<documentID-CLB-Dataset="198853"ID-DOI="http://dx.doi.org/10.3897/zookeys.1133.94530"ID-GBIF-Dataset="c5acfacc-834a-4abc-ad38-d6d2964f72f5"ID-PMC="PMC9836506"ID-Pensoft-Pub="1313-2970-1133-1"ID-Pensoft-UUID="4C6A7FCDB0B55088AED4D861AF5BE503"ID-PubMed="36760608"ID-ZooBank="D7A37FE0DC2A4ECCA6A10E873C7C7A5A"ModsDocID="1313-2970-1133-1"checkinTime="1669899240784"checkinUser="pensoft"docAuthor="Kato, Makoto, Yamamori, Luna & Imada, Yume"docDate="2022"docId="4ED5243AA8C65E0EB190495FA77138D7"docLanguage="en"docName="ZooKeys 1133: 1-164"docOrigin="ZooKeys 1133"docPubDate="2022-11-30"docSource="http://dx.doi.org/10.3897/zookeys.1133.94530"docTitle="Phytoliriomyza Hendel 1931"docType="treatment"docVersion="7"id="4C6A7FCDB0B55088AED4D861AF5BE503"lastPageNumber="1"masterDocId="4C6A7FCDB0B55088AED4D861AF5BE503"masterDocTitle="Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts"masterLastPageNumber="164"masterPageNumber="1"pageNumber="1"updateTime="1720471835971"updateUser="ExternalLinkService">
<mods:titleid="FDEBEFCB8A5B6EDE5DE465C12C468F12">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
<bibRefCitationid="08A3D21952B57A40FC52269594DEF13B"author="Frick, KE"journalOrPublisher="University of California Publications in Entomology"pageId="0"pageNumber="1"pagination="339 - 452"refId="B8"refString="Frick, KE, 1952. A generic revision of the family Agromyzidae (Diptera) with a catalogue of New World species. University of California Publications in Entomology 8: 339 - 452"title="A generic revision of the family Agromyzidae (Diptera) with a catalogue of New World species."volume="8"year="1952">Frick 1952</bibRefCitation>
<bibRefCitationid="D344DEECD44FA7634033609590825584"DOI="https://doi.org/10.5479/si.00963801.108-3407.347"author="Frick, KE"journalOrPublisher="Proceedings of the United States National Museum"pageId="0"pageNumber="1"pagination="347 - 465"refId="B9"refString="Frick, KE, 1959. Synopsis of the species of agromyzid leaf miners described from North America (Diptera). Proceedings of the United States National Museum 108 (3407): 347 - 465, DOI: https://doi.org/10.5479/si.00963801.108-3407.347"title="Synopsis of the species of agromyzid leaf miners described from North America (Diptera)."url="https://doi.org/10.5479/si.00963801.108-3407.347"volume="108"year="1959">1959</bibRefCitation>
<bibRefCitationid="ADCD931B2F9DAA52450747A4D9F12845"DOI="https://doi.org/10.4039/entm10164fv"author="Spencer, KA"journalOrPublisher="Memoirs of the Entomological Society of Canada"pageId="0"pageNumber="1"pagination="1 - 311"refId="B35"refString="Spencer, KA, 1969. The Agromyzidae of Canada and Alaska. Memoirs of the Entomological Society of Canada 64: 1 - 311, DOI: https://doi.org/10.4039/entm10164fv"title="The Agromyzidae of Canada and Alaska."url="https://doi.org/10.4039/entm10164fv"volume="64"year="1969">Spencer 1969</bibRefCitation>
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<bibRefCitationid="4B4789B23923D51F79963D6E4DFAE458"author="Frick, KE"journalOrPublisher="University of California Publications in Entomology"pageId="0"pageNumber="1"pagination="339 - 452"refId="B8"refString="Frick, KE, 1952. A generic revision of the family Agromyzidae (Diptera) with a catalogue of New World species. University of California Publications in Entomology 8: 339 - 452"title="A generic revision of the family Agromyzidae (Diptera) with a catalogue of New World species."volume="8"year="1952">Frick 1952</bibRefCitation>
<bibRefCitationid="52ECE3A9F30AAE486C77D5761E72708F"DOI="https://doi.org/10.5962/bhl.part.21862"author="Spencer, KA"journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology"pageId="0"pageNumber="1"pagination="25 - 31"refId="B34"refString="Spencer, KA, 1965. Agromyzidae. Diptera from Nepal. Bulletin of the British Museum (Natural History), Entomology 16: 25 - 31, DOI: https://doi.org/10.5962/bhl.part.21862"title="Agromyzidae. Diptera from Nepal."url="https://doi.org/10.5962/bhl.part.21862"volume="16"year="1965">Spencer 1965</bibRefCitation>
<bibRefCitationid="F2BD0CDC58B2325AEB6A5C32C6D45F84"author="von Tschirnhaus, M"journalOrPublisher="Beitraege zur Entomologie"pageId="0"pageNumber="1"pagination="551 - 579"refId="B44"refString="von Tschirnhaus, M, 1971. Unbekannte Stridulationsorgane bei Dipteren und ihre Bedeutung fuer Taxonomie und Phylogenetik der Agromyziden. Beitraege zur Entomologie 21: 551 - 579"title="Unbekannte Stridulationsorgane bei Dipteren und ihre Bedeutung fuer Taxonomie und Phylogenetik der Agromyziden."volume="21"year="1971">von Tschirnhaus 1971</bibRefCitation>
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Head yellow, frons often with pruinosity. Orbital plates more or less emerge from the plane of frons. Front orbitals three pairs; one ori directed inward and two ors directed up. 1-2(3) pairs of medio-clinate lower orbital setae, two pairs of reclinate upper orbital setae. Orbital setulae proclinate, upright or partly reclinate. Postocellar and ocellar setae well developed. First flagellomere round.
Scutum with 1+3 dorsocentrals. Acrostichal setulae in two rows, but lacking in some species. Postpronotal, two propleural, presutural, and propleural setae normal or strong. Anepisternum with a long posterior seta and with two or three shorter posterior setae. Katepisternum with one or two long setae. Legs simple, only ventroapical seta on middle tibia present. Costal vein ends at apex of M1. The ratio of ultimate and penultimate sections of M4 various.
Epandrium round apically, often with a comb of several fused tubercle-like setae and/or elongated tubercle-like setae on the inner surface; surstylus setigerous and sometimes with one or a few tubercle-like setae. Subepandrial sclerite short but very broad, connecting bases of surstyli through a special sclerite. Hypandrium very long but very thin. Hypophallus membranous often with a pair of parallel sclerites medially. Paraphallus present or absent. Mesophallus cylindrical and well sclerotized. Distiphallus with basal part formed by usually a paired sclerite and an apical part, which terminates in paired tubes or in extremely long less sclerotized tubules. Ejaculatory apodeme variable, but basal part usually broad and blade not too large.
<subSubSectionid="653E6C07A1426BB360444CB578F5F94B"pageId="0"pageNumber="1"type="classification of phytoliriomyza species associated with bryophytes">
. In addition to these characteristics of adult males, we found characteristics of adult females unique to the bryophyte-associated species, in which tergite 10 was trifurcate or cruciform and each cercus bears two stout, apical, trichoid sensilla.
<paragraphid="66BC936B5888030856C614A82F5A8F62"pageId="0"pageNumber="1">All the species associated with bryophytes are distinguished by color of antenna, color of maxillary palpus, color pattern of scutum and morphology of male genitalia. Colors of 1st flagellomere of antenna and maxillary palpus are dark or yellow, and unique to each species. Scutum was yellow or dark, and the yellow scutum often had three pairs of dark longitudinal stripes: medial stripes on anterior 2/3 (always merged together except in one species), intra-alar stripes (inner lateral stripes) and supra-alar stripes (outer lateral stripes) on anterior 4/5. In some species, the intra-alar stripe and the supra-alar stripe are merged together to shape a wide band, and often merged with the medial stripe. The color pattern of scutum is unique to each species at least among the species sharing the same host bryophyte genus. Although some species are very similar in external morphology, they could be clearly distinguished by the number of tubercle-like setae in a comb on the male epandrium and other male genital morphological characters.</paragraph>
group) based on presumed synapomorphy of the following characteristics: distribution of acrostichal setulae on scutum, morphology of epandrium and distiphallus of male genitalia, and color of legs. By assessing the morphological characteristics of the
<tableCitationid="CB3FB577CBE765543DD6E4579BDCD387"captionStart="Table 3"captionStartId="T3"captionText="Table 3. A synopsis of morphological characteristics of Japanese bryophyte-associated Phytoliriomyza species."httpUri="http://table.plazi.org/id/4D0D409101DD66831F665248707BEDB5"pageId="0"pageNumber="1"tableUuid="4D0D409101DD66831F665248707BEDB5">3</tableCitation>
<tableNoteid="16460583E8BD4F51DC4B3D9B53E2D400"pageId="0"pageNumber="1">1 1, present; 0, absent; 1-1, stripes unfused; 1/1, stripes fused.<br/> 2 Number of unfused tubercle-like setae forming a row is shown in parenthesis; -, lacking tubercle-like setae forming a row.<br/> * Number of setulae in four rows.</tableNote>
<figureCitationid="6C55B38D3CFBFD2DB8A863D315A273E1"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74D</figureCitation>
<figureCitationid="8B19662AE9A4E06F1FFF0C02BA09CE26"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74I</figureCitation>
<figureCitationid="7305B5B1C19DD425B5D714D1685A7AD5"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74G</figureCitation>
<figureCitationid="D244FF12EE905C8C3D5DEDF9E42B8791"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="24DCD12C33C75B82073E06A2B3B3DA7A"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1J</figureCitation>
<figureCitationid="ABA454585202A27B090A7682E003B3BB"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1H</figureCitation>
<figureCitationid="9543691EFA2D43714CB242645A084CC8"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74I</figureCitation>
<figureCitationid="35F82AFFE70DC4849860FFC7078278E3"captionStart="Figure 63"captionStartId="F63"captionText="Figure 63. Phytoliriomyza ricciae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female, frontal (MK-AG- 201) G-K male genitalia (G-J on Riccia nipponica K on R. huebeneriana) G whole genitalia, ventral H phallic complex, lateral I phallic complex, ventral L ejaculatory apodeme, lateral M epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure63"httpUri="https://binary.pensoft.net/fig/775437"pageId="0"pageNumber="1">63K</figureCitation>
<figureCitationid="328E1AEC406F86072D81C23B4A7A3626"captionStart="Figure 72"captionStartId="F72"captionText="Figure 72. Phytoliriomyza megacerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 14), dorsal G-J male genitalia (G at type locality H, I at Tashiro J at Yasukawa-keikoku) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral J phallic complex, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure72"httpUri="https://binary.pensoft.net/fig/775446"pageId="0"pageNumber="1">72A</figureCitation>
<figureCitationid="A76DA9ECC9220B7894FD6F22AF332387"captionStart="Figure 72"captionStartId="F72"captionText="Figure 72. Phytoliriomyza megacerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 14), dorsal G-J male genitalia (G at type locality H, I at Tashiro J at Yasukawa-keikoku) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral J phallic complex, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure72"httpUri="https://binary.pensoft.net/fig/775446"pageId="0"pageNumber="1">72H</figureCitation>
<figureCitationid="B5C8176F8662387056145EE7546F8FF4"captionStart="Figure 72"captionStartId="F72"captionText="Figure 72. Phytoliriomyza megacerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 14), dorsal G-J male genitalia (G at type locality H, I at Tashiro J at Yasukawa-keikoku) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral J phallic complex, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure72"httpUri="https://binary.pensoft.net/fig/775446"pageId="0"pageNumber="1">72G</figureCitation>
<figureCitationid="F9A976EF7E0ACF141E3BC2A5CC3AEC5B"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74A</figureCitation>
<figureCitationid="F3101C67309E126BD975667FBF59C51C"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74I</figureCitation>
<figureCitationid="D205F6CF60A6B99A44EB6D3D8D81FE84"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74G</figureCitation>
<figureCitationid="AA6AA03742B74D9F6C80CEDF99AEFFA0"captionStart="Figure 70"captionStartId="F70"captionText="Figure 70. Phytoliriomyza foliocerotis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 450) E dorsal F posterior G-J male genitalia G whole genitalia, ventral H ejaculatory apodeme, lateral I phallic complex, lateral J epandrium, posterior-ventral K, L female postabdomen K oviscape and spermatheca with spermatheca L tergite 10 M habitat at type locality N mined thallus of Folioceros fuciformis"figureDoi="10.3897/zookeys.1133.94530.figure70"httpUri="https://binary.pensoft.net/fig/775444"pageId="0"pageNumber="1">70B</figureCitation>
<figureCitationid="EA715CDCEAC5DED962BD42E4C4E9FE13"captionStart="Figure 70"captionStartId="F70"captionText="Figure 70. Phytoliriomyza foliocerotis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 450) E dorsal F posterior G-J male genitalia G whole genitalia, ventral H ejaculatory apodeme, lateral I phallic complex, lateral J epandrium, posterior-ventral K, L female postabdomen K oviscape and spermatheca with spermatheca L tergite 10 M habitat at type locality N mined thallus of Folioceros fuciformis"figureDoi="10.3897/zookeys.1133.94530.figure70"httpUri="https://binary.pensoft.net/fig/775444"pageId="0"pageNumber="1">70D</figureCitation>
<figureCitationid="FA8743C1098DB6C54785FFD7F94E95C4"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74B</figureCitation>
<figureCitationid="CC25A1C48902FD51C891B8B1BD299663"captionStart="Figure 74"captionStartId="F74"captionText="Figure 74. Phytoliriomyza phaeocerotis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 10), dorsal G-K male genitalia (G, H emerged from Notothylas temperata at type locality I emerged from Phaeoceros carolinianus at Kakeroma Is. J emerged from Notothylas temperata at Kibi-kogen K emerged from Anthoceros punctatus at Inago) G whole genitalia, ventral H, J phallic complex, lateral and ventral I epandrium, ventral K ejaculatory apodeme, lateral L phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure74"httpUri="https://binary.pensoft.net/fig/775448"pageId="0"pageNumber="1">74D</figureCitation>
<taxonomicNameid="2204931EB105FFA807A0ACC21F41A69A"authorityName="Sullivant ex A.Gray"authorityYear="1846"class="Anthocerotopsida"family="Notothyladaceae"genus="Notothylas"kingdom="Plantae"lsidName="Notothylas"order="Notothyladales"pageId="0"pageNumber="1"phylum="Anthocerotophyta"rank="genus">
<figureCitationid="501B3B4B39FB1AABF1217AAD35BBB223"captionStart="Figure 63"captionStartId="F63"captionText="Figure 63. Phytoliriomyza ricciae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female, frontal (MK-AG- 201) G-K male genitalia (G-J on Riccia nipponica K on R. huebeneriana) G whole genitalia, ventral H phallic complex, lateral I phallic complex, ventral L ejaculatory apodeme, lateral M epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure63"httpUri="https://binary.pensoft.net/fig/775437"pageId="0"pageNumber="1">63C</figureCitation>
<figureCitationid="EF28DF974EDD8FD8F65BA235E9033E82"captionStart="Figure 63"captionStartId="F63"captionText="Figure 63. Phytoliriomyza ricciae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female, frontal (MK-AG- 201) G-K male genitalia (G-J on Riccia nipponica K on R. huebeneriana) G whole genitalia, ventral H phallic complex, lateral I phallic complex, ventral L ejaculatory apodeme, lateral M epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure63"httpUri="https://binary.pensoft.net/fig/775437"pageId="0"pageNumber="1">63A</figureCitation>
<figureCitationid="7016588E7738DF22870B89D5180F1A87"captionStart="Figure 63"captionStartId="F63"captionText="Figure 63. Phytoliriomyza ricciae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female, frontal (MK-AG- 201) G-K male genitalia (G-J on Riccia nipponica K on R. huebeneriana) G whole genitalia, ventral H phallic complex, lateral I phallic complex, ventral L ejaculatory apodeme, lateral M epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure63"httpUri="https://binary.pensoft.net/fig/775437"pageId="0"pageNumber="1">63D</figureCitation>
<figureCitationid="7E62BA6463B94FA811EFE8CE09F04B96"captionStart="Figure 68"captionStartId="F68"captionText="Figure 68. Phytoliriomyza caerulescens sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 558) F frontal G habitus H lateral I dorsal J posterior K female fly ovipositing to thallus of Riccia billardieri L-O male genitalia L whole genitalia, ventral M phallic complex, lateral N epandrium, ventral O ejaculatory apodeme, ventral."figureDoi="10.3897/zookeys.1133.94530.figure68"httpUri="https://binary.pensoft.net/fig/775442"pageId="0"pageNumber="1">68F</figureCitation>
<figureCitationid="ACE4E1636E4FE673086427DADA34119B"captionStart="Figure 68"captionStartId="F68"captionText="Figure 68. Phytoliriomyza caerulescens sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 558) F frontal G habitus H lateral I dorsal J posterior K female fly ovipositing to thallus of Riccia billardieri L-O male genitalia L whole genitalia, ventral M phallic complex, lateral N epandrium, ventral O ejaculatory apodeme, ventral."figureDoi="10.3897/zookeys.1133.94530.figure68"httpUri="https://binary.pensoft.net/fig/775442"pageId="0"pageNumber="1">68A</figureCitation>
<figureCitationid="716DCE073683AC1730C3570650B073B4"captionStart="Figure 66"captionStartId="F66"captionText="Figure 66. Phytoliriomyza sexfasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 573) F habitus F lateral G frontal H dorsal I posterior K-O male genitalia K whole genitalia, ventral L epandrium, ventral M phallic complex, lateral N genitalia, lateral O ejaculatory apodeme, lateral P live female fly walking on soil."figureDoi="10.3897/zookeys.1133.94530.figure66"httpUri="https://binary.pensoft.net/fig/775440"pageId="0"pageNumber="1">66I</figureCitation>
<figureCitationid="0513D8C0B105BD95B4D47EB76E247F70"captionStart="Figure 21"captionStartId="F21"captionText="Figure 21. Locality records of three Phytoliriomyza species associated with Asterella and Reboulia spp.: P. iriomotensis and P. cometiformis."figureDoi="10.3897/zookeys.1133.94530.figure21"httpUri="https://binary.pensoft.net/fig/775394"pageId="0"pageNumber="1">21D</figureCitation>
<figureCitationid="03DF9D2E5EC152F08EF053BBB4E81E63"captionStart="Figure 21"captionStartId="F21"captionText="Figure 21. Locality records of three Phytoliriomyza species associated with Asterella and Reboulia spp.: P. iriomotensis and P. cometiformis."figureDoi="10.3897/zookeys.1133.94530.figure21"httpUri="https://binary.pensoft.net/fig/775394"pageId="0"pageNumber="1">21D</figureCitation>
<figureCitationid="652EF42AF7B9FA6F12AAF478D488B94B"captionStart="Figure 66"captionStartId="F66"captionText="Figure 66. Phytoliriomyza sexfasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 573) F habitus F lateral G frontal H dorsal I posterior K-O male genitalia K whole genitalia, ventral L epandrium, ventral M phallic complex, lateral N genitalia, lateral O ejaculatory apodeme, lateral P live female fly walking on soil."figureDoi="10.3897/zookeys.1133.94530.figure66"httpUri="https://binary.pensoft.net/fig/775440"pageId="0"pageNumber="1">66I</figureCitation>
<figureCitationid="D3F42329CC9ED0AD7D2299EB03C3C888"captionStart="Figure 66"captionStartId="F66"captionText="Figure 66. Phytoliriomyza sexfasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 573) F habitus F lateral G frontal H dorsal I posterior K-O male genitalia K whole genitalia, ventral L epandrium, ventral M phallic complex, lateral N genitalia, lateral O ejaculatory apodeme, lateral P live female fly walking on soil."figureDoi="10.3897/zookeys.1133.94530.figure66"httpUri="https://binary.pensoft.net/fig/775440"pageId="0"pageNumber="1">66I</figureCitation>
<figureCitationid="4751C591DEBED8E621A262F704D427C3"captionStart="Figure 66"captionStartId="F66"captionText="Figure 66. Phytoliriomyza sexfasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 573) F habitus F lateral G frontal H dorsal I posterior K-O male genitalia K whole genitalia, ventral L epandrium, ventral M phallic complex, lateral N genitalia, lateral O ejaculatory apodeme, lateral P live female fly walking on soil."figureDoi="10.3897/zookeys.1133.94530.figure66"httpUri="https://binary.pensoft.net/fig/775440"pageId="0"pageNumber="1">66I</figureCitation>
<figureCitationid="13E83AA37BF9FE4863E2730E6BAC2F21"captionStart="Figure 66"captionStartId="F66"captionText="Figure 66. Phytoliriomyza sexfasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 573) F habitus F lateral G frontal H dorsal I posterior K-O male genitalia K whole genitalia, ventral L epandrium, ventral M phallic complex, lateral N genitalia, lateral O ejaculatory apodeme, lateral P live female fly walking on soil."figureDoi="10.3897/zookeys.1133.94530.figure66"httpUri="https://binary.pensoft.net/fig/775440"pageId="0"pageNumber="1">66L</figureCitation>
<figureCitationid="4718839D9A58B96C0DB0B68FA71394E3"captionStart="Figure 68"captionStartId="F68"captionText="Figure 68. Phytoliriomyza caerulescens sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 558) F frontal G habitus H lateral I dorsal J posterior K female fly ovipositing to thallus of Riccia billardieri L-O male genitalia L whole genitalia, ventral M phallic complex, lateral N epandrium, ventral O ejaculatory apodeme, ventral."figureDoi="10.3897/zookeys.1133.94530.figure68"httpUri="https://binary.pensoft.net/fig/775442"pageId="0"pageNumber="1">68I</figureCitation>
<figureCitationid="459B23FA0E5A74EBF043C8172995DC2A"captionStart="Figure 68"captionStartId="F68"captionText="Figure 68. Phytoliriomyza caerulescens sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F-J paratype female (MK-AG-a 558) F frontal G habitus H lateral I dorsal J posterior K female fly ovipositing to thallus of Riccia billardieri L-O male genitalia L whole genitalia, ventral M phallic complex, lateral N epandrium, ventral O ejaculatory apodeme, ventral."figureDoi="10.3897/zookeys.1133.94530.figure68"httpUri="https://binary.pensoft.net/fig/775442"pageId="0"pageNumber="1">68N</figureCitation>
<figureCitationid="D61A4CBB6DE5483BC1A07CC6D26F05EE"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56B</figureCitation>
<figureCitationid="50A0B18F44E1B2462355A25E7C31A6C4"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56 D</figureCitation>
<figureCitationid="387856EA25D055EB799EFF0923882E86"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1B</figureCitation>
<figureCitationid="F7A3D914A6784863F47018D1BBCBB8D4"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="B742F51381988950AB9C509BE88C7317"captionStart="Figure 44"captionStartId="F44"captionText="Figure 44. Phytoliriomyza ugetsu sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 554), frontal G-J male genitalia (G, H at type locality I Yaku Is. J Mikisato) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K a live female fly foraging on Conocephalum orientalis thallus at Tategasaki L, M female postabdomen L oviscape and spermatheca M tergite 10 N puparia O mined thallus of Conocephalum orientalis at type locality P habitat at Tategasaki."figureDoi="10.3897/zookeys.1133.94530.figure44"httpUri="https://binary.pensoft.net/fig/775418"pageId="0"pageNumber="1">44D, F</figureCitation>
<figureCitationid="468E77B33B93EF678CD5548649550104"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Phytoliriomyza dumortierae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG- 241) F-L male genitalia (F at type locality G-I at Ichiu J-L at Tashiro) F whole genitalia, ventral G whole genitalia, lateral H phallic complex, lateral I ejaculatory apodeme, lateral J phallic complex, ventral K-L epandrium, posterior and anterior."figureDoi="10.3897/zookeys.1133.94530.figure10"httpUri="https://binary.pensoft.net/fig/775383"pageId="0"pageNumber="1">10E</figureCitation>
<figureCitationid="CE526C236512AB95A8552B3CF431BD8E"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56B</figureCitation>
<figureCitationid="65FC8A488FE25D8C3EE90C2546DEA553"captionStart="Figure 8"captionStartId="F8"captionText="Figure 8. Phytoliriomyza nubatama sp. nov. A-D, G holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG- 323), E dorsal F frontal G-L male genitalia (G-K type locality L Kume Island) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme lateral K postgonite L phallic complex, ventral."figureDoi="10.3897/zookeys.1133.94530.figure8"httpUri="https://binary.pensoft.net/fig/775381"pageId="0"pageNumber="1">8C</figureCitation>
<figureCitationid="E94AD8EB6E6FFD33CFF9FCF55390644C"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Phytoliriomyza dumortierae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG- 241) F-L male genitalia (F at type locality G-I at Ichiu J-L at Tashiro) F whole genitalia, ventral G whole genitalia, lateral H phallic complex, lateral I ejaculatory apodeme, lateral J phallic complex, ventral K-L epandrium, posterior and anterior."figureDoi="10.3897/zookeys.1133.94530.figure10"httpUri="https://binary.pensoft.net/fig/775383"pageId="0"pageNumber="1">10B</figureCitation>
<figureCitationid="0FAE68EEB1276284826624A79A00DDD0"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Phytoliriomyza dumortierae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG- 241) F-L male genitalia (F at type locality G-I at Ichiu J-L at Tashiro) F whole genitalia, ventral G whole genitalia, lateral H phallic complex, lateral I ejaculatory apodeme, lateral J phallic complex, ventral K-L epandrium, posterior and anterior."figureDoi="10.3897/zookeys.1133.94530.figure10"httpUri="https://binary.pensoft.net/fig/775383"pageId="0"pageNumber="1">10B</figureCitation>
<figureCitationid="0AEE432F8FC2AF7C6AFDA8434175DA46"captionStart="Figure 10"captionStartId="F10"captionText="Figure 10. Phytoliriomyza dumortierae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG- 241) F-L male genitalia (F at type locality G-I at Ichiu J-L at Tashiro) F whole genitalia, ventral G whole genitalia, lateral H phallic complex, lateral I ejaculatory apodeme, lateral J phallic complex, ventral K-L epandrium, posterior and anterior."figureDoi="10.3897/zookeys.1133.94530.figure10"httpUri="https://binary.pensoft.net/fig/775383"pageId="0"pageNumber="1">10K</figureCitation>
<figureCitationid="945B6902E03E7BAA70CDA6C501E1EB4D"captionStart="Figure 35"captionStartId="F35"captionText="Figure 35. Phytoliriomyza wiesnerellae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG-a 342), posterior F-J male genitalia F whole genitalia, ventral G epandrium, lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure35"httpUri="https://binary.pensoft.net/fig/775408"pageId="0"pageNumber="1">35B</figureCitation>
<figureCitationid="37209D53B3E21F501506E4CB140C4970"captionStart="Figure 35"captionStartId="F35"captionText="Figure 35. Phytoliriomyza wiesnerellae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG-a 342), posterior F-J male genitalia F whole genitalia, ventral G epandrium, lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure35"httpUri="https://binary.pensoft.net/fig/775408"pageId="0"pageNumber="1">35I</figureCitation>
<figureCitationid="8F117F0F8A3B140B8E697B893C89B273"captionStart="Figure 62"captionStartId="F62"captionText="Figure 62. Phytoliriomyza suetsugui sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E-H paratype female (MK-AG-a 434) E dorsal F frontal G lateral H posterior I-L male genitalia I whole genitalia, ventral J phallic complex, lateral K epandrium, ventral L ejaculatory apodeme M, N female postabdomen M oviscape and spermatheca N tergite 10 O, P habitat at type locality Q mined thallus of Conocephalum orientalis."figureDoi="10.3897/zookeys.1133.94530.figure62"httpUri="https://binary.pensoft.net/fig/775436"pageId="0"pageNumber="1">62D</figureCitation>
<figureCitationid="B8363887D55E703780799F561ACFBA5C"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56B</figureCitation>
<figureCitationid="8043DC015DDE2475C1613E1BC80E841F"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Phytoliriomyza tsukuyomi sp. nov. A-E holotype male A habitus, B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442) G-J male genitalia G whole genitalia, ventral H phallus complex, lateral I epandrium J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Marchantia polymorpha."figureDoi="10.3897/zookeys.1133.94530.figure5"httpUri="https://binary.pensoft.net/fig/775378"pageId="0"pageNumber="1">5C</figureCitation>
<figureCitationid="859BB7B397DCBBA01E506B6EC41D29C3"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Phytoliriomyza tsukuyomi sp. nov. A-E holotype male A habitus, B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442) G-J male genitalia G whole genitalia, ventral H phallus complex, lateral I epandrium J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Marchantia polymorpha."figureDoi="10.3897/zookeys.1133.94530.figure5"httpUri="https://binary.pensoft.net/fig/775378"pageId="0"pageNumber="1">5D</figureCitation>
<figureCitationid="5BAA1602ECE9E43027431D165F5F8A53"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Phytoliriomyza tsukuyomi sp. nov. A-E holotype male A habitus, B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442) G-J male genitalia G whole genitalia, ventral H phallus complex, lateral I epandrium J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Marchantia polymorpha."figureDoi="10.3897/zookeys.1133.94530.figure5"httpUri="https://binary.pensoft.net/fig/775378"pageId="0"pageNumber="1">5I</figureCitation>
<figureCitationid="2A477FB7774589A950D34C3203CEA3D7"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56C</figureCitation>
<figureCitationid="CD736AE3A4D92484EA1E38A7FD394497"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56D</figureCitation>
<figureCitationid="22DA7C7BBBB44766ED488C0F048EA833"captionStart="Figure 58"captionStartId="F58"captionText="Figure 58. Phytoliriomyza lanternaria sp. nov. A-D holotype male A habitus B dorsal C frontal D dorsal E paratype female (MK-AG- 430), dorsal F-I male genitalia (F-H at type locality I Mt. Horoiwa) F whole genitalia, ventral G phallic complex, lateral H, I epandrium, ventral, J, K female postabdomen J oviscape and spermatheca K tergite 10 L habitat at type locality M, N mined thalli of Conocephalum orientalis. An arrow in N indicates an internal puparium O live female fly at Yakumo."figureDoi="10.3897/zookeys.1133.94530.figure58"httpUri="https://binary.pensoft.net/fig/775432"pageId="0"pageNumber="1">58D</figureCitation>
<figureCitationid="3B013255300A8997B9968C707EF61998"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Phytoliriomyza tsukuyomi sp. nov. A-E holotype male A habitus, B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442) G-J male genitalia G whole genitalia, ventral H phallus complex, lateral I epandrium J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Marchantia polymorpha."figureDoi="10.3897/zookeys.1133.94530.figure5"httpUri="https://binary.pensoft.net/fig/775378"pageId="0"pageNumber="1">5I</figureCitation>
<figureCitationid="F3B917C17A9A716DADA06B9AA5BB4B74"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56B</figureCitation>
<figureCitationid="CD8229D6B57DC3B386CFD73692E8D502"captionStart="Figure 56"captionStartId="F56"captionText="Figure 56. Phytoliriomyza alpicola A-E male at Ikawa-toge A habitus B dorsal C frontal D dorsal E posterior F live female fly at Mitsumine-Jinja G-K male genitalia (G-J at Ikawa-toge K at Samani) G whole genitalia H phallic complex I epandrium J, K ejaculatory apodeme, ventral and lateral."figureDoi="10.3897/zookeys.1133.94530.figure56"httpUri="https://binary.pensoft.net/fig/775430"pageId="0"pageNumber="1">56D</figureCitation>
<figureCitationid="D356C98560EAB00BA0E9EF539F3FBD22"captionStart="Figure 58"captionStartId="F58"captionText="Figure 58. Phytoliriomyza lanternaria sp. nov. A-D holotype male A habitus B dorsal C frontal D dorsal E paratype female (MK-AG- 430), dorsal F-I male genitalia (F-H at type locality I Mt. Horoiwa) F whole genitalia, ventral G phallic complex, lateral H, I epandrium, ventral, J, K female postabdomen J oviscape and spermatheca K tergite 10 L habitat at type locality M, N mined thalli of Conocephalum orientalis. An arrow in N indicates an internal puparium O live female fly at Yakumo."figureDoi="10.3897/zookeys.1133.94530.figure58"httpUri="https://binary.pensoft.net/fig/775432"pageId="0"pageNumber="1">58B</figureCitation>
<figureCitationid="D9050CA67EA62EA7147EE10B1C75C6BF"captionStart="Figure 58"captionStartId="F58"captionText="Figure 58. Phytoliriomyza lanternaria sp. nov. A-D holotype male A habitus B dorsal C frontal D dorsal E paratype female (MK-AG- 430), dorsal F-I male genitalia (F-H at type locality I Mt. Horoiwa) F whole genitalia, ventral G phallic complex, lateral H, I epandrium, ventral, J, K female postabdomen J oviscape and spermatheca K tergite 10 L habitat at type locality M, N mined thalli of Conocephalum orientalis. An arrow in N indicates an internal puparium O live female fly at Yakumo."figureDoi="10.3897/zookeys.1133.94530.figure58"httpUri="https://binary.pensoft.net/fig/775432"pageId="0"pageNumber="1">58D</figureCitation>
<figureCitationid="8253C9AC52EE04F79E7F2A9DD6F968CF"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure6"httpUri="https://binary.pensoft.net/fig/775379"pageId="0"pageNumber="1">6I</figureCitation>
<figureCitationid="E330CC2D6934DAAA2A8B371AD37FBACF"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure6"httpUri="https://binary.pensoft.net/fig/775379"pageId="0"pageNumber="1">6I</figureCitation>
<figureCitationid="94F9E96FFA19C946BC5A43DD1A004712"captionStart="Figure 59"captionStartId="F59"captionText="Figure 59. Phytoliriomyza conocephali sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 444), dorsal G-K male genitalia (G on Conocephalum orientalis H-K on C. japonicum) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, dorsal K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure59"httpUri="https://binary.pensoft.net/fig/775433"pageId="0"pageNumber="1">59K</figureCitation>
<figureCitationid="EAC4562CCCE0B4366787D922614A768D"captionStart="Figure 59"captionStartId="F59"captionText="Figure 59. Phytoliriomyza conocephali sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 444), dorsal G-K male genitalia (G on Conocephalum orientalis H-K on C. japonicum) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, dorsal K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure59"httpUri="https://binary.pensoft.net/fig/775433"pageId="0"pageNumber="1">59K</figureCitation>
<figureCitationid="1E73A4FCC633F5C58CE2BBBC85BAC42B"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure6"httpUri="https://binary.pensoft.net/fig/775379"pageId="0"pageNumber="1">6I</figureCitation>
<figureCitationid="C7B44A992DFD31B63AA2F862FC795F8F"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure6"httpUri="https://binary.pensoft.net/fig/775379"pageId="0"pageNumber="1">6G</figureCitation>
<figureCitationid="E1889A3F236F094EEA04B9F49C44C089"captionStart="Figure 33"captionStartId="F33"captionText="Figure 33. Phytoliriomyza rebouliae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 424) E posterior F lateral G live female fly H-K male genitalia H whole genitalia, ventral I phallic complex, lateral J epandrium, ventral K ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure33"httpUri="https://binary.pensoft.net/fig/775406"pageId="0"pageNumber="1">33J</figureCitation>
<figureCitationid="EFFCE2A30A83B4F2727A116B75753166"captionStart="Figure 33"captionStartId="F33"captionText="Figure 33. Phytoliriomyza rebouliae sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 424) E posterior F lateral G live female fly H-K male genitalia H whole genitalia, ventral I phallic complex, lateral J epandrium, ventral K ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure33"httpUri="https://binary.pensoft.net/fig/775406"pageId="0"pageNumber="1">33H</figureCitation>
<figureCitationid="3B09692A72468E54A6C87D065B2257C6"captionStart="Figure 59"captionStartId="F59"captionText="Figure 59. Phytoliriomyza conocephali sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 444), dorsal G-K male genitalia (G on Conocephalum orientalis H-K on C. japonicum) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, dorsal K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure59"httpUri="https://binary.pensoft.net/fig/775433"pageId="0"pageNumber="1">59I</figureCitation>
<figureCitationid="19657B0842E15DB9647BBF3624490263"captionStart="Figure 59"captionStartId="F59"captionText="Figure 59. Phytoliriomyza conocephali sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 444), dorsal G-K male genitalia (G on Conocephalum orientalis H-K on C. japonicum) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, dorsal K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure59"httpUri="https://binary.pensoft.net/fig/775433"pageId="0"pageNumber="1">59K</figureCitation>
<figureCitationid="00A9CDC0821A1740D47C5291D5500087"captionStart="Figure 58"captionStartId="F58"captionText="Figure 58. Phytoliriomyza lanternaria sp. nov. A-D holotype male A habitus B dorsal C frontal D dorsal E paratype female (MK-AG- 430), dorsal F-I male genitalia (F-H at type locality I Mt. Horoiwa) F whole genitalia, ventral G phallic complex, lateral H, I epandrium, ventral, J, K female postabdomen J oviscape and spermatheca K tergite 10 L habitat at type locality M, N mined thalli of Conocephalum orientalis. An arrow in N indicates an internal puparium O live female fly at Yakumo."figureDoi="10.3897/zookeys.1133.94530.figure58"httpUri="https://binary.pensoft.net/fig/775432"pageId="0"pageNumber="1">58I</figureCitation>
<figureCitationid="93DF26D9E62F6B0EF38D966158320804"captionStart="Figure 58"captionStartId="F58"captionText="Figure 58. Phytoliriomyza lanternaria sp. nov. A-D holotype male A habitus B dorsal C frontal D dorsal E paratype female (MK-AG- 430), dorsal F-I male genitalia (F-H at type locality I Mt. Horoiwa) F whole genitalia, ventral G phallic complex, lateral H, I epandrium, ventral, J, K female postabdomen J oviscape and spermatheca K tergite 10 L habitat at type locality M, N mined thalli of Conocephalum orientalis. An arrow in N indicates an internal puparium O live female fly at Yakumo."figureDoi="10.3897/zookeys.1133.94530.figure58"httpUri="https://binary.pensoft.net/fig/775432"pageId="0"pageNumber="1">58I</figureCitation>
<figureCitationid="B38C233C357B270FF0A07D9FA41B5B5C"captionStart="Figure 44"captionStartId="F44"captionText="Figure 44. Phytoliriomyza ugetsu sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 554), frontal G-J male genitalia (G, H at type locality I Yaku Is. J Mikisato) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K a live female fly foraging on Conocephalum orientalis thallus at Tategasaki L, M female postabdomen L oviscape and spermatheca M tergite 10 N puparia O mined thallus of Conocephalum orientalis at type locality P habitat at Tategasaki."figureDoi="10.3897/zookeys.1133.94530.figure44"httpUri="https://binary.pensoft.net/fig/775418"pageId="0"pageNumber="1">44D</figureCitation>
<figureCitationid="3C03E50F1F1158D0763440E168308467"captionStart="Figure 44"captionStartId="F44"captionText="Figure 44. Phytoliriomyza ugetsu sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 554), frontal G-J male genitalia (G, H at type locality I Yaku Is. J Mikisato) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K a live female fly foraging on Conocephalum orientalis thallus at Tategasaki L, M female postabdomen L oviscape and spermatheca M tergite 10 N puparia O mined thallus of Conocephalum orientalis at type locality P habitat at Tategasaki."figureDoi="10.3897/zookeys.1133.94530.figure44"httpUri="https://binary.pensoft.net/fig/775418"pageId="0"pageNumber="1">44F</figureCitation>
<figureCitationid="95A4F98D25747375E9FB780E3FF2A705"captionStart="Figure 44"captionStartId="F44"captionText="Figure 44. Phytoliriomyza ugetsu sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 554), frontal G-J male genitalia (G, H at type locality I Yaku Is. J Mikisato) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K a live female fly foraging on Conocephalum orientalis thallus at Tategasaki L, M female postabdomen L oviscape and spermatheca M tergite 10 N puparia O mined thallus of Conocephalum orientalis at type locality P habitat at Tategasaki."figureDoi="10.3897/zookeys.1133.94530.figure44"httpUri="https://binary.pensoft.net/fig/775418"pageId="0"pageNumber="1">44D</figureCitation>
<figureCitationid="65D157F81EA6DC2D64B3A3583ABF37E1"captionStart="Figure 44"captionStartId="F44"captionText="Figure 44. Phytoliriomyza ugetsu sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 554), frontal G-J male genitalia (G, H at type locality I Yaku Is. J Mikisato) G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K a live female fly foraging on Conocephalum orientalis thallus at Tategasaki L, M female postabdomen L oviscape and spermatheca M tergite 10 N puparia O mined thallus of Conocephalum orientalis at type locality P habitat at Tategasaki."figureDoi="10.3897/zookeys.1133.94530.figure44"httpUri="https://binary.pensoft.net/fig/775418"pageId="0"pageNumber="1">44F</figureCitation>
<figureCitationid="BC86F1CBD5C450A0CB4E687B81D948BE"captionStart="Figure 49"captionStartId="F49"captionText="Figure 49. Phytoliriomyza pallidofasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 520), dorsal G-J male genitalia G whole genitalia H ejaculatory apodeme, lateral I phallic complex, lateral J epandrium, ventral K live fly L, M female postabdomen L oviscape and spermatheca M tergite 10 N-P mined thalli of Conocephalum orientalis. An arrow in O indicates an internal puparium."figureDoi="10.3897/zookeys.1133.94530.figure49"httpUri="https://binary.pensoft.net/fig/775423"pageId="0"pageNumber="1">49D</figureCitation>
<figureCitationid="FF2DBE6F9139053761348AF7BA7818AF"captionStart="Figure 49"captionStartId="F49"captionText="Figure 49. Phytoliriomyza pallidofasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 520), dorsal G-J male genitalia G whole genitalia H ejaculatory apodeme, lateral I phallic complex, lateral J epandrium, ventral K live fly L, M female postabdomen L oviscape and spermatheca M tergite 10 N-P mined thalli of Conocephalum orientalis. An arrow in O indicates an internal puparium."figureDoi="10.3897/zookeys.1133.94530.figure49"httpUri="https://binary.pensoft.net/fig/775423"pageId="0"pageNumber="1">49D</figureCitation>
<figureCitationid="4746A0BB58208903633E818EEEC1998C"captionStart="Figure 51"captionStartId="F51"captionText="Figure 51. Phytoliriomyza luteola sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 1), dorsal G-K male genitalia (G at type locality K-M at Mt. Horoiwa K at Sounkyo) G, H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral L, M female postabdomen L oviscape and spermatheca M tergite 10 N habitat at type locality O mined thallus of Conocephalum salebrosum P live female fly at Eniwa."figureDoi="10.3897/zookeys.1133.94530.figure51"httpUri="https://binary.pensoft.net/fig/775425"pageId="0"pageNumber="1">51B</figureCitation>
<figureCitationid="059156F87792AF52034C931196779C4F"captionStart="Figure 51"captionStartId="F51"captionText="Figure 51. Phytoliriomyza luteola sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 1), dorsal G-K male genitalia (G at type locality K-M at Mt. Horoiwa K at Sounkyo) G, H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral L, M female postabdomen L oviscape and spermatheca M tergite 10 N habitat at type locality O mined thallus of Conocephalum salebrosum P live female fly at Eniwa."figureDoi="10.3897/zookeys.1133.94530.figure51"httpUri="https://binary.pensoft.net/fig/775425"pageId="0"pageNumber="1">51I</figureCitation>
<figureCitationid="60F19DBF75C908A7E688273154932692"captionStart="Figure 52"captionStartId="F52"captionText="Figure 52. Phytoliriomyza helva sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 541), frontal G-J male genitalia G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure52"httpUri="https://binary.pensoft.net/fig/775426"pageId="0"pageNumber="1">52B</figureCitation>
<figureCitationid="6E1DD1C60F9E07C5E279CABF8211FAE9"captionStart="Figure 52"captionStartId="F52"captionText="Figure 52. Phytoliriomyza helva sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 541), frontal G-J male genitalia G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure52"httpUri="https://binary.pensoft.net/fig/775426"pageId="0"pageNumber="1">52I</figureCitation>
<figureCitationid="487A946CDDBFC0A770805AB6FA4DE2AF"captionStart="Figure 49"captionStartId="F49"captionText="Figure 49. Phytoliriomyza pallidofasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 520), dorsal G-J male genitalia G whole genitalia H ejaculatory apodeme, lateral I phallic complex, lateral J epandrium, ventral K live fly L, M female postabdomen L oviscape and spermatheca M tergite 10 N-P mined thalli of Conocephalum orientalis. An arrow in O indicates an internal puparium."figureDoi="10.3897/zookeys.1133.94530.figure49"httpUri="https://binary.pensoft.net/fig/775423"pageId="0"pageNumber="1">49D</figureCitation>
<figureCitationid="5F511E26EFA96CFEC8BFE310DB60578D"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="D1F817886B0914C5ADCF85A1F1CF2D53"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="39F12DD2366EFF83BA167C2BED08E071"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Phytoliriomyza igniculus sp. nov. A-E holotype male A habitus B lateral body C frontal head and ventral body D dorsal thorax E thorax in posterior view F-H male genitalia F whole genitalia G phallic complex H postgonite I epandrium J a living female fly on a thallus of Marchantia polymorpha K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure3"httpUri="https://binary.pensoft.net/fig/775376"pageId="0"pageNumber="1">3D</figureCitation>
<figureCitationid="5B7AE2582F90C0B227B1B54490FBDB63"captionStart="Figure 46"captionStartId="F46"captionText="Figure 46. Phytoliriomyza nigroflava sp. nov. A-C holotype male A habitus B dorsal C frontal D-G paratype female (MK-AG-a 22) D habitus E dorsal F frontal G posterior H-K male genitalia H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral B postgonite."figureDoi="10.3897/zookeys.1133.94530.figure46"httpUri="https://binary.pensoft.net/fig/775420"pageId="0"pageNumber="1">46B</figureCitation>
<figureCitationid="640CD3A08C1BCB3CCE8FCBEAA70A8384"captionStart="Figure 46"captionStartId="F46"captionText="Figure 46. Phytoliriomyza nigroflava sp. nov. A-C holotype male A habitus B dorsal C frontal D-G paratype female (MK-AG-a 22) D habitus E dorsal F frontal G posterior H-K male genitalia H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral B postgonite."figureDoi="10.3897/zookeys.1133.94530.figure46"httpUri="https://binary.pensoft.net/fig/775420"pageId="0"pageNumber="1">46B</figureCitation>
<figureCitationid="6651F4C80B9F431A9AF02930F22145C9"captionStart="Figure 54"captionStartId="F54"captionText="Figure 54. Phytoliriomyza bifasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 272), dorsal G-J male genitalia G whole genitalia H phallic complex I epandrium J ejaculatory apodeme, dorsal K, L female postabdomen K oviscape and spermatheca L tergite 10 M live female fly at Hozaka N habitat at Hachimori O-Q mined thalli (O Conocephalum orientalis at Hachimori P C. salebrosum at Haccho-toge Q C. salebrosum at Akka)."figureDoi="10.3897/zookeys.1133.94530.figure54"httpUri="https://binary.pensoft.net/fig/775428"pageId="0"pageNumber="1">54D</figureCitation>
<figureCitationid="4A98C7694B41FECF82B123ADAC7466EE"captionStart="Figure 54"captionStartId="F54"captionText="Figure 54. Phytoliriomyza bifasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 272), dorsal G-J male genitalia G whole genitalia H phallic complex I epandrium J ejaculatory apodeme, dorsal K, L female postabdomen K oviscape and spermatheca L tergite 10 M live female fly at Hozaka N habitat at Hachimori O-Q mined thalli (O Conocephalum orientalis at Hachimori P C. salebrosum at Haccho-toge Q C. salebrosum at Akka)."figureDoi="10.3897/zookeys.1133.94530.figure54"httpUri="https://binary.pensoft.net/fig/775428"pageId="0"pageNumber="1">54D</figureCitation>
<figureCitationid="48527AB3B0FBE2421DAF793662CE2B3F"captionStart="Figure 46"captionStartId="F46"captionText="Figure 46. Phytoliriomyza nigroflava sp. nov. A-C holotype male A habitus B dorsal C frontal D-G paratype female (MK-AG-a 22) D habitus E dorsal F frontal G posterior H-K male genitalia H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral B postgonite."figureDoi="10.3897/zookeys.1133.94530.figure46"httpUri="https://binary.pensoft.net/fig/775420"pageId="0"pageNumber="1">46B</figureCitation>
<figureCitationid="A7C031136BA7035A2FC2CCE0B783612C"captionStart="Figure 46"captionStartId="F46"captionText="Figure 46. Phytoliriomyza nigroflava sp. nov. A-C holotype male A habitus B dorsal C frontal D-G paratype female (MK-AG-a 22) D habitus E dorsal F frontal G posterior H-K male genitalia H whole genitalia, ventral I epandrium, ventral J phallic complex, lateral K ejaculatory apodeme, lateral B postgonite."figureDoi="10.3897/zookeys.1133.94530.figure46"httpUri="https://binary.pensoft.net/fig/775420"pageId="0"pageNumber="1">46A, D</figureCitation>
<figureCitationid="9E837FBAA0790DEEF327E540B428755F"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="8E44A27492C492BB4C01EA98917481BD"captionStart="Figure 24"captionStartId="F24"captionText="Figure 24. Phytoliriomyza argentifasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 436) G-J male genitalia (G-J at type locality K at Kume Is.) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral, J phallic complex, ventral K phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure24"httpUri="https://binary.pensoft.net/fig/775397"pageId="0"pageNumber="1">24B</figureCitation>
<figureCitationid="E882445D98EAF17655230F6C882AA3D1"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1B</figureCitation>
<figureCitationid="F6D75D1869F2B20A649639C8689285F8"captionStart="Figure 24"captionStartId="F24"captionText="Figure 24. Phytoliriomyza argentifasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 436) G-J male genitalia (G-J at type locality K at Kume Is.) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral, J phallic complex, ventral K phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure24"httpUri="https://binary.pensoft.net/fig/775397"pageId="0"pageNumber="1">24D</figureCitation>
<figureCitationid="9FED30B982DA8B3158F89E394C71857F"captionStart="Figure 24"captionStartId="F24"captionText="Figure 24. Phytoliriomyza argentifasciata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 436) G-J male genitalia (G-J at type locality K at Kume Is.) G whole genitalia, ventral H epandrium, ventral I ejaculatory apodeme, lateral, J phallic complex, ventral K phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure24"httpUri="https://binary.pensoft.net/fig/775397"pageId="0"pageNumber="1">24H</figureCitation>
<figureCitationid="3ACB40B41C9C043246108CF20FDA5407"captionStart="Figure 27"captionStartId="F27"captionText="Figure 27. Phytoliriomyza longifurcae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F, G paratype female (MK-AG-a 497) F dorsal G lateral H-L male genitalia H epandrium, ventral I whole genitalia, ventral J, K phallic complex, ventral and lateral L ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure27"httpUri="https://binary.pensoft.net/fig/775400"pageId="0"pageNumber="1">27D</figureCitation>
<figureCitationid="CC359F8E3BC49B11255CC6E9F5C24C66"captionStart="Figure 27"captionStartId="F27"captionText="Figure 27. Phytoliriomyza longifurcae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F, G paratype female (MK-AG-a 497) F dorsal G lateral H-L male genitalia H epandrium, ventral I whole genitalia, ventral J, K phallic complex, ventral and lateral L ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure27"httpUri="https://binary.pensoft.net/fig/775400"pageId="0"pageNumber="1">27I</figureCitation>
<figureCitationid="38F6F24FA246916D49B930AC2CCE34D3"captionStart="Figure 27"captionStartId="F27"captionText="Figure 27. Phytoliriomyza longifurcae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F, G paratype female (MK-AG-a 497) F dorsal G lateral H-L male genitalia H epandrium, ventral I whole genitalia, ventral J, K phallic complex, ventral and lateral L ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure27"httpUri="https://binary.pensoft.net/fig/775400"pageId="0"pageNumber="1">27H</figureCitation>
<figureCitationid="C1179557F2A9CE93336192AE3BFF021E"captionStart="Figure 27"captionStartId="F27"captionText="Figure 27. Phytoliriomyza longifurcae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F, G paratype female (MK-AG-a 497) F dorsal G lateral H-L male genitalia H epandrium, ventral I whole genitalia, ventral J, K phallic complex, ventral and lateral L ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure27"httpUri="https://binary.pensoft.net/fig/775400"pageId="0"pageNumber="1">27K</figureCitation>
<figureCitationid="D38AAE11726A8B93BBA826DF32FA40E9"captionStart="Figure 18"captionStartId="F18"captionText="Figure 18. Phytoliriomyza calcicola sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 222) E dorsal F posterior G, H male genitalia G epandrium H phallic complex in lateral view I live female fly."figureDoi="10.3897/zookeys.1133.94530.figure18"httpUri="https://binary.pensoft.net/fig/775391"pageId="0"pageNumber="1">18G</figureCitation>
<figureCitationid="87CEBBA478C4EFC0EC5FC040C87E075E"captionStart="Figure 18"captionStartId="F18"captionText="Figure 18. Phytoliriomyza calcicola sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 222) E dorsal F posterior G, H male genitalia G epandrium H phallic complex in lateral view I live female fly."figureDoi="10.3897/zookeys.1133.94530.figure18"httpUri="https://binary.pensoft.net/fig/775391"pageId="0"pageNumber="1">18G</figureCitation>
<figureCitationid="6B72FE120F21891ABF92798FBCA4B162"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1D</figureCitation>
<figureCitationid="0B38D357B117D751DCEA8178CB0E50D9"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Phytoliriomyza dorsata A-E male at Yakumo A habitus B lateral body C frontal head D dorsal thorax E thorax in posterior view F, G female at Yakumo F dorsal thorax G lateral body H-J male genitalia H whole genitalia, ventral I phallic complex, ventral J epandrium, ventral K, L female postabdomen K oviscape and spermatheca L tergite 10 M-O mined thalli of Marchantia polymorpha M mined thallus dissected to show interior and puparium (arrow) N mined thallus with exit hole O the same thallus with the mine dissected to show a puparium (arrow)."figureDoi="10.3897/zookeys.1133.94530.figure1"httpUri="https://binary.pensoft.net/fig/775374"pageId="0"pageNumber="1">1J</figureCitation>
<figureCitationid="82DE8E612FB8AA0399E50073D7CA1CFD"captionStart="Figure 48"captionStartId="F48"captionText="Figure 48. Phytoliriomyza brunofasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 42), dorsal G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, dorsal L, M female postabdomen L oviscape and spermatheca M tergite 10 N habitat at Nishiyama-onsen O-R mined thalli of Conocephalum salebrosum. Arrows in Q and R indicate puparia."figureDoi="10.3897/zookeys.1133.94530.figure48"httpUri="https://binary.pensoft.net/fig/775422"pageId="0"pageNumber="1">48D</figureCitation>
<figureCitationid="59CB9DF9F63E0D61FC6A8E434E5F9D2F"captionStart="Figure 48"captionStartId="F48"captionText="Figure 48. Phytoliriomyza brunofasciata sp. nov. A-E holotype male A habitus B dorsal C frontal D dorsal E posterior F paratype female (MK-AG-a 42), dorsal G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, dorsal L, M female postabdomen L oviscape and spermatheca M tergite 10 N habitat at Nishiyama-onsen O-R mined thalli of Conocephalum salebrosum. Arrows in Q and R indicate puparia."figureDoi="10.3897/zookeys.1133.94530.figure48"httpUri="https://binary.pensoft.net/fig/775422"pageId="0"pageNumber="1">48I</figureCitation>
<figureCitationid="9A12154F47F1B57F3B5FEF301CDFC1D8"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37C</figureCitation>
<figureCitationid="2E9B90ABAFCD54E10B8A3A2C9F60BA2E"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37I</figureCitation>
<figureCitationid="C600F59345CE6BDCD0647B03DF1683A9"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14B</figureCitation>
<figureCitationid="4E680DAC92783AE99E39B18F41227DD5"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14K</figureCitation>
<figureCitationid="6939EFEED1410ED819ECC72AF368DF41"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
<figureCitationid="F6A80339702D7480FC83115ECD56DC33"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
<figureCitationid="AA81784B1CC1620BEBB6746C04534209"captionStart="Figure 47"captionStartId="F47"captionText="Figure 47. Female morphology and larval ecology of Phytoliriomyza nigroflava sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C, D habitat (C type locality D at Shirabiso-toge) E-H mined thalli of Conocephalum salebrosum at type locality. Arrows in G and H shows internal puparia."figureDoi="10.3897/zookeys.1133.94530.figure47"httpUri="https://binary.pensoft.net/fig/775421"pageId="0"pageNumber="1">47D</figureCitation>
<figureCitationid="B36F3EC14C4ACFC408F66991CB761A1B"captionStart="Figure 47"captionStartId="F47"captionText="Figure 47. Female morphology and larval ecology of Phytoliriomyza nigroflava sp. nov. A, B female postabdomen A oviscape and spermatheca B tergite 10 C, D habitat (C type locality D at Shirabiso-toge) E-H mined thalli of Conocephalum salebrosum at type locality. Arrows in G and H shows internal puparia."figureDoi="10.3897/zookeys.1133.94530.figure47"httpUri="https://binary.pensoft.net/fig/775421"pageId="0"pageNumber="1">47D</figureCitation>
<figureCitationid="5E23AF3EF8C4CA0F413DD04E7453D17A"captionStart="Figure 22"captionStartId="F22"captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure22"httpUri="https://binary.pensoft.net/fig/775395"pageId="0"pageNumber="1">22D</figureCitation>
<figureCitationid="314F9771B8009618579C5F2C2A40D773"captionStart="Figure 22"captionStartId="F22"captionText="Figure 22. Phytoliriomyza cometiformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female, dorsal (MK-AG-a 428) F-J male genitalia (F-H at type locality I, J at Sagiura) F whole genitalia, ventral G phallic complex, lateral H, J epandrium, ventral I ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure22"httpUri="https://binary.pensoft.net/fig/775395"pageId="0"pageNumber="1">22J</figureCitation>
<figureCitationid="EFB629B7F0118D6EF63A6EAC19215A8F"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37D</figureCitation>
<figureCitationid="D549AD1B0179D4A0969397FB4C2EB13F"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37I</figureCitation>
<figureCitationid="AC013B58BE13502C64791E184E004821"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37D</figureCitation>
<figureCitationid="D6F350C28001FF2B06C4CF437711E704"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37D</figureCitation>
<figureCitationid="C244DF8D8A5DC815E79F30038880C4B3"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37G</figureCitation>
<figureCitationid="BD37119E239A282F629C035EAE1DEC37"captionStart="Figure 43"captionStartId="F43"captionText="Figure 43. Phytoliriomyza caliginosa sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 224) G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Conocephalum orientalis at Tsurara-kannon."figureDoi="10.3897/zookeys.1133.94530.figure43"httpUri="https://binary.pensoft.net/fig/775417"pageId="0"pageNumber="1">43D</figureCitation>
<figureCitationid="FCB3719D3D5A13EFEC189A01E8FCA917"captionStart="Figure 43"captionStartId="F43"captionText="Figure 43. Phytoliriomyza caliginosa sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 224) G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Conocephalum orientalis at Tsurara-kannon."figureDoi="10.3897/zookeys.1133.94530.figure43"httpUri="https://binary.pensoft.net/fig/775417"pageId="0"pageNumber="1">43D</figureCitation>
<figureCitationid="2992A336AE88315762EF1931D0691DF0"captionStart="Figure 43"captionStartId="F43"captionText="Figure 43. Phytoliriomyza caliginosa sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 224) G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Conocephalum orientalis at Tsurara-kannon."figureDoi="10.3897/zookeys.1133.94530.figure43"httpUri="https://binary.pensoft.net/fig/775417"pageId="0"pageNumber="1">43G</figureCitation>
<figureCitationid="4F98763D7833F2BCF4E3D206F99311AE"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37D</figureCitation>
<figureCitationid="F3BAEE83CEA5B558B826F0C149B8DF47"captionStart="Figure 37"captionStartId="F37"captionText="Figure 37. Phytoliriomyza luna sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG- 512), dorsal G-K male genitalia G whole genitalia, ventral and lateral H phallic complex, lateral I epandrium, ventral J ejaculatory apodeme, lateral K postgonite L-N female postabdomen at Aizankei L oviscape and spermatheca M tergite 10 N spermatheca."figureDoi="10.3897/zookeys.1133.94530.figure37"httpUri="https://binary.pensoft.net/fig/775410"pageId="0"pageNumber="1">37I</figureCitation>
<figureCitationid="F71F2C61759E23E5FAAC500D403E62C7"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40.. Phytoliriomyza izayoi sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 262) E posterior F lateral G-K male genitalia (G-I type locality J, K Renge-onsen) G whole genitalia, ventral H phallic complex, ventral I ejaculatory apodeme, dorsal J epandrium, ventral K phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure40"httpUri="https://binary.pensoft.net/fig/775414"pageId="0"pageNumber="1">40D</figureCitation>
<figureCitationid="FBF1B9878FC54D08F2C8A5E2BEED0B67"captionStart="Figure 40"captionStartId="F40"captionText="Figure 40.. Phytoliriomyza izayoi sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E, F paratype female (MK-AG-a 262) E posterior F lateral G-K male genitalia (G-I type locality J, K Renge-onsen) G whole genitalia, ventral H phallic complex, ventral I ejaculatory apodeme, dorsal J epandrium, ventral K phallic complex, lateral."figureDoi="10.3897/zookeys.1133.94530.figure40"httpUri="https://binary.pensoft.net/fig/775414"pageId="0"pageNumber="1">40J</figureCitation>
<figureCitationid="F28DFDB7688CDB19939D186F65DE37FE"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Phytoliriomyza igniculus sp. nov. A-E holotype male A habitus B lateral body C frontal head and ventral body D dorsal thorax E thorax in posterior view F-H male genitalia F whole genitalia G phallic complex H postgonite I epandrium J a living female fly on a thallus of Marchantia polymorpha K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure3"httpUri="https://binary.pensoft.net/fig/775376"pageId="0"pageNumber="1">3E</figureCitation>
<figureCitationid="4C4A089657057C06F61B7D5804860600"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Phytoliriomyza igniculus sp. nov. A-E holotype male A habitus B lateral body C frontal head and ventral body D dorsal thorax E thorax in posterior view F-H male genitalia F whole genitalia G phallic complex H postgonite I epandrium J a living female fly on a thallus of Marchantia polymorpha K, L female postabdomen K oviscape and spermatheca L tergite 10."figureDoi="10.3897/zookeys.1133.94530.figure3"httpUri="https://binary.pensoft.net/fig/775376"pageId="0"pageNumber="1">3I</figureCitation>
<figureCitationid="A05A4A067F05486F1F658FA5E42465EE"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Phytoliriomyza chichibuensis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F live male fly G-K male genitalia G whole genitalia, ventral H, I phallic complex, ventral and lateral J epandrium, ventral K ejaculatory apodeme, lateral, L, M female postabdomen L oviscape and spermatheca M tergite 10 N mined thallus of Conocephalum sp. 2."figureDoi="10.3897/zookeys.1133.94530.figure42"httpUri="https://binary.pensoft.net/fig/775416"pageId="0"pageNumber="1">42B</figureCitation>
<figureCitationid="0E4AEC7C386E772BF603E9344AD17667"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Phytoliriomyza chichibuensis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F live male fly G-K male genitalia G whole genitalia, ventral H, I phallic complex, ventral and lateral J epandrium, ventral K ejaculatory apodeme, lateral, L, M female postabdomen L oviscape and spermatheca M tergite 10 N mined thallus of Conocephalum sp. 2."figureDoi="10.3897/zookeys.1133.94530.figure42"httpUri="https://binary.pensoft.net/fig/775416"pageId="0"pageNumber="1">42J</figureCitation>
<figureCitationid="947D95AD8FCDE9868C9256C626F1BB65"captionStart="Figure 42"captionStartId="F42"captionText="Figure 42. Phytoliriomyza chichibuensis sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F live male fly G-K male genitalia G whole genitalia, ventral H, I phallic complex, ventral and lateral J epandrium, ventral K ejaculatory apodeme, lateral, L, M female postabdomen L oviscape and spermatheca M tergite 10 N mined thallus of Conocephalum sp. 2."figureDoi="10.3897/zookeys.1133.94530.figure42"httpUri="https://binary.pensoft.net/fig/775416"pageId="0"pageNumber="1">42J</figureCitation>
<figureCitationid="EA2369B60F2F723AAD28C099FAFE9520"captionStart="Figure 43"captionStartId="F43"captionText="Figure 43. Phytoliriomyza caliginosa sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 224) G-J male genitalia G whole genitalia, ventral H phallic complex, ventral I epandrium, ventral J ejaculatory apodeme, lateral K, L female postabdomen K oviscape and spermatheca L tergite 10 M mined thallus of Conocephalum orientalis at Tsurara-kannon."figureDoi="10.3897/zookeys.1133.94530.figure43"httpUri="https://binary.pensoft.net/fig/775417"pageId="0"pageNumber="1">43I</figureCitation>
<figureCitationid="53F6B73D3295E52895779E1EE1731AFE"captionStart="Figure 32"captionStartId="F32"captionText="Figure 32. Phytoliriomyza aratriformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG-a 346) F-I male genitalia (K-H type locality I, J Okuchihibu) F, I phallic complex, ventral (distiphallus lost) G phallic complex, lateral H epandrium, ventral J postgonite K, L female postabdomen K oviscape and spermatheca (one spermatheca split into two) L tergite 10 M live female N habitat at Nakatsugawa O mined thallus of Reboulia hemisphaerica orientalis at Nakatsugawa."figureDoi="10.3897/zookeys.1133.94530.figure32"httpUri="https://binary.pensoft.net/fig/775405"pageId="0"pageNumber="1">32D</figureCitation>
<figureCitationid="90D8EAD9F0FFEF6DC19517D2EBDAA004"captionStart="Figure 32"captionStartId="F32"captionText="Figure 32. Phytoliriomyza aratriformis sp. nov. A-D holotype male A habitus B lateral C frontal D dorsal E paratype female (MK-AG-a 346) F-I male genitalia (K-H type locality I, J Okuchihibu) F, I phallic complex, ventral (distiphallus lost) G phallic complex, lateral H epandrium, ventral J postgonite K, L female postabdomen K oviscape and spermatheca (one spermatheca split into two) L tergite 10 M live female N habitat at Nakatsugawa O mined thallus of Reboulia hemisphaerica orientalis at Nakatsugawa."figureDoi="10.3897/zookeys.1133.94530.figure32"httpUri="https://binary.pensoft.net/fig/775405"pageId="0"pageNumber="1">32E</figureCitation>
<figureCitationid="7744550CDEB6DE2BBD7A2322D00CBD8E"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14D</figureCitation>
<figureCitationid="834994B73B672C076885394509D69C01"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14E</figureCitation>
<figureCitationid="0A6B8470FCC0D32F6825D54E5C9A1F98"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14K</figureCitation>
<figureCitationid="47DB243451B7860999D7CA1DF989B007"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Phytoliriomyza plagiochasmatos sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 523), dorsal G-J male genitalia G whole genitalia, ventral H phallic complex, lateral I, J epandrium, ventral and anterior K ejaculatory apodeme, lateral L live male fly."figureDoi="10.3897/zookeys.1133.94530.figure17"httpUri="https://binary.pensoft.net/fig/775390"pageId="0"pageNumber="1">17I</figureCitation>
<figureCitationid="B962BCF707A3B1C4E610521C3F90EB64"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Phytoliriomyza arcus sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 16), dorsal G-K male genitalia (G at type locality H-K at Todai-shiraiwa) G, H whole genitalia, ventral I phallic complex, lateral J ejaculatory apodeme, lateral K epandrium, ventral."figureDoi="10.3897/zookeys.1133.94530.figure14"httpUri="https://binary.pensoft.net/fig/775387"pageId="0"pageNumber="1">14K</figureCitation>
<figureCitationid="C5BA4F13399F7BF6AAB877CD028E2C06"captionStart="Figure 29"captionStartId="F29"captionText="Figure 29. Phytoliriomyza falcata sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 18) dorsal G-L male genitalia (G-K, at type locality L at Tatsuyama) G whole genitalia, ventral H phallic complex, ventral I, J, L epandrium (I anterior J anterior-ventral L ventral) K ejaculatory apodeme, lateral."figureDoi="10.3897/zookeys.1133.94530.figure29"httpUri="https://binary.pensoft.net/fig/775402"pageId="0"pageNumber="1">29I</figureCitation>
<tableCitationid="A48BCC28404C32396B09FFFD876CF847"captionStart="Table 1"captionStartId="T1"captionText="Table 1. Thalloid liverwort and hornwort genera, with number of species of each genus and phytophagous insect species associated with the genera in Japan."httpUri="http://table.plazi.org/id/B0659C7F15A64A3632814688D4A74986"pageId="0"pageNumber="1"tableUuid="B0659C7F15A64A3632814688D4A74986">1</tableCitation>
<tableCitationid="2C793FE7B4EF469032F010915E8B5714"captionStart="Table 2"captionStartId="T2"captionText="Table 2. Agromyzid species associated with thalloid liverworts and hornworts in Japan, with their host plants and distributions."httpUri="http://table.plazi.org/id/4D2040899F03849E19AA6350A5EF1917"pageId="0"pageNumber="1"tableUuid="4D2040899F03849E19AA6350A5EF1917">2</tableCitation>
