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<document id="7FE839FE2E0CD8D6E6219FAFA51B8C18" ID-DOI="10.11646/zootaxa.3709.3.2" ID-GBIF-Dataset="05206976-826d-4c9d-b858-bf8c11bcb9ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="222475" ID-ZooBank="1FC2BF4A-79A8-4C20-B95A-4B8CC0D45877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673675043" checkinUser="plazi" docAuthor="Lima, Lucas R. C., Mariano, Rodolfo &amp; Pinheiro, Ulisses" docDate="2013" docId="03F587C3FFC7B234FF46FD09C277FFA5" docLanguage="en" docName="zt03709p242.pdf" docOrigin="Zootaxa 3709 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thraulodes luizgonzagai Lima, Mariano &amp; Pinheiro, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="235" masterDocId="FFCCFFBBFFC5B232FFD1FFBBC653FF52" masterDocTitle="New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil" masterLastPageNumber="242" masterPageNumber="230" pageNumber="232" updateTime="1698348997805" updateUser="plazi">
<mods:mods id="92B36C272398B8E81EB8F159D4F460D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="666BD69950BCCDA32ED622562759AF5F">
<mods:title id="268C05DA962B4075F38A5CD18AC59375">New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil</mods:title>
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<mods:name id="DBF81B5350CB23B73EFAA3884F680679" type="personal">
<mods:role id="C292579E506D465F93E0104346D1F512">
<mods:roleTerm id="DC323B2D29D6D6F62757CAD6A0CE912C">Author</mods:roleTerm>
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<mods:namePart id="B31448842E8B5B36CC71EE98B63C2134">Lima, Lucas R. C.</mods:namePart>
</mods:name>
<mods:name id="9246D298632F12D0C506A782A48DF2CE" type="personal">
<mods:role id="16BA4922636C555B939F0B535746E48C">
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<mods:namePart id="5362E5490F3033E6C49BBF5DBDBEB576">Mariano, Rodolfo</mods:namePart>
</mods:name>
<mods:name id="DA5DAB910320BD4BCB6FC70AAA102294" type="personal">
<mods:role id="BD836FEEB5D6C8DC3152C74615D3950E">
<mods:roleTerm id="C66160FD69644CABBA8A041CBA3F5E6A">Author</mods:roleTerm>
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<mods:namePart id="1B10BE48E702A7C64419AFF34651A5E0">Pinheiro, Ulisses</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C503930E31BB27F4D56A59E464D5D29">text</mods:typeOfResource>
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<mods:title id="99B23B687067F6B1BFB58252E1D1B39B">Zootaxa</mods:title>
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<mods:part id="C16FDD5D172E8FC9979F69361B983948">
<mods:date id="0D6CECD6CC26F8DAB720D832FEEC9AE6">2013</mods:date>
<mods:detail id="0BA51A7B3838880EF08C5EB2E27A377E" type="volume">
<mods:number id="304417ECF10174B1B382B5C788D0FC91">3709</mods:number>
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<mods:detail id="85FAFDC20B018B6643C531B09C593B81" type="issue">
<mods:number id="6422A615DB80C5145048300FDD57D79D">3</mods:number>
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<mods:start id="07A0BCF5C273109D4E71C01AAD7E11A3">230</mods:start>
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<mods:classification id="758CAFF8F2B6C28060A060E8175E7090">journal article</mods:classification>
<mods:identifier id="1E0EB253E88CF29D50AEAEC12C3C2355" type="DOI">10.11646/zootaxa.3709.3.2</mods:identifier>
<mods:identifier id="27EDA91064202FB34EE223BA6783B1E9" type="GBIF-Dataset">05206976-826d-4c9d-b858-bf8c11bcb9ff</mods:identifier>
<mods:identifier id="EE295460F97AA712BE7969B182370386" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E7913DA598852578EB46BFDE22AD4CB3" type="Zenodo-Dep">222475</mods:identifier>
<mods:identifier id="8B8353C822A971DDDBE97DF9030C0D56" type="ZooBank">1FC2BF4A-79A8-4C20-B95A-4B8CC0D45877</mods:identifier>
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<treatment id="03F587C3FFC7B234FF46FD09C277FFA5" ID-DOI="http://doi.org/10.5281/zenodo.6156361" ID-GBIF-Taxon="119578472" ID-Zenodo-Dep="6156361" LSID="urn:lsid:plazi:treatment:03F587C3FFC7B234FF46FD09C277FFA5" httpUri="http://treatment.plazi.org/id/03F587C3FFC7B234FF46FD09C277FFA5" lastPageId="6" lastPageNumber="235" pageId="2" pageNumber="232">
<subSubSection id="C346655EFFC7B230FF46FD09C7E9FDBC" pageId="2" pageNumber="232" type="nomenclature">
<paragraph id="8BE336D5FFC7B230FF46FD09C537FD99" blockId="2.[151,868,690,750]" box="[151,868,690,715]" pageId="2" pageNumber="232">
<heading id="D0AB81B9FFC7B230FF46FD09C537FD99" bold="true" box="[151,868,690,715]" fontSize="10" level="2" pageId="2" pageNumber="232" reason="2">
<emphasis id="B928EAC7FFC7B230FF46FD09C537FD99" bold="true" box="[151,868,690,715]" pageId="2" pageNumber="232">
<taxonomicName id="4C5C4D56FFC7B230FF46FD09C7E7FD99" box="[151,436,690,715]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="232" phylum="Arthropoda" rank="species" species="luizgonzagai" status="sp. nov.">
<emphasis id="B928EAC7FFC7B230FF46FD09C7E7FD99" bold="true" box="[151,436,690,715]" italics="true" pageId="2" pageNumber="232">Thraulodes luizgonzagai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC7B230FE6AFD09C445FD99" box="[443,534,690,715]" pageId="2" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
, Lima, Mariano &amp; Pinheiro
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE336D5FFC7B230FF46FD6EC7E9FDBC" blockId="2.[151,868,690,750]" box="[151,442,725,750]" pageId="2" pageNumber="232">
(
<figureCitation id="13672A50FFC7B230FF4EFD6EC758FDBC" box="[159,267,725,750]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[286,1309,193,1928]" captionTargetId="figure@3.[286,1309,193,1930]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Wings of male imagos. 2, Thraulodes luizgonzagai sp. nov., forewing; 3, idem, hind wing; 4, T. sternimaculatus sp. nov., forewing; 5, idem, hind wing; 6, T. alapictus sp. nov., forewing; 7, idem, hind wing; 8, T. pinhoi sp, nov., forewing; 9, idem, hind wing." httpUri="https://zenodo.org/record/222477/files/figure.png" pageId="2" pageNumber="232">Figs. 2–3</figureCitation>
; 10–11; 18–22)
</paragraph>
</subSubSection>
<subSubSection id="C346655EFFC7B230FF46FCAEC5CEFCB0" pageId="2" pageNumber="232" type="diagnosis">
<paragraph id="8BE336D5FFC7B230FF46FCAEC5CEFCB0" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF46FCAEC742FC7C" bold="true" box="[151,273,789,814]" pageId="2" pageNumber="232">Diagnosis.</emphasis>
The male adult of this species can be separated from all the other congeners by the combination of the following characters: (1) costal membrane basal to bulla without cross veins (
<figureCitation id="13672A50FFC7B230FC27FC82C268FC00" box="[1014,1083,825,850]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[286,1309,193,1928]" captionTargetId="figure@3.[286,1309,193,1930]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Wings of male imagos. 2, Thraulodes luizgonzagai sp. nov., forewing; 3, idem, hind wing; 4, T. sternimaculatus sp. nov., forewing; 5, idem, hind wing; 6, T. alapictus sp. nov., forewing; 7, idem, hind wing; 8, T. pinhoi sp, nov., forewing; 9, idem, hind wing." httpUri="https://zenodo.org/record/222477/files/figure.png" pageId="2" pageNumber="232">Fig. 2</figureCitation>
); (2) costal and subcostal zones of forewing translucent (
<figureCitation id="13672A50FFC7B230FE10FCE6C45FFC24" box="[449,524,861,886]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[286,1309,193,1928]" captionTargetId="figure@3.[286,1309,193,1930]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Wings of male imagos. 2, Thraulodes luizgonzagai sp. nov., forewing; 3, idem, hind wing; 4, T. sternimaculatus sp. nov., forewing; 5, idem, hind wing; 6, T. alapictus sp. nov., forewing; 7, idem, hind wing; 8, T. pinhoi sp, nov., forewing; 9, idem, hind wing." httpUri="https://zenodo.org/record/222477/files/figure.png" pageId="2" pageNumber="232">Fig. 2</figureCitation>
); (3) two dark brown bands on femora (
<figureCitation id="13672A50FFC7B230FBD5FCE6C233FC24" box="[1028,1120,861,886]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Fig. 22</figureCitation>
); (4) terga in dorsal view predominantly light brown, in lateral view translucent whitish, with three distinct spots on terga II–VI and one spot on terga VII–VIII (
<figureCitation id="13672A50FFC7B230FEA5FC1EC7ADFCEC" box="[372,510,933,958]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Figs. 19–20</figureCitation>
; (5) subapical spines of penes long and narrow, apicolateral area not forming an “ear”, lateral pouch present, and without recurved folds (
<figureCitation id="13672A50FFC7B230FCC2FC72C5C3FCB0" box="[787,912,969,994]" captionStart="FIGURES 10 – 17" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[195,1406,193,1776]" captionTargetId="figure@4.[170,1412,193,1932]" captionTargetPageId="4" captionText="FIGURES 10 – 17. Genitalia of male imagos. 10, Thraulodes luizgonzagai sp. nov., genitalia (ventral view); 11, idem, penes (ventral view); 12, T. sternimaculatus sp. nov., genitalia (ventral view); 13, idem, penes (ventral view); 14, T. alapictus sp. nov., genitalia (ventral view); 15, idem, penes (ventral view); 16, T. pinhoi sp, nov. genitalia (ventral view); 17, idem, penes (ventral view)." httpUri="https://zenodo.org/record/222478/files/figure.png" pageId="2" pageNumber="232">Fig. 10–11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346655EFFC7B230FF16FC56C462F84C" pageId="2" pageNumber="232" type="description">
<paragraph id="8BE336D5FFC7B230FF16FC56C311FB54" blockId="2.[151,1437,789,2038]" box="[199,1346,1005,1030]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FC56C708FB54" bold="true" box="[199,347,1005,1030]" pageId="2" pageNumber="232">Male imago.</emphasis>
<emphasis id="B928EAC7FFC7B230FEB3FC56C7E3FB54" box="[354,432,1005,1030]" italics="true" pageId="2" pageNumber="232">Length</emphasis>
: body,
<quantity id="4CA49B30FFC7B230FDD3FC56C408FB57" box="[514,603,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="232" unit="mm" value="5.5">5.5 mm</quantity>
; forewing,
<quantity id="4CA49B30FFC7B230FD0BFC56C560FB57" box="[730,819,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="232" unit="mm" value="5.8">5.8 mm</quantity>
; hind wing,
<quantity id="4CA49B30FFC7B230FC6CFC55C245FB57" box="[957,1046,1006,1030]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="232" unit="mm" value="1.1">1.1 mm</quantity>
; general coloration brown.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16FBAAC52AFB1C" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FBAAC757FB78" box="[199,260,1041,1066]" italics="true" pageId="2" pageNumber="232">Head</emphasis>
(
<figureCitation id="13672A50FFC7B230FEC5FBAAC7CCFB78" box="[276,415,1041,1066]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Figs. 18–19</figureCitation>
): dark brown; antenna yellowish brown with scape lighter; upper portion of eye yellowish brown, lower portion black; ocelli white with a black ring at base.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16FBE2C2D8FBE8" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FBE2C74BFB20" box="[199,280,1113,1138]" italics="true" pageId="2" pageNumber="232">Thorax</emphasis>
(
<figureCitation id="13672A50FFC7B230FEFEFBE2C793FB20" box="[303,448,1113,1138]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Figs. 19–21</figureCitation>
): general color of pro-, meso and metanotum chestnut, with sutures dark brown. Mesonotum with whitish cup-shaped spot on mesoscutelum, posterior region with scuto scutelar suture whitish. Pleura whitish washed with red. Pro- and mesosternum whitish; furcasternum dark brown.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16FB7DC26FFA74" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FB7DC759FB8F" box="[199,266,1222,1245]" italics="true" pageId="2" pageNumber="232">Wings</emphasis>
(
<figureCitation id="13672A50FFC7B230FEF2FB7EC7CBFB8C" box="[291,408,1221,1246]" captionStart="FIGURES 2 – 9" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[286,1309,193,1928]" captionTargetId="figure@3.[286,1309,193,1930]" captionTargetPageId="3" captionText="FIGURES 2 – 9. Wings of male imagos. 2, Thraulodes luizgonzagai sp. nov., forewing; 3, idem, hind wing; 4, T. sternimaculatus sp. nov., forewing; 5, idem, hind wing; 6, T. alapictus sp. nov., forewing; 7, idem, hind wing; 8, T. pinhoi sp, nov., forewing; 9, idem, hind wing." httpUri="https://zenodo.org/record/222477/files/figure.png" pageId="2" pageNumber="232">Figs. 2–3</figureCitation>
): forewing membrane hyaline, with dark spot near base; costal and subcostal zones translucent; veins ICu2 and CuP not joined basally. Costal membrane basal to bulla without cross veins and 8 cross veins distal to bulla.
<collectingCountry id="F34B7645FFC7B230FEAEFAB6C7E4FA74" box="[383,439,1293,1318]" name="India" pageId="2" pageNumber="232">Hind</collectingCountry>
wing hyaline, with dark spot near base and 4 cross veins.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16FA89C52CFA3C" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FA89C6AFFA1B" box="[199,252,1330,1353]" italics="true" pageId="2" pageNumber="232">Legs</emphasis>
(
<figureCitation id="13672A50FFC7B230FEDBFA8AC70DFA18" box="[266,350,1329,1354]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Fig. 22</figureCitation>
): femur with two dark brown bands; apical 1/3 of femur tinged with light brown and inner margin washed with black. Tibiae and tarsi light brown. Tarsi pale yellow.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16FAC2C2E5FAAC" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16FAC2C762FAC0" box="[199,305,1401,1426]" italics="true" pageId="2" pageNumber="232">Abdomen</emphasis>
(
<figureCitation id="13672A50FFC7B230FE91FAC2C798FAC0" box="[320,459,1401,1426]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Figs. 18–20</figureCitation>
): terga predominantly light brown. In lateral view translucent brown, with two isolated spots and two stigmatic dots on terga III–VI, and one isolated spot and a blackish stripe on terga VII–IX. In dorsal view, terga segments I, VII–IX brown and II–VI translucent with posterior margin light brown. Terga segments I– III and VI–VII medially with blackish dorsal mark. Sterna I–VI translucent and VII–X brown.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16F9B2C78CF9E0" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16F9B2C762F970" box="[199,305,1545,1570]" italics="true" pageId="2" pageNumber="232">Genitalia</emphasis>
(
<figureCitation id="13672A50FFC7B230FE92F9B2C79CF970" box="[323,463,1545,1570]" captionStart="FIGURES 10 – 17" captionStartId="4.[151,264,1863,1886]" captionTargetBox="[195,1406,193,1776]" captionTargetId="figure@4.[170,1412,193,1932]" captionTargetPageId="4" captionText="FIGURES 10 – 17. Genitalia of male imagos. 10, Thraulodes luizgonzagai sp. nov., genitalia (ventral view); 11, idem, penes (ventral view); 12, T. sternimaculatus sp. nov., genitalia (ventral view); 13, idem, penes (ventral view); 14, T. alapictus sp. nov., genitalia (ventral view); 15, idem, penes (ventral view); 16, T. pinhoi sp, nov. genitalia (ventral view); 17, idem, penes (ventral view)." httpUri="https://zenodo.org/record/222478/files/figure.png" pageId="2" pageNumber="232">Figs. 10–11</figureCitation>
): forceps pale yellow, segment II washed with black. Posterior margin of subgenital plate almost straight, with a medial rounded projection. Penes pale yellow; 2/3 of penes divided apically, with a pair of long and narrow subapical spines, 2.0x the width of penes; apicolateral area not forming an “ear”; external margin forming lateral pouch and without recurved folds parallel to inner margin. Terminal filament whitish, basal area of segments black.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16F905C44FF9A8" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16F905C769F987" box="[199,314,1726,1749]" italics="true" pageId="2" pageNumber="232">Variations</emphasis>
: Some specimens show color patterns overall darker (
<figureCitation id="13672A50FFC7B230FC7DF906C5ACF984" box="[940,1023,1725,1750]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Fig. 18</figureCitation>
) or lighter (
<figureCitation id="13672A50FFC7B230FB5DF906C2B3F984" box="[1164,1248,1725,1750]" captionStart="FIGURES 18 – 22" captionStartId="5.[151,264,1791,1814]" captionTargetBox="[166,1436,193,1741]" captionTargetId="figure@5.[166,1451,193,1744]" captionTargetPageId="5" captionText="FIGURES 18 – 22. Thraulodes luizgonzagai sp. nov.: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1 mm)." httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="2" pageNumber="232">Fig. 19</figureCitation>
), due to alcohol fixation period making them paler.
</paragraph>
<paragraph id="8BE336D5FFC7B230FF16F8BEC462F84C" blockId="2.[151,1437,789,2038]" box="[199,561,1797,1822]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16F8BEC7E7F84C" bold="true" box="[199,436,1797,1822]" pageId="2" pageNumber="232">Female and nymph.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C346655EFFC7B230FF16F892C515F8A4" pageId="2" pageNumber="232" type="materials_examined">
<paragraph id="8BE336D5FFC7B230FF16F892C515F8A4" blockId="2.[151,1437,789,2038]" pageId="2" pageNumber="232">
<emphasis id="B928EAC7FFC7B230FF16F892C7AAF810" bold="true" box="[199,505,1833,1858]" pageId="2" pageNumber="232">
<typeStatus id="54E78877FFC7B230FF16F892C751F810" box="[199,258,1833,1858]" pageId="2" pageNumber="232">Type</typeStatus>
material.
<typeStatus id="54E78877FFC7B230FE52F892C7A7F810" box="[387,500,1833,1858]" pageId="2" pageNumber="232" type="holotype">Holotype</typeStatus>
:
</emphasis>
one male imago,
<collectingCountry id="F34B7645FFC7B230FD1EF892C545F810" box="[719,790,1833,1858]" name="Brazil" pageId="2" pageNumber="232">Brazil</collectingCountry>
, Bahia State, Juazeiro, Rio São Francisco,
<geoCoordinate id="EE685012FFC7B230FADEF892C3C4F810" box="[1295,1431,1833,1858]" direction="south" orientation="latitude" pageId="2" pageNumber="232" precision="1" value="-9.417861">9°25'04.3&quot;S</geoCoordinate>
,
<geoCoordinate id="EE685012FFC7B230FF46F8F6C765F834" box="[151,310,1869,1894]" direction="west" orientation="longitude" pageId="2" pageNumber="232" precision="1" value="-40.480804">40°28'50.9&quot;W</geoCoordinate>
,
<quantity id="4CA49B30FFC7B230FE97F8F6C7DBF834" box="[326,392,1869,1894]" metricMagnitude="2" metricUnit="m" metricValue="3.62" pageId="2" pageNumber="232" unit="m" value="362.0">362m</quantity>
,
<date id="FFE21015FFC7B230FE49F8F6C459F834" box="[408,522,1869,1894]" pageId="2" pageNumber="232" value="2011-10-05">05.x.2011</date>
, L.R.C. Lima coll. (
<collectionCode id="ED4DAE10FFC7B230FD2DF8F5C527F834" box="[764,884,1870,1894]" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="232">MZUESC</collectionCode>
).
<typeStatus id="54E78877FFC7B230FC5AF8F6C5ABF834" box="[907,1016,1869,1894]" pageId="2" pageNumber="232" type="paratype">
<emphasis id="B928EAC7FFC7B230FC5AF8F6C5ABF834" bold="true" box="[907,1016,1869,1894]" pageId="2" pageNumber="232">Paratype</emphasis>
</typeStatus>
: same data as
<typeStatus id="54E78877FFC7B230FB7CF8F6C341F834" box="[1197,1298,1869,1894]" pageId="2" pageNumber="232" type="holotype">holotype</typeStatus>
, three male imagos (1
<collectionCode id="ED4DAE10FFC7B230FEDCF8C9C70BF8D8" box="[269,344,1906,1930]" pageId="2" pageNumber="232">CZNC</collectionCode>
and 2
<collectionCode id="ED4DAE10FFC7B230FE75F8C9C448F8D8" box="[420,539,1906,1930]" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="232">MZUESC</collectionCode>
); one male,
<collectingCountry id="F34B7645FFC7B230FD72F8CAC4B9F8D8" box="[675,746,1905,1930]" name="Brazil" pageId="2" pageNumber="232">Brazil</collectingCountry>
, Pernambuco State, Petrolina, Base Fluvial Ilha do Fogo, Rio São Francisco,
<geoCoordinate id="EE685012FFC7B230FE92F82DC79FF8FC" box="[323,460,1942,1966]" direction="south" orientation="latitude" pageId="2" pageNumber="232" precision="1" value="-9.406361">9°24'22.9&quot;S</geoCoordinate>
,
<geoCoordinate id="EE685012FFC7B230FE09F82DC424F8FC" box="[472,631,1942,1966]" direction="west" orientation="longitude" pageId="2" pageNumber="232" precision="1" value="-40.504833">40°30'17.4&quot;W</geoCoordinate>
,
<quantity id="4CA49B30FFC7B230FD52F82EC496F8FC" box="[643,709,1941,1966]" metricMagnitude="2" metricUnit="m" metricValue="3.62" pageId="2" pageNumber="232" unit="m" value="362.0">362m</quantity>
,
<date id="FFE21015FFC7B230FD00F82DC510F8FC" box="[721,835,1942,1966]" pageId="2" pageNumber="232" value="2011-10-09">09.x.2011</date>
, L.R.C. Lima coll. (
<collectionCode id="ED4DAE10FFC7B230FBF5F82DC23CF8FC" box="[1060,1135,1942,1966]" pageId="2" pageNumber="232">UFPE</collectionCode>
); two male imagos,
<collectingCountry id="F34B7645FFC7B230FA81F82EC3C4F8FC" box="[1360,1431,1941,1966]" name="Brazil" pageId="2" pageNumber="232">Brazil</collectingCountry>
, Pernambuco, Petrolina, Rio da Vitória, afluente do Rio São Francisco, 09°21'814''S, 40°35'409''W,
<quantity id="4CA49B30FFC7B230FA97F801C3C4F883" box="[1350,1431,1978,2002]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="232" unit="m" value="440.0">440 m</quantity>
,
<date id="FFE21015FFC7B230FF46F865C754F8A4" box="[151,263,2014,2038]" pageId="2" pageNumber="232" value="2008-10-22">22.x.2008</date>
- luz UV e branca, Calor, Mariano &amp; Mateus colls.
</paragraph>
</subSubSection>
<caption id="DF23665DFFC6B231FF46F81AC7AEF8A7" httpUri="https://zenodo.org/record/222477/files/figure.png" pageId="3" pageNumber="233" targetBox="[286,1309,193,1928]" targetPageId="3">
<paragraph id="8BE336D5FFC6B231FF46F81AC7AEF8A7" blockId="3.[151,1436,1953,2037]" pageId="3" pageNumber="233">
<emphasis id="B928EAC7FFC6B231FF46F81AC710F8E5" bold="true" box="[151,323,1953,1976]" pageId="3" pageNumber="233">FIGURES 2–9.</emphasis>
Wings of male imagos. 2,
<taxonomicName id="4C5C4D56FFC6B231FD57F819C5D9F8EA" box="[646,906,1954,1976]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="luizgonzagai" status="sp. nov.">
<emphasis id="B928EAC7FFC6B231FD57F819C5D9F8EA" box="[646,906,1954,1976]" italics="true" pageId="3" pageNumber="233">Thraulodes luizgonzagai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC6B231FC48F819C5A1F8E5" box="[921,1010,1954,1975]" pageId="3" pageNumber="233" rank="species">
<emphasis id="B928EAC7FFC6B231FC48F819C5A1F8E5" bold="true" box="[921,1010,1954,1975]" pageId="3" pageNumber="233">sp. nov.</emphasis>
</taxonomicNameLabel>
, forewing; 3, idem, hind wing; 4,
<taxonomicName id="4C5C4D56FFC6B231FA58F818C76AF884" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="sternimaculatus" status="sp. nov.">
<emphasis id="B928EAC7FFC6B231FA58F818C76AF884" italics="true" pageId="3" pageNumber="233">T. sternimaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC6B231FEEEF87BC7C3F887" box="[319,400,1984,2005]" pageId="3" pageNumber="233" rank="species">
<emphasis id="B928EAC7FFC6B231FEEEF87BC7C3F887" bold="true" box="[319,400,1984,2005]" pageId="3" pageNumber="233">sp. nov.</emphasis>
</taxonomicNameLabel>
, forewing; 5, idem, hind wing; 6,
<taxonomicName id="4C5C4D56FFC6B231FD21F87AC534F884" box="[752,871,1984,2006]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="alapictus" status="sp. nov.">
<emphasis id="B928EAC7FFC6B231FD21F87AC534F884" box="[752,871,1984,2006]" italics="true" pageId="3" pageNumber="233">T. alapictus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC6B231FCBCF87BC5EDF887" box="[877,958,1984,2005]" pageId="3" pageNumber="233" rank="species">
<emphasis id="B928EAC7FFC6B231FCBCF87BC5EDF887" bold="true" box="[877,958,1984,2005]" pageId="3" pageNumber="233">sp. nov.</emphasis>
</taxonomicNameLabel>
, forewing; 7, idem, hind wing; 8,
<taxonomicName id="4C5C4D56FFC6B231FACFF87AC32BF884" box="[1310,1400,1984,2006]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="pinhoi">
<emphasis id="B928EAC7FFC6B231FACFF87AC32BF884" box="[1310,1400,1984,2006]" italics="true" pageId="3" pageNumber="233">T. pinhoi</emphasis>
</taxonomicName>
sp, nov., forewing; 9, idem, hind wing.
</paragraph>
</caption>
<caption id="DF23665DFFC1B236FF46F8FCC77EF8E8" httpUri="https://zenodo.org/record/222478/files/figure.png" pageId="4" pageNumber="234" targetBox="[195,1406,193,1776]" targetPageId="4">
<paragraph id="8BE336D5FFC1B236FF46F8FCC77EF8E8" blockId="4.[151,1436,1863,1978]" pageId="4" pageNumber="234">
<emphasis id="B928EAC7FFC1B236FF46F8FCC706F80F" bold="true" box="[151,341,1863,1886]" pageId="4" pageNumber="234">FIGURES 10–17.</emphasis>
Genitalia of male imagos. 10,
<taxonomicName id="4C5C4D56FFC1B236FD46F8F3C5C7F80C" box="[663,916,1864,1886]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="234" phylum="Arthropoda" rank="species" species="luizgonzagai" status="sp. nov.">
<emphasis id="B928EAC7FFC1B236FD46F8F3C5C7F80C" box="[663,916,1864,1886]" italics="true" pageId="4" pageNumber="234">Thraulodes luizgonzagai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC1B236FC4DF8F3C5BEF80F" box="[924,1005,1864,1885]" pageId="4" pageNumber="234" rank="species">
<emphasis id="B928EAC7FFC1B236FC4DF8F3C5BEF80F" bold="true" box="[924,1005,1864,1885]" pageId="4" pageNumber="234">sp. nov.</emphasis>
</taxonomicNameLabel>
, genitalia (ventral view); 11, idem, penes (ventral view); 12,
<taxonomicName id="4C5C4D56FFC1B236FEB1F8D3C44EF82E" box="[352,541,1894,1917]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="234" phylum="Arthropoda" rank="species" species="sternimaculatus" status="sp. nov.">
<emphasis id="B928EAC7FFC1B236FEB1F8D3C44EF82E" box="[352,541,1894,1917]" italics="true" pageId="4" pageNumber="234">T. sternimaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC1B236FDF4F8DCC42BF82E" box="[549,632,1895,1916]" pageId="4" pageNumber="234" rank="species">
<emphasis id="B928EAC7FFC1B236FDF4F8DCC42BF82E" bold="true" box="[549,632,1895,1916]" pageId="4" pageNumber="234">sp. nov.</emphasis>
</taxonomicNameLabel>
, genitalia (ventral view); 13, idem, penes (ventral view); 14,
<taxonomicName id="4C5C4D56FFC1B236FB2CF8D3C326F82E" box="[1277,1397,1894,1917]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="234" phylum="Arthropoda" rank="species" species="alapictus" status="sp. nov.">
<emphasis id="B928EAC7FFC1B236FB2CF8D3C326F82E" box="[1277,1397,1894,1917]" italics="true" pageId="4" pageNumber="234">T. alapictus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC1B236FAACF8DCC690F8C9" pageId="4" pageNumber="234" rank="species">
<emphasis id="B928EAC7FFC1B236FAACF8DCC690F8C9" bold="true" pageId="4" pageNumber="234">sp. nov.</emphasis>
</taxonomicNameLabel>
, genitalia (ventral view); 15, idem, penes (ventral view); 16,
<taxonomicName id="4C5C4D56FFC1B236FC90F83DC5CFF8C9" box="[833,924,1925,1947]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="234" phylum="Arthropoda" rank="species" species="pinhoi">
<emphasis id="B928EAC7FFC1B236FC90F83DC5CFF8C9" box="[833,924,1925,1947]" italics="true" pageId="4" pageNumber="234">T. pinhoi</emphasis>
</taxonomicName>
sp, nov. genitalia (ventral view); 17, idem, penes (ventral view).
</paragraph>
</caption>
<caption id="DF23665DFFC0B237FF46F944C299F866" httpUri="https://zenodo.org/record/222479/files/figure.png" pageId="5" pageNumber="235" targetBox="[166,1436,193,1741]" targetPageId="5">
<paragraph id="8BE336D5FFC0B237FF46F944C299F866" blockId="5.[151,1436,1791,1844]" pageId="5" pageNumber="235">
<emphasis id="B928EAC7FFC0B237FF46F944C70AF846" bold="true" box="[151,345,1791,1814]" pageId="5" pageNumber="235">FIGURES 18–22.</emphasis>
<taxonomicName id="4C5C4D56FFC0B237FEB4F944C436F847" box="[357,613,1791,1813]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="235" phylum="Arthropoda" rank="species" species="luizgonzagai" status="sp. nov.">
<emphasis id="B928EAC7FFC0B237FEB4F944C436F847" box="[357,613,1791,1813]" italics="true" pageId="5" pageNumber="235">Thraulodes luizgonzagai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC0B237FDA3F8BBC494F847" box="[626,711,1792,1813]" pageId="5" pageNumber="235" rank="species">
<emphasis id="B928EAC7FFC0B237FDA3F8BBC494F847" bold="true" box="[626,711,1792,1813]" pageId="5" pageNumber="235">sp. nov.</emphasis>
</taxonomicNameLabel>
: 18, lateral view (freshly killed specimen); 19, lateral view (fixed specimen); 20, dorsal view (fixed specimen); 21, detail of prosternum; 22, detail of foreleg (scales: 1mm).
</paragraph>
</caption>
<subSubSection id="C346655EFFC0B237FF16F8DBC34AF8CF" pageId="5" pageNumber="235" type="etymology">
<paragraph id="8BE336D5FFC0B237FF16F8DBC34AF8CF" blockId="5.[151,1436,1888,2022]" pageId="5" pageNumber="235">
<emphasis id="B928EAC7FFC0B237FF16F8DBC71DF82B" bold="true" box="[199,334,1888,1913]" pageId="5" pageNumber="235">Etymology.</emphasis>
The specific epithet is a tribute to Luiz Gonzaga, famous singer from the state of Pernambuco, who sang beautiful songs about the semi-arid region of Pernambuco, collection site of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C346655EFFC0B234FF16F813C277FFA5" lastPageId="6" lastPageNumber="236" pageId="5" pageNumber="235" type="discussion">
<paragraph id="8BE336D5FFC0B234FF16F813C277FFA5" blockId="5.[151,1436,1888,2022]" lastBlockId="6.[151,1436,151,248]" lastPageId="6" lastPageNumber="236" pageId="5" pageNumber="235">
<emphasis id="B928EAC7FFC0B237FF16F813C768F893" bold="true" box="[199,315,1960,1985]" pageId="5" pageNumber="235">Remarks.</emphasis>
<taxonomicName id="4C5C4D56FFC0B237FE93F813C40BF893" box="[322,600,1960,1985]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="235" phylum="Arthropoda" rank="species" species="luizgonzagai" status="sp. nov.">
<emphasis id="B928EAC7FFC0B237FE93F813C40BF893" box="[322,600,1960,1985]" italics="true" pageId="5" pageNumber="235">Thraulodes luizgonzagai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57BCFFC0B237FDB1F813C4E9F893" box="[608,698,1960,1985]" pageId="5" pageNumber="235" rank="species">
<emphasis id="B928EAC7FFC0B237FDB1F813C4E9F893" bold="true" box="[608,698,1960,1985]" pageId="5" pageNumber="235">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be more related to
<taxonomicName id="4C5C4D56FFC0B237FBDEF811C304F893" authority="Traver, 1964" authorityName="Traver" authorityYear="1964" box="[1039,1367,1960,1985]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="235" phylum="Arthropoda" rank="species" species="paysandensis">
<emphasis id="B928EAC7FFC0B237FBDEF811C293F893" box="[1039,1216,1960,1985]" italics="true" pageId="5" pageNumber="235">T. paysandensis</emphasis>
Traver, 1964
</taxonomicName>
. Both are small species (with body length up to
<quantity id="4CA49B30FFC0B237FDABF876C492F8B7" box="[634,705,1997,2021]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="235" unit="mm" value="8.0">8 mm</quantity>
), with femora twice banded, and lobes of penes showing lateral pouch and long and narrow subapical spines. Nevertheless, the new species differs from
<taxonomicName id="4C5C4D56FFC3B234FB46FF22C319FFE2" box="[1175,1354,151,176]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="236" phylum="Arthropoda" rank="species" species="paysandensis">
<emphasis id="B928EAC7FFC3B234FB46FF22C319FFE2" box="[1175,1354,151,176]" italics="true" pageId="6" pageNumber="236">T. paysandensis</emphasis>
</taxonomicName>
by the abdominal color pattern, the shape of medial projection of styliger plate (narrow and rounded in the new species, dome-shaped in
<taxonomicName id="4C5C4D56FFC3B234FE81FF5AC453FFAA" box="[336,512,223,248]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="236" phylum="Arthropoda" rank="species" species="paysandensis">
<emphasis id="B928EAC7FFC3B234FE81FF5AC453FFAA" box="[336,512,223,248]" italics="true" pageId="6" pageNumber="236">T. paysandensis</emphasis>
</taxonomicName>
), and by the absence of recurved folds on penes.
</paragraph>
</subSubSection>
</treatment>
</document>
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