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<document id="5CC5476FE2A94620110B0A04C934E984" ID-CLB-Dataset="102626" ID-DOI="10.11646/zootaxa.5168.1.1" ID-GBIF-Dataset="4e90e71f-ff39-4569-8fdf-6f8f5edb73d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6876161" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658417337616" checkinUser="plazi" docAuthor="Mather, Ellen K., Lee, Michael S. Y. &amp; Worthy, Trevor H." docDate="2022" docId="DC3487FAFFD0B04AFF075198FF32F90E" docLanguage="en" docName="zootaxa.5168.1.1.pdf" docOrigin="Zootaxa 5168 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gypaetinae Storr 1784" docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="200DFF82FFDDB047FF905236FB6AFFEF" masterDocTitle="A new look at an old Australian raptor places “ Taphaetus ” lacertosus de Vis 1905 in the Old World vultures (Accipitridae: Aegypiinae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1699361215756" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8D03870632942BB9E8D7929D18D7794A">A new look at an old Australian raptor places “ Taphaetus ” lacertosus de Vis 1905 in the Old World vultures (Accipitridae: Aegypiinae)</mods:title>
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<mods:namePart id="D96093B3C8D3DDE5CFB03E12F19ADA99">Mather, Ellen K.</mods:namePart>
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<mods:affiliation id="C61C4D29BBF77B564D5B3A7718A7EE4F">Palaeontology Group, College of Science and Engineering, Flinders University, GPO 2100, Adelaide, SA 5001, Australia &amp; ellen. mather @ flinders. edu. au, math 0083 @ flinders. edu. au, https: // orcid. org / 0000 - 0001 - 9437 - 1395</mods:affiliation>
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<mods:namePart id="F23218E5037CD40554DC50D1A96C251E">Lee, Michael S. Y.</mods:namePart>
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<mods:affiliation id="C84B5996BF2EED97E5285C260A8DD3B8">Palaeontology Group, College of Science and Engineering, Flinders University, GPO 2100, Adelaide, SA 5001, Australia &amp; Earth Sciences Section, South Australian Museum, North Terrace, Adelaide 5000, Australia &amp; mike. lee @ flinders. edu. au; https: // orcid. org / 0000 - 0002 - 3905 - 0887</mods:affiliation>
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<mods:namePart id="33D13DEC06094613256F4BEAA6A21BA3">Worthy, Trevor H.</mods:namePart>
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<mods:affiliation id="26D17BA54FE9F1518922F386B3B642E6">Palaeontology Group, College of Science and Engineering, Flinders University, GPO 2100, Adelaide, SA 5001, Australia &amp; trevor. worthy @ flinders. edu. au; https: // orcid. org / 0000 - 0001 - 7047 - 4680</mods:affiliation>
<mods:nameIdentifier id="0CC2298ABBEA9414CA673EC10F457D68" type="email">trevor.worthy@flinders.edu.au</mods:nameIdentifier>
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<mods:title id="77C71A72A4261CAF4080530AACCD9814">Zootaxa</mods:title>
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<mods:date id="D18A7E32838643DF9AC06E6ACF7ACACC">2022</mods:date>
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<mods:number id="89029C5C877A45BA82C4BFEFD5A001C7">2022-07-20</mods:number>
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<treatment id="DC3487FAFFD0B04AFF075198FF32F90E" ID-DOI="http://doi.org/10.5281/zenodo.6884270" ID-GBIF-Taxon="197839018" ID-Zenodo-Dep="6884270" LSID="urn:lsid:plazi:treatment:DC3487FAFFD0B04AFF075198FF32F90E" httpUri="http://treatment.plazi.org/id/DC3487FAFFD0B04AFF075198FF32F90E" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="1C876567FFD0B04AFF075198FA4DFC27" box="[151,295,942,968]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="542236ECFFD0B04AFF075198FA4DFC27" blockId="13.[151,1437,942,1761]" box="[151,295,942,968]" pageId="13" pageNumber="14">
<heading id="0F6A8180FFD0B04AFF075198FA4DFC27" bold="true" box="[151,295,942,968]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="66E9EAFEFFD0B04AFF075198FA4DFC27" bold="true" box="[151,295,942,968]" pageId="13" pageNumber="14">
<taxonomicName id="939D4D6FFFD0B04AFF075198FA48FC27" authorityName="Storr" authorityYear="1784" box="[151,290,942,968]" class="Aves" family="Accipitridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="subFamily" subFamily="Gypaetinae">Gypaetinae</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1C876567FFD0B04AFF5751E5FF32F90E" pageId="13" pageNumber="14" type="description">
<paragraph id="542236ECFFD0B04AFF5751E5F956FB2A" blockId="13.[151,1437,942,1761]" pageId="13" pageNumber="14">
The gypaetines share with the fossil the states of the following six tarsometatarsi characters:
<emphasis id="66E9EAFEFFD0B04AFB4C51E4FF96FC03" bold="true" box="[1244,1276,978,1004]" pageId="13" pageNumber="14">(1)</emphasis>
the eminentia intercotylaris projects proximally as in the fossil in all species except
<taxonomicName id="939D4D6FFFD0B04AFC0B51C1FF1BFBFF" box="[923,1137,1015,1041]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFC0B51C1FF1BFBFF" box="[923,1137,1015,1041]" italics="true" pageId="13" pageNumber="14">
Polyboroides
<typeStatus id="8B26884EFFD0B04AFBA651C1FF1BFBFF" box="[1078,1137,1015,1040]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
, where it is hyper-protruding proximally;
<emphasis id="66E9EAFEFFD0B04AFEC0562CFA1BFBDB" bold="true" box="[336,369,1050,1076]" pageId="13" pageNumber="14">(2)</emphasis>
a narrow sulcus hypotarsus;
<emphasis id="66E9EAFEFFD0B04AFD52562CF989FBDB" bold="true" box="[706,739,1050,1076]" pageId="13" pageNumber="14">(3)</emphasis>
the base of the sulcus hypotarsus is set plantar to the sulcus flexorius, though in
<taxonomicName id="939D4D6FFFD0B04AFEEC5609F9EAFBB6" box="[380,640,1087,1113]" class="Aves" family="Accipitridae" genus="Gypohierax" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="66E9EAFEFFD0B04AFEEC5609F9EAFBB6" box="[380,640,1087,1113]" italics="true" pageId="13" pageNumber="14">Gypohierax angolensis</emphasis>
</taxonomicName>
it is to a lesser degree than in the fossil;
<emphasis id="66E9EAFEFFD0B04AFBDD5608FF04FBB7" bold="true" box="[1101,1134,1086,1112]" pageId="13" pageNumber="14">(8)</emphasis>
a deep fossa infracotylaris dorsalis;
<emphasis id="66E9EAFEFFD0B04AFF6C5654FA41FB93" bold="true" box="[252,299,1122,1148]" pageId="13" pageNumber="14">(21)</emphasis>
the sulcus for the m. abductor digit IV being broad;
<emphasis id="66E9EAFEFFD0B04AFCFF5654F8F4FB93" bold="true" box="[879,926,1122,1148]" pageId="13" pageNumber="14">(22)</emphasis>
and the trochlea metatarsi II is slightly longer than trochlea metatarsi IV in
<taxonomicName id="939D4D6FFFD0B04AFE4956B1F9B1FB4E" box="[473,731,1159,1185]" class="Aves" family="Accipitridae" genus="Gypohierax" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="66E9EAFEFFD0B04AFE4956B1F9B1FB4E" box="[473,731,1159,1185]" italics="true" pageId="13" pageNumber="14">Gypohierax angolensis</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFC9F56B1F80EFB4F" box="[783,868,1159,1184]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFC9F56B1F80EFB4F" box="[783,868,1159,1184]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFCB956B1F80EFB4F" box="[809,868,1159,1184]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
and is longer by at least half its length in
<taxonomicName id="939D4D6FFFD0B04AFABD56B1FA40FB2B" authorityName="SAMA B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Aves" family="Accipitridae" genus="Neophron" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="percnopterus">
<emphasis id="66E9EAFEFFD0B04AFABD56B1FA40FB2B" italics="true" pageId="13" pageNumber="14">Neophron percnopterus</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFEF0569DF95FFB2A" authorityName="NHMUK S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[352,565,1195,1221]" class="Aves" family="Accipitridae" genus="Gypaetus" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="barbatus">
<emphasis id="66E9EAFEFFD0B04AFEF0569DF95FFB2A" box="[352,565,1195,1221]" italics="true" pageId="13" pageNumber="14">Gypaetus barbatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="542236ECFFD0B04AFF5756F9FF32F90E" blockId="13.[151,1437,942,1761]" pageId="13" pageNumber="14">
Tarsometatarsi of gypaetines differ from the fossil as follows:
<emphasis id="66E9EAFEFFD0B04AFC0256F8F8D9FB07" bold="true" box="[914,947,1230,1256]" pageId="13" pageNumber="14">(5)</emphasis>
the notch for the nervus peroneus forms a shallow yet distinct notch in all species except
<taxonomicName id="939D4D6FFFD0B04AFD3F56C5F8EFFAE2" authorityName="NHMUK S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[687,901,1267,1293]" class="Aves" family="Accipitridae" genus="Gypaetus" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="barbatus">
<emphasis id="66E9EAFEFFD0B04AFD3F56C5F8EFFAE2" box="[687,901,1267,1293]" italics="true" pageId="13" pageNumber="14">Gypaetus barbatus</emphasis>
</taxonomicName>
, where it is deep;
<emphasis id="66E9EAFEFFD0B04AFBC056C4FF1AFAE3" bold="true" box="[1104,1136,1266,1292]" pageId="13" pageNumber="14">(6)</emphasis>
the fossa parahypotarsalis lateralis spans a quarter of the shaft length or less in all species except
<taxonomicName id="939D4D6FFFD0B04AFC595721FFB9FADF" authorityName="SAMA B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[969,1235,1303,1329]" class="Aves" family="Accipitridae" genus="Neophron" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="percnopterus">
<emphasis id="66E9EAFEFFD0B04AFC595721FFB9FADF" box="[969,1235,1303,1329]" italics="true" pageId="13" pageNumber="14">Neophron percnopterus</emphasis>
</taxonomicName>
, where it spans a third of shaft length;
<emphasis id="66E9EAFEFFD0B04AFE19570CFAC0FABB" bold="true" box="[393,426,1338,1364]" pageId="13" pageNumber="14">(9)</emphasis>
the impressio ligamentum collateralis lateralis is indistinct in all species except
<taxonomicName id="939D4D6FFFD0B04AFAAD570DFEFEFABB" box="[1341,1428,1339,1364]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFAAD570DFEFEFABB" box="[1341,1428,1339,1364]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFAC9570DFEFEFABB" box="[1369,1428,1339,1364]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
;
<emphasis id="66E9EAFEFFD0B04AFF075768FBAFFA97" bold="true" box="[151,197,1374,1400]" pageId="13" pageNumber="14">(10)</emphasis>
the plantar facies of the hypotarsus crista lateralis is wider than it is long in all species except
<taxonomicName id="939D4D6FFFD0B04AFB4E5769FE5EFA97" box="[1246,1332,1375,1400]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFB4E5769FE5EFA97" box="[1246,1332,1375,1400]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFB695769FE5EFA97" box="[1273,1332,1375,1400]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
, where it is longer than wide;
<emphasis id="66E9EAFEFFD0B04AFEED57B4FAC7FA73" bold="true" box="[381,429,1410,1436]" pageId="13" pageNumber="14">(13)</emphasis>
the medial shaft margin is thin dorsal to the fossa parahypotarsalis medialis in all species except
<taxonomicName id="939D4D6FFFD0B04AFF765791FAF3FA2F" authorityName="SAMA B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[230,409,1447,1472]" class="Aves" family="Accipitridae" genus="Neophron" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="percnopterus">
<emphasis id="66E9EAFEFFD0B04AFF765791FAF3FA2F" box="[230,409,1447,1472]" italics="true" pageId="13" pageNumber="14">N. percnopterus</emphasis>
</taxonomicName>
, where it is thick as in the fossil;
<emphasis id="66E9EAFEFFD0B04AFC965790F85EFA2F" bold="true" box="[774,820,1446,1472]" pageId="13" pageNumber="14">(14)</emphasis>
the proximal end of the fossa parahypotarsalis medialis takes up at least a third of the shaft width in all species, and is deep in all species except
<taxonomicName id="939D4D6FFFD0B04AFBE757FDFF97FA0A" authorityName="NHMUK S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1143,1277,1483,1509]" class="Aves" family="Accipitridae" genus="Gypaetus" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="barbatus">
<emphasis id="66E9EAFEFFD0B04AFBE757FDFF97FA0A" box="[1143,1277,1483,1509]" italics="true" pageId="13" pageNumber="14">G. barbatus</emphasis>
</taxonomicName>
, in which it is shallow;
<emphasis id="66E9EAFEFFD0B04AFF6E57D8FA44F9E7" bold="true" box="[254,302,1518,1544]" pageId="13" pageNumber="14">(15)</emphasis>
the impressiones retinaculi extensorii are present as small ridges in
<taxonomicName id="939D4D6FFFD0B04AFBA857D9FFFAF9E7" box="[1080,1168,1519,1544]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFBA857D9FFFAF9E7" box="[1080,1168,1519,1544]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFBC557D9FFFAF9E7" box="[1109,1168,1519,1544]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFB5957D9FE0FF9E6" box="[1225,1381,1519,1545]" class="Aves" family="Accipitridae" genus="Gypohierax" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="66E9EAFEFFD0B04AFB5957D9FE0FF9E6" box="[1225,1381,1519,1545]" italics="true" pageId="13" pageNumber="14">G. angolensis</emphasis>
</taxonomicName>
, and practically absent in
<taxonomicName id="939D4D6FFFD0B04AFE115425F96DF9C2" authorityName="NHMUK S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[385,519,1555,1581]" class="Aves" family="Accipitridae" genus="Gypaetus" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="barbatus">
<emphasis id="66E9EAFEFFD0B04AFE115425F96DF9C2" box="[385,519,1555,1581]" italics="true" pageId="13" pageNumber="14">G. barbatus</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFDAE5425F998F9C3" authorityName="SAMA B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[574,754,1555,1580]" class="Aves" family="Accipitridae" genus="Neophron" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="percnopterus">
<emphasis id="66E9EAFEFFD0B04AFDAE5425F998F9C3" box="[574,754,1555,1580]" italics="true" pageId="13" pageNumber="14">N. percnopterus</emphasis>
</taxonomicName>
;
<emphasis id="66E9EAFEFFD0B04AFC915424F845F9C3" bold="true" box="[769,815,1554,1580]" pageId="13" pageNumber="14">(16)</emphasis>
the tuberositas m. tibialis cranialis is separated distally from the proximal foramina by a distance equivalent to its length;
<emphasis id="66E9EAFEFFD0B04AFC1E5400F8D6F9BF" bold="true" box="[910,956,1590,1616]" pageId="13" pageNumber="14">(17)</emphasis>
the tuberositas is oval in
<taxonomicName id="939D4D6FFFD0B04AFB745401FE06F9BE" authorityName="NHMUK S." baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1252,1388,1591,1617]" class="Aves" family="Accipitridae" genus="Gypaetus" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="barbatus">
<emphasis id="66E9EAFEFFD0B04AFB745401FE06F9BE" box="[1252,1388,1591,1617]" italics="true" pageId="13" pageNumber="14">G. barbatus</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFF07546DFA21F99B" authorityName="SAMA B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,331,1627,1652]" class="Aves" family="Accipitridae" genus="Neophron" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="percnopterus">
<emphasis id="66E9EAFEFFD0B04AFF07546DFA21F99B" box="[151,331,1627,1652]" italics="true" pageId="13" pageNumber="14">N. percnopterus</emphasis>
</taxonomicName>
, and an elongate narrow ridge in
<taxonomicName id="939D4D6FFFD0B04AFD53546DF873F99B" box="[707,793,1627,1652]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFD53546DF873F99B" box="[707,793,1627,1652]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFD4E546DF873F99B" box="[734,793,1627,1652]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
and
<taxonomicName id="939D4D6FFFD0B04AFCC0546DF880F99A" box="[848,1002,1627,1653]" class="Aves" family="Accipitridae" genus="Gypohierax" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="66E9EAFEFFD0B04AFCC0546DF880F99A" box="[848,1002,1627,1653]" italics="true" pageId="13" pageNumber="14">G. angolensis</emphasis>
</taxonomicName>
;
<emphasis id="66E9EAFEFFD0B04AFC68546CFF4DF99B" bold="true" box="[1016,1063,1626,1652]" pageId="13" pageNumber="14">(20)</emphasis>
the sulcus flexorius is shallow in all species except
<taxonomicName id="939D4D6FFFD0B04AFEF35449FAD0F977" box="[355,442,1663,1688]" class="Aves" family="Accipitridae" genus="Polyboroides" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typus">
<emphasis id="66E9EAFEFFD0B04AFEF35449FAD0F977" box="[355,442,1663,1688]" italics="true" pageId="13" pageNumber="14">
P.
<typeStatus id="8B26884EFFD0B04AFEEF5449FAD0F977" box="[383,442,1663,1688]" pageId="13" pageNumber="14">typus</typeStatus>
</emphasis>
</taxonomicName>
, in which it is deep;
<emphasis id="66E9EAFEFFD0B04AFD365448F9BFF977" bold="true" box="[678,725,1662,1688]" pageId="13" pageNumber="14">(22)</emphasis>
the trochlea metatarsi III has greater distal extent than trochlea metatarsi II in all species except
<taxonomicName id="939D4D6FFFD0B04AFD9A5495F864F952" box="[522,782,1699,1725]" class="Aves" family="Accipitridae" genus="Gypohierax" kingdom="Animalia" order="Accipitriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="66E9EAFEFFD0B04AFD9A5495F864F952" box="[522,782,1699,1725]" italics="true" pageId="13" pageNumber="14">Gypohierax angolensis</emphasis>
</taxonomicName>
, in which trochlea metatarsi II is longer;
<emphasis id="66E9EAFEFFD0B04AFB725494FE7AF953" bold="true" box="[1250,1296,1698,1724]" pageId="13" pageNumber="14">(27)</emphasis>
the trochlea metatarsi II is relatively broad;
<emphasis id="66E9EAFEFFD0B04AFE6754F0F94CF90F" bold="true" box="[503,550,1734,1760]" pageId="13" pageNumber="14">(28)</emphasis>
and the trochlea metatarsi IV is relatively narrow.
</paragraph>
</subSubSection>
</treatment>
</document>