAB96FFBFFF986EE144D7FCF1F917" ID-DOI="http://doi.org/10.5281/zenodo.10135760" ID-Zenodo-Dep="10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="2" pageNumber="18" startId="2.[144,197,1697,1721]" targetBox="[148,1425,184,1633]" targetPageId="2" targetType="figure">
<paragraph id="8BECFB1EFFBFFF986EE144D7FCF1F917" blockId="2.[144,1429,1696,1785]" pageId="2" pageNumber="18">
<emphasis id="B927270CFFBFFF986EE144D7FF62F957" bold="true" box="[144,254,1697,1721]" pageId="2" pageNumber="18">Figs 1–9.</emphasis>
<taxonomicName id="4C53809DFFBFFF986F7544D6FE72F957" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[260,494,1696,1721]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="18" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFBFFF986F7544D6FE72F957" box="[260,494,1696,1721]" italics="true" pageId="2" pageNumber="18">Urophora sevanensis</emphasis>
</taxonomicName>
<emphasis id="B927270CFFBFFF986F8444D7FDCCF957" bold="true" box="[501,592,1697,1721]" pageId="2" pageNumber="18">
<taxonomicNameLabel id="A2149A77FFBFFF986F8444D7FDCCF957" box="[501,592,1697,1721]" pageId="2" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratypes (1–8) and holotype (9).
<emphasis id="B927270CFFBFFF986D9744D7FC68F957" bold="true" box="[998,1012,1697,1721]" pageId="2" pageNumber="18">1</emphasis>
, male habitus (in lateral view);
<emphasis id="B927270CFFBFFF986B2A44D7FAF5F957" bold="true" box="[1371,1385,1697,1721]" pageId="2" pageNumber="18">2</emphasis>
, female habitus (in lateral view);
<emphasis id="B927270CFFBFFF986F9844B7FE6BF937" bold="true" box="[489,503,1729,1753]" pageId="2" pageNumber="18">3</emphasis>
, male wing;
<emphasis id="B927270CFFBFFF986CF444B7FD0FF937" bold="true" box="[645,659,1729,1753]" pageId="2" pageNumber="18">4</emphasis>
, female wing;
<emphasis id="B927270CFFBFFF986D4744B7FCD8F937" bold="true" box="[822,836,1729,1753]" pageId="2" pageNumber="18">5</emphasis>
, aculeus;
<emphasis id="B927270CFFBFFF986DC344B7FC5CF937" bold="true" box="[946,960,1729,1753]" pageId="2" pageNumber="18">6</emphasis>
, apex of aculeus;
<emphasis id="B927270CFFBFFF986AF244B7FB0CF937" bold="true" box="[1155,1168,1729,1753]" pageId="2" pageNumber="18">7</emphasis>
, female thorax (in dorsal view);
<emphasis id="B927270CFFBFFF986F734497FE8CF917" bold="true" box="[258,272,1761,1785]" pageId="2" pageNumber="18">8</emphasis>
, glans of phallus;
<emphasis id="B927270CFFBFFF986FAA4497FE75F917" bold="true" box="[475,489,1761,1785]" pageId="2" pageNumber="18">9</emphasis>
, female abdomen (in dorsal view).
</paragraph>
</caption>
<caption id="DF2CAB96FFBEFF996EEE475FFDC8FA4C" ID-DOI="http://doi.org/10.5281/zenodo.10135764" ID-Zenodo-Dep="10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="3" pageNumber="19" startId="3.[159,211,1321,1345]" targetBox="[161,1442,183,1252]" targetPageId="3" targetType="figure">
<paragraph id="8BECFB1EFFBEFF996EEE475FFDC8FA4C" blockId="3.[159,1444,1321,1442]" pageId="3" pageNumber="19">
<emphasis id="B927270CFFBEFF996EEE475FFEB6FAAC" bold="true" box="[159,298,1321,1346]" pageId="3" pageNumber="19">Figs 10–18.</emphasis>
<taxonomicName id="4C53809DFFBEFF996F42475FFD00FAAC" authority="(Fallen, 1820)" baseAuthorityName="Fallen" baseAuthorityYear="1820" box="[307,668,1321,1346]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="19" phylum="Arthropoda" rank="species" species="aprica">
<emphasis id="B927270CFFBEFF996F42475FFE73FAAF" box="[307,495,1321,1346]" italics="true" pageId="3" pageNumber="19">Urophora aprica</emphasis>
(Fallén, 1820)
</taxonomicName>
from European Russia (Ulyanovsk Province; reared from
<taxonomicName id="4C53809DFFBEFF996B39475CFE84FA8F" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="19" phylum="Tracheophyta" rank="species" species="cyanus">
<emphasis id="B927270CFFBEFF996B39475CFE84FA8F" italics="true" pageId="3" pageNumber="19">Centaurea cyanus</emphasis>
</taxonomicName>
), females (10, 12, 15–18) and males (11, 13, 14).
<emphasis id="B927270CFFBEFF996D3D473CFCFAFA8C" bold="true" box="[844,870,1354,1378]" pageId="3" pageNumber="19">10</emphasis>
, female habitus (in lateral view);
<emphasis id="B927270CFFBEFF996A9B473CFA98FA8C" bold="true" box="[1258,1284,1354,1378]" pageId="3" pageNumber="19">11</emphasis>
, male habitus (in lateral view);
<emphasis id="B927270CFFBEFF996F16471CFE1FFA6C" bold="true" box="[359,387,1386,1410]" pageId="3" pageNumber="19">12</emphasis>
, female abdomen (in dorsal view);
<emphasis id="B927270CFFBEFF996D64471CFCACFA6C" bold="true" box="[789,816,1386,1410]" pageId="3" pageNumber="19">13</emphasis>
, male wing;
<emphasis id="B927270CFFBEFF996DB5471CFC43FA6C" bold="true" box="[964,991,1386,1410]" pageId="3" pageNumber="19">14</emphasis>
, glans of phallus;
<emphasis id="B927270CFFBEFF996ADE471CFB56FA6C" bold="true" box="[1199,1226,1386,1410]" pageId="3" pageNumber="19">15</emphasis>
, aculeus;
<emphasis id="B927270CFFBEFF996B4A471CFACAFA6C" bold="true" box="[1339,1366,1386,1410]" pageId="3" pageNumber="19">16</emphasis>
, distal part of aculeus;
<emphasis id="B927270CFFBEFF996F2247FCFEF1FA4C" bold="true" box="[339,365,1418,1442]" pageId="3" pageNumber="19">17</emphasis>
,
<emphasis id="B927270CFFBEFF996F0847FCFE08FA4C" bold="true" box="[377,404,1418,1442]" pageId="3" pageNumber="19">18</emphasis>
, apex of aculeus.
</paragraph>
</caption>
<caption id="DF2CAB96FFB9FF9E6EE14763FC95FA83" ID-DOI="http://doi.org/10.5281/zenodo.10135766" ID-Zenodo-Dep="10135766" httpUri="https://zenodo.org/record/10135766/files/figure.png" pageId="4" pageNumber="20" startId="4.[144,195,1301,1325]" targetBox="[146,1425,185,1214]" targetPageId="4" targetType="figure">
<paragraph id="8BECFB1EFFB9FF9E6EE14763FC95FA83" blockId="4.[143,1428,1300,1389]" pageId="4" pageNumber="20">
<emphasis id="B927270CFFB9FF9E6EE14763FE86FAC3" bold="true" box="[144,282,1301,1325]" pageId="4" pageNumber="20">Figs 19–26.</emphasis>
<taxonomicName id="4C53809DFFB9FF9E6F514762FD2FFAC3" authority="Korneyev, 1985" authorityName="Korneyev" authorityYear="1985" box="[288,691,1300,1325]" class="Insecta" family="Tephritidae" genus="Terellia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="ermolenkoi">
<emphasis id="B927270CFFB9FF9E6F514762FE64FAC3" box="[288,504,1300,1325]" italics="true" pageId="4" pageNumber="20">Terellia ermolenkoi</emphasis>
<bibRefCitation id="EFC286EFFFB9FF9E6F8C4763FD2FFAC3" author="Korneyev V. A." box="[509,691,1300,1325]" pageId="4" pageNumber="20" pagination="35 - 55" refId="ref3362" refString="Korneyev V. A. 1985. Fruit flies of the tribe Terelliini Hendel, 1927 (Diptera, Tephritidae) of the fauna of the USSR. Entomologicheskoe Obozrenie, 64 (3): 626 - 644. (In Russian; English translation: Entomological Review, 1986, 65 (1): 35 - 55)." type="journal article" year="1985">Korneyev, 1985</bibRefCitation>
</taxonomicName>
, females (19, 20, 22, 25, 26) and males (20, 21, 23, 24).
<emphasis id="B927270CFFB9FF9E6B524763FAA2FAC3" bold="true" box="[1315,1342,1301,1325]" pageId="4" pageNumber="20">19</emphasis>
, female habitus (in lateral view);
<emphasis id="B927270CFFB9FF9E6FC94743FE4FFAA3" bold="true" box="[440,467,1333,1357]" pageId="4" pageNumber="20">20</emphasis>
, male wing;
<emphasis id="B927270CFFB9FF9E6C184743FD1FFAA3" bold="true" box="[617,643,1333,1357]" pageId="4" pageNumber="20">21</emphasis>
, glans of phallus;
<emphasis id="B927270CFFB9FF9E6D264743FCEEFAA3" bold="true" box="[855,882,1333,1357]" pageId="4" pageNumber="20">22</emphasis>
, female abdomen (in dorsal view);
<emphasis id="B927270CFFB9FF9E6B784743FAB8FAA3" bold="true" box="[1289,1316,1333,1357]" pageId="4" pageNumber="20">23</emphasis>
,
<emphasis id="B927270CFFB9FF9E6B424743FACCFAA3" bold="true" box="[1331,1360,1333,1357]" pageId="4" pageNumber="20">24</emphasis>
, male thorax (in dorsal view);
<emphasis id="B927270CFFB9FF9E6FD34723FE23FA83" bold="true" box="[418,447,1365,1389]" pageId="4" pageNumber="20">25</emphasis>
, aculeus;
<emphasis id="B927270CFFB9FF9E6C5C4723FDD5FA83" bold="true" box="[557,585,1365,1389]" pageId="4" pageNumber="20">26</emphasis>
, apex of aculeus.
</paragraph>
</caption>
<subSubSection id="C349A895FFB9FF9E6EC447AFFA09F868" pageId="4" pageNumber="20" type="key">
<paragraph id="8BECFB1EFFB9FF9E6EC447AFFE95F9D2" blockId="4.[144,764,1497,1926]" pageId="4" pageNumber="20">
The following key may be used to differentiate
<taxonomicName id="4C53809DFFB9FF9E6EB04788FEF0F9F9" ID-CoL="BDPJK" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[193,364,1534,1559]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFB9FF9E6EB04788FEF0F9F9" box="[193,364,1534,1559]" italics="true" pageId="4" pageNumber="20">U. sevanensis</emphasis>
</taxonomicName>
<emphasis id="B927270CFFB9FF9E6F094788FE7DF9F6" bold="true" box="[376,481,1534,1560]" pageId="4" pageNumber="20">
<taxonomicNameLabel id="A2149A77FFB9FF9E6F094788FE7DF9F6" box="[376,481,1534,1560]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C53809DFFB9FF9E6C454788FD5DF9F9" ID-CoL="8FL2V" authorityName="Korneyev et White" authorityYear="1996" box="[564,705,1534,1559]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="trinervii">
<emphasis id="B927270CFFB9FF9E6C454788FD5DF9F9" box="[564,705,1534,1559]" italics="true" pageId="4" pageNumber="20">U. trinervii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C53809DFFB9FF9E6EE14455FE9DF9D2" ID-CoL="89C4Y" baseAuthorityName="Fallen" baseAuthorityYear="1820" box="[144,257,1571,1596]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="aprica">
<emphasis id="B927270CFFB9FF9E6EE14455FE9DF9D2" box="[144,257,1571,1596]" italics="true" pageId="4" pageNumber="20">U. aprica</emphasis>
</taxonomicName>
:
</paragraph>
<keyStep id="30A7E3BBFFB9FF9E6EE14433FD67F8CD" pageId="4" pageNumber="20">
<paragraph id="8BECFB1EFFB9FF9E6EE14433FD60F971" blockId="4.[144,764,1497,1926]" pageId="4" pageNumber="20">
<keyLead id="30A2582BFFB9FF9E6EE14433FD60F971" pageId="4" pageNumber="20">
<key id="FFC2AC2FFFB9FF9E6EE14433FD60F971" pageId="4" pageNumber="20">
1. Wing pattern characterised by reduced, faint crossbands. Larvae develop in the capitula of
<taxonomicName id="4C53809DFFB9FF9E6CF74413FEB9F970" ID-CoL="4NGQC" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="20" phylum="Tracheophyta" rank="species" species="trinervius">
<emphasis id="B927270CFFB9FF9E6CF74413FEB9F970" italics="true" pageId="4" pageNumber="20">Psephellus trinervius</emphasis>
</taxonomicName>
..........................
<taxonomicName id="4C53809DFFB9FF9E6C0144F1FD60F971" ID-CoL="8FL2V" authorityName="Korneyev et White" authorityYear="1996" box="[624,764,1671,1695]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="trinervii">
<emphasis id="B927270CFFB9FF9E6C0144F1FD60F971" bold="true" box="[624,764,1671,1695]" italics="true" pageId="4" pageNumber="20">U. trinervii</emphasis>
</taxonomicName>
</key>
</keyLead>
</paragraph>
<paragraph id="8BECFB1EFFB9FF9E6EE144DEFD67F8CD" blockId="4.[144,764,1497,1926]" pageId="4" pageNumber="20">
<keyLead id="30A2582BFFB9FF9E6EE144DEFD67F8CD" pageId="4" pageNumber="20">– Wing with four distinct crossbands: separated subbasal and discal crossbands, separated discal and preapical crossbands, anteriorly connected preapical and apical crossbands...................... 2</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BECFB1EFFB9FF9E6EE1455AFA09F98E" blockId="4.[144,764,1497,1926]" lastBlockId="4.[809,1430,1496,1926]" pageId="4" pageNumber="20">
<keyLead id="30A2582BFFB9FF9E6EE1455AFA09F98E" pageId="4" pageNumber="20">
<keyStep id="30A7E3BBFFB9FF9E6EE1455AFC31FA1F" pageId="4" pageNumber="20">
2. Male posterior process of subbasal crossband long (
<figureCitation id="1368E79BFFB9FF9E6ECF453AFE98F88B" box="[190,260,1868,1893]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="4" pageNumber="20">Fig. 3</figureCitation>
). Usually male femora entirely orange. Aculeus with distinct preapical primary and small secondary
</keyStep>
steps (
<figureCitation id="1368E79BFFB9FF9E6D8C47AEFBD8FA1F" box="[1021,1092,1496,1521]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="4" pageNumber="20">Fig. 6</figureCitation>
). Apical notch small (
<figureCitation id="1368E79BFFB9FF9E6B4E47AEFA1AFA1F" box="[1343,1414,1496,1521]" captionStart="Figs 1–9" captionStartId="2.[144,197,1697,1721]" captionTargetBox="[148,1425,184,1633]" captionTargetId="figure-86@2.[144,1427,181,1636]" captionTargetPageId="2" captionText="Figs 1–9. Urophora sevanensis sp. nov., paratypes (1–8) and holotype (9). 1, male habitus (in lateral view); 2, female habitus (in lateral view); 3, male wing; 4, female wing; 5, aculeus; 6, apex of aculeus; 7, female thorax (in dorsal view); 8, glans of phallus; 9, female abdomen (in dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10135760" httpUri="https://zenodo.org/record/10135760/files/figure.png" pageId="4" pageNumber="20">Fig. 6</figureCitation>
). Aculeus length
<quantity id="4CAB56FBFFB9FF9E6A774788FBFDF9FB" box="[1030,1121,1534,1558]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="20" unit="mm" value="2.7">2.7 mm</quantity>
. Larvae develop in the capitula of
<taxonomicName id="4C53809DFFB9FF9E6DC54454FBD3F9D4" box="[948,1103,1570,1595]" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B927270CFFB9FF9E6DC54454FBB6F9D5" box="[948,1066,1570,1595]" italics="true" pageId="4" pageNumber="20">Psephellus</emphasis>
sp.
</taxonomicName>
..................................................
<emphasis id="B927270CFFB9FF9E6AFA443EFA09F98E" bold="true" box="[1163,1429,1607,1632]" pageId="4" pageNumber="20">
<taxonomicName id="4C53809DFFB9FF9E6AFA443EFAAEF9B1" ID-CoL="BDPJK" authority="Evstigneev, 2023" authorityName="Evstigneev" authorityYear="2023" box="[1163,1330,1607,1631]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="sevanensis" status="sp. nov.">
<emphasis id="B927270CFFB9FF9E6AFA443EFAAEF9B1" bold="true" box="[1163,1330,1607,1631]" italics="true" pageId="4" pageNumber="20">U. sevanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2149A77FFB9FF9E6B48443EFA09F98E" box="[1337,1429,1608,1632]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BECFB1EFFB9FF9E6D58441BFA09F868" blockId="4.[809,1430,1496,1926]" pageId="4" pageNumber="20">
<keyLead id="30A2582BFFB9FF9E6D58441BFA09F868" pageId="4" pageNumber="20">
– Male posterior process of subbasal crossband short (
<figureCitation id="1368E79BFFB9FF9E6D2944E6FC78F947" box="[856,996,1680,1705]" captionStart="Figs 10–18" captionStartId="3.[159,211,1321,1345]" captionTargetBox="[161,1442,183,1252]" captionTargetId="figure-277@3.[159,1443,181,1254]" captionTargetPageId="3" captionText="Figs 10–18. Urophora aprica (Fallén, 1820) from European Russia (Ulyanovsk Province; reared from Centaurea cyanus), females (10, 12, 15–18) and males (11, 13, 14). 10, female habitus (in lateral view); 11, male habitus (in lateral view); 12, female abdomen (in dorsal view); 13, male wing; 14, glans of phallus; 15, aculeus; 16, distal part of aculeus; 17, 18, apex of aculeus." figureDoi="http://doi.org/10.5281/zenodo.10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="4" pageNumber="20">Figs 11, 13</figureCitation>
). Male femora with extensive black marks up to predominantly black, orange apically (
<figureCitation id="1368E79BFFB9FF9E6D2644ACFC36F91D" box="[855,938,1754,1779]" captionStart="Figs 10–18" captionStartId="3.[159,211,1321,1345]" captionTargetBox="[161,1442,183,1252]" captionTargetId="figure-277@3.[159,1443,181,1254]" captionTargetPageId="3" captionText="Figs 10–18. Urophora aprica (Fallén, 1820) from European Russia (Ulyanovsk Province; reared from Centaurea cyanus), females (10, 12, 15–18) and males (11, 13, 14). 10, female habitus (in lateral view); 11, male habitus (in lateral view); 12, female abdomen (in dorsal view); 13, male wing; 14, glans of phallus; 15, aculeus; 16, distal part of aculeus; 17, 18, apex of aculeus." figureDoi="http://doi.org/10.5281/zenodo.10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="4" pageNumber="20">Fig. 11</figureCitation>
). Aculeus with two pairs of very prominent preapical steps (
<figureCitation id="1368E79BFFB9FF9E6A7D4489FB0EF8F6" box="[1036,1170,1791,1816]" captionStart="Figs 10–18" captionStartId="3.[159,211,1321,1345]" captionTargetBox="[161,1442,183,1252]" captionTargetId="figure-277@3.[159,1443,181,1254]" captionTargetPageId="3" captionText="Figs 10–18. Urophora aprica (Fallén, 1820) from European Russia (Ulyanovsk Province; reared from Centaurea cyanus), females (10, 12, 15–18) and males (11, 13, 14). 10, female habitus (in lateral view); 11, male habitus (in lateral view); 12, female abdomen (in dorsal view); 13, male wing; 14, glans of phallus; 15, aculeus; 16, distal part of aculeus; 17, 18, apex of aculeus." figureDoi="http://doi.org/10.5281/zenodo.10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="4" pageNumber="20">Figs 17, 18</figureCitation>
). Apical notch deeper (
<figureCitation id="1368E79BFFB9FF9E6D264552FC41F8D2" box="[855,989,1828,1853]" captionStart="Figs 10–18" captionStartId="3.[159,211,1321,1345]" captionTargetBox="[161,1442,183,1252]" captionTargetId="figure-277@3.[159,1443,181,1254]" captionTargetPageId="3" captionText="Figs 10–18. Urophora aprica (Fallén, 1820) from European Russia (Ulyanovsk Province; reared from Centaurea cyanus), females (10, 12, 15–18) and males (11, 13, 14). 10, female habitus (in lateral view); 11, male habitus (in lateral view); 12, female abdomen (in dorsal view); 13, male wing; 14, glans of phallus; 15, aculeus; 16, distal part of aculeus; 17, 18, apex of aculeus." figureDoi="http://doi.org/10.5281/zenodo.10135764" httpUri="https://zenodo.org/record/10135764/files/figure.png" pageId="4" pageNumber="20">Figs 17, 18</figureCitation>
). Aculeus length
<quantity id="4CAB56FBFFB9FF9E6ADD4552FAABF8D2" box="[1196,1335,1828,1852]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="4" pageNumber="20" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 mm</quantity>
. Larvae develop in the capitula of
<taxonomicName id="4C53809DFFB9FF9E6A03453FFAA3F88F" ID-CoL="69GPL" box="[1138,1343,1865,1889]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="20" phylum="Tracheophyta" rank="species" species="cyanus">
<emphasis id="B927270CFFB9FF9E6A03453FFAA3F88F" box="[1138,1343,1865,1889]" italics="true" pageId="4" pageNumber="20">Centaurea cyanus</emphasis>
</taxonomicName>
...........................................
<taxonomicName id="4C53809DFFB9FF9E6B534518FA09F868" ID-CoL="89C4Y" baseAuthorityName="Fallen" baseAuthorityYear="1820" box="[1314,1429,1902,1926]" class="Insecta" family="Tephritidae" genus="Urophora" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="aprica">
<emphasis id="B927270CFFB9FF9E6B534518FA09F868" bold="true" box="[1314,1429,1902,1926]" italics="true" pageId="4" pageNumber="20">U. aprica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFB8FF9F6EB442CCFEF2FE6B" pageId="5" pageNumber="21" type="biology_ecology">
<paragraph id="8BECFB1EFFB8FF9F6EB442CCFEF2FE6B" blockId="5.[159,779,185,567]" pageId="5" pageNumber="21">
<emphasis id="B927270CFFB8FF9F6EB442CCFED4FF3D" box="[197,328,185,211]" italics="true" pageId="5" pageNumber="21">Host plant</emphasis>
. Larvae of the new species develop in the capitula of
<taxonomicName id="4C53809DFFB8FF9F6FF342AAFDB2FF19" box="[386,558,220,247]" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="21" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B927270CFFB8FF9F6FF342AAFD9FFF18" box="[386,515,220,246]" italics="true" pageId="5" pageNumber="21">Psephellus</emphasis>
sp.
</taxonomicName>
(Fig. 27, see Addenda).
<taxonomicName id="4C53809DFFB8FF9F6F764376FD52FEF5" authority="Korneyev, 1985" authorityName="Korneyev" authorityYear="1985" box="[263,718,255,283]" class="Insecta" family="Tephritidae" genus="Terellia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="ermolenkoi">
<emphasis id="B927270CFFB8FF9F6F764376FE64FEF4" box="[263,504,256,282]" italics="true" pageId="5" pageNumber="21">Terellia ermolenkoi</emphasis>
<bibRefCitation id="EFC286EFFFB8FF9F6C744377FD52FEF5" author="Korneyev V. A." box="[517,718,255,283]" pageId="5" pageNumber="21" pagination="35 - 55" refId="ref3362" refString="Korneyev V. A. 1985. Fruit flies of the tribe Terelliini Hendel, 1927 (Diptera, Tephritidae) of the fauna of the USSR. Entomologicheskoe Obozrenie, 64 (3): 626 - 644. (In Russian; English translation: Entomological Review, 1986, 65 (1): 35 - 55)." type="journal article" year="1985">Korneyev, 1985</bibRefCitation>
</taxonomicName>
was reared from the capitula of the same species of
<taxonomicName id="4C53809DFFB8FF9F6EEE4331FEBCFE8F" box="[159,288,327,353]" class="Magnoliopsida" family="Asteraceae" genus="Psephellus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B927270CFFB8FF9F6EEE4331FEBCFE8F" box="[159,288,327,353]" italics="true" pageId="5" pageNumber="21">Psephellus</emphasis>
</taxonomicName>
. Information about this fly species is presented below.
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFB8FF9F6EB443F9FD78FE1E" pageId="5" pageNumber="21" type="etymology">
<paragraph id="8BECFB1EFFB8FF9F6EB443F9FD78FE1E" blockId="5.[159,779,185,567]" pageId="5" pageNumber="21">
<emphasis id="B927270CFFB8FF9F6EB443F9FEDBFE47" box="[197,327,399,425]" italics="true" pageId="5" pageNumber="21">Etymology</emphasis>
. The species name is an adjective derived from Lake Sevan (Սևան in Armenian), on the coast of which the
<typeStatus id="54E845BCFFB8FF9F6FC943A0FE6DFE1E" box="[440,497,470,496]" pageId="5" pageNumber="21">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="C349A895FFB8FF9F6EB4438FFD15FDD9" pageId="5" pageNumber="21" type="distribution">
<paragraph id="8BECFB1EFFB8FF9F6EB4438FFD15FDD9" blockId="5.[159,779,185,567]" pageId="5" pageNumber="21">
<emphasis id="B927270CFFB8FF9F6EB4438FFEC2FDFD" box="[197,350,505,531]" italics="true" pageId="5" pageNumber="21">Distribution</emphasis>
. The new species is known from the
<collectingRegion id="499735FCFFB8FF9F6EBF406BFE73FDD9" box="[206,495,541,567]" country="Armenia" name="Geghark'unik'" pageId="5" pageNumber="21">Gegharkunik Province</collectingRegion>
of
<collectingCountry id="F344BB8EFFB8FF9F6C65406BFD19FDD9" box="[532,645,541,567]" name="Armenia" pageId="5" pageNumber="21">Armenia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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