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<document id="95F4AC0DB6C0EE48B2403F1DCBEF0C06" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625278689308" checkinUser="felipe" docAuthor="Paulsen, M. J. &amp; Mondaca, José" docDate="2006" docId="03D03943FFF1FFAEFEF4BB87FCD3FC51" docLanguage="en" docName="zt01191p019.pdf" docOrigin="Zootaxa 1191 (1191)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hilophyllus argentinensis Paulsen &amp; Mondaca 2006, new combination" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="FFE9413BFFFAFFA0FFFCB923FF85FFDF" masterDocTitle="Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="12" updateTime="1698985734564" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BA722BDBCC6BE22BFCD3567E74259FF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6DF0D42F5C6D530FC7B8FCF517B686E0">
<mods:title id="133559C78F0F37B5F5C798D960D83799">Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species</mods:title>
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<mods:name id="445D169A03E1F6956F901781528CBA54" type="personal">
<mods:role id="12B873BFCF12EC14C0CCC49B32CFBAED">
<mods:roleTerm id="7B8491A586FEE3D7AF4424BD4B6AC564">Author</mods:roleTerm>
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<mods:namePart id="278A8CE5D010C24EF7D3937B772368AD">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="82DD8433BE5B6BC6896B4FBAAE3B9044">Division of Entomology, University of Nebraska State Museum, W 436 Nebraska Hall, Lincoln, NE, USA,</mods:affiliation>
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<mods:namePart id="06935666DEDCE359E9ABF730ACB61A0D">Mondaca, José</mods:namePart>
</mods:name>
<mods:typeOfResource id="336E73303ECC6CB82584FCEDA4D30E11">text</mods:typeOfResource>
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<mods:title id="AEC4D08BAD961B28AB4AB39018C6D8E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2A92731B189DA9543C0F9ECCB38E213E">
<mods:date id="8C0317856F9F67C281EC2D60E4B4FB63">2006</mods:date>
<mods:detail id="D63AC7976560BA1BF20DFCFE73936EB3" type="pubDate">
<mods:number id="8767288F30D66CF97B8FDBB1387B1DB6">2006-05-01</mods:number>
</mods:detail>
<mods:detail id="B6C78554C6647AA2A7D74C24FC155E0C" type="volume">
<mods:number id="2E176929D8F0B200450B6131BF093B99">1191</mods:number>
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<mods:detail id="C8934D1D64E840E93B39152DF3A4271E" type="issue">
<mods:number id="D8AD20AEF09E730FD837AB79F06C24C7">1191</mods:number>
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<mods:start id="5566E3A7674D9C2934F30D0B3EF60867">1</mods:start>
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<mods:classification id="C38CAA378F415DBFAAA010B80BEBF279">journal article</mods:classification>
<mods:identifier id="8BDDAB2E7C31439284B036D3DA0AF40C" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="550CA2B28CFC5401B712275D70D1C44B" type="ZooBank">829B214A-C06F-42CB-B1A8-D729C631DF55</mods:identifier>
</mods:mods>
<treatment id="03D03943FFF1FFAEFEF4BB87FCD3FC51" LSID="urn:lsid:plazi:treatment:03D03943FFF1FFAEFEF4BB87FCD3FC51" httpUri="http://treatment.plazi.org/id/03D03943FFF1FFAEFEF4BB87FCD3FC51" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C363DBDEFFF1FFABFEF4BB87FC78FD61" box="[264,1021,676,702]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BC68855FFF1FFABFEF4BB87FC78FD61" blockId="11.[264,1021,676,737]" box="[264,1021,676,702]" pageId="11" pageNumber="12">
<heading id="D08E3F39FFF1FFABFEF4BB87FC78FD61" bold="true" box="[264,1021,676,702]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B90D5447FFF1FFABFEF4BB87FC78FD61" bold="true" box="[264,1021,676,702]" pageId="11" pageNumber="12">
<taxonomicName id="4C79F3D6FFF1FFABFEF4BB87FC9CFD61" ID-CoL="3LXHY" authority="(Martinez, 1981)" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" baseAuthorityName="Martinez" baseAuthorityYear="1981" box="[264,793,676,702]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis" status="comb. nov.">
<emphasis id="B90D5447FFF1FFABFEF4BB87FDC5FD61" bold="true" box="[264,576,676,702]" italics="true" pageId="11" pageNumber="12">Hilophyllus argentinensis</emphasis>
(
<bibRefCitation id="EFE8F5A4FFF1FFABFDB3BB87FC95FD61" author="Martinez, A." box="[591,784,676,702]" pageId="11" pageNumber="12" pagination="13 - 19" refId="ref8179" refString="Martinez, A. (1981) Un nuevo Ceratognathus neotropical (Coleoptera: Lucanidae). Boletin de Entomologia Venezolana N. S., 2 (2), 13 - 19." type="journal article" year="1981">Martínez, 1981</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A23EE93CFFF1FFABFCDABB87FC78FD61" box="[806,1021,676,702]" pageId="11" pageNumber="12" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363DBDEFFF1FFABFEF4BBE4FD07FD3E" box="[264,642,711,737]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BC68855FFF1FFABFEF4BBE4FD07FD3E" blockId="11.[264,1021,676,737]" box="[264,642,711,737]" pageId="11" pageNumber="12">
(
<figureCitation id="134294D0FFF1FFABFEEDBBE4FE04FD3E" box="[273,385,711,737]" captionStart="FIGURES 1–6" captionStartId="8.[264,382,1611,1635]" captionTargetBox="[272,1285,617,1590]" captionTargetId="figure-176@8.[264,1323,609,1604]" captionTargetPageId="8" captionText="FIGURES 1–6. 1, dorsal habitus of male Hilophyllus penai. 2, dorsal habitus of male H. argentinensis. 3, dorsal habitus of male holotype H. martinezi, n.sp. 4, dorsal view of head of female H. penai showing smaller antennal club, eyes, and mandibles. 5, dorsal view of head of male H. penai showing eyes and mandibles. 6, dorsal view of head of male H. argentinensis showing large eyes and bent, interlocking mandibles. Scale bar = 2 mm (Figs. 1–3) or 1 mm (Figs. 4–6)." pageId="11" pageNumber="12">Figs. 2, 6</figureCitation>
,
<figureCitation id="134294D0FFF1FFABFE71BBE4FE18FD3E" box="[397,413,711,737]" captionStart="FIGURES 7–12" captionStartId="9.[264,382,1737,1761]" captionTargetBox="[295,1303,331,1716]" captionTargetId="figure-74@9.[295,1303,326,1716]" captionTargetPageId="9" captionText="FIGURES 7–12. 7, right protibiae of Hilophyllus penai, female (a) and male (b). 8, right protibiae of H. argentinensis female (a) and male (b). 9, right protibia of H. martinezi holotype male. 10, right protibia of Mitophyllus irroratus male. 11, right protibia of Ceratognathus niger female. 12, right protibia of Holloceratognathus helotoides male." pageId="11" pageNumber="12">8</figureCitation>
,
<figureCitation id="134294D0FFF1FFABFE57BBE4FE73FD3E" box="[427,502,711,737]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutely­wrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="11" pageNumber="12">13, 15</figureCitation>
,
<figureCitation id="134294D0FFF1FFABFDFFBBE4FDCBFD3E" box="[515,590,711,737]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="11" pageNumber="12">20, 23</figureCitation>
,
<figureCitation id="134294D0FFF1FFABFDA7BBE4FDFCFD3E" box="[603,633,711,737]" captionStart="FIGURE 25" captionStartId="15.[264,368,1372,1396]" captionTargetBox="[296,1310,326,1355]" captionTargetId="figure-88@15.[264,1323,326,1367]" captionTargetPageId="15" captionText="FIGURE 25. Localities from which Hilophyllus specimens were examined: H. penai (asterisks), H. argentinensis (squares), and H. martinezi (triangle)." pageId="11" pageNumber="12">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C363DBDEFFF1FFABFEF4BA31FC90FCF7" box="[264,789,785,809]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BC68855FFF1FFABFEF4BA31FC90FCF7" blockId="11.[264,789,785,809]" box="[264,789,785,809]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB69AF7BFFF1FFABFEF4BA31FC90FCF7" box="[264,789,785,809]" pageId="11" pageNumber="12">
<treatmentCitation id="0AD8AE44FFF1FFABFEF4BA31FC94FCF7" author="Martinez, A." box="[264,785,785,809]" page="13" pageId="11" pageNumber="12" year="1981">
<taxonomicName id="4C79F3D6FFF1FFABFEF4BA31FC94FCF7" ID-CoL="SN5Z" authority="Martinez, 1981: 13" authorityName="Martinez" authorityPageNumber="13" authorityYear="1981" box="[264,785,785,809]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFEF4BA31FDB9FCF6" box="[264,572,786,809]" italics="true" pageId="11" pageNumber="12">Ceratognathus argentinensis</emphasis>
<bibRefCitation id="EFE8F5A4FFF1FFABFDBFBA32FC94FCF7" author="Martinez, A." box="[579,785,785,808]" pageId="11" pageNumber="12" pagination="13 - 19" refId="ref8179" refString="Martinez, A. (1981) Un nuevo Ceratognathus neotropical (Coleoptera: Lucanidae). Boletin de Entomologia Venezolana N. S., 2 (2), 13 - 19." type="journal article" year="1981">Martínez, 1981: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C363DBDEFFF1FFAEFEF4BA77FCD3FC51" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BC68855FFF1FFABFEF4BA77FE3BFCB2" blockId="11.[264,1325,852,1878]" box="[264,446,852,877]" pageId="11" pageNumber="12">
<emphasis id="B90D5447FFF1FFABFEF4BA77FE3BFCB2" box="[264,446,852,877]" italics="true" pageId="11" pageNumber="12">Type specimens</emphasis>
</paragraph>
<paragraph id="8BC68855FFF1FFACFEC4BA5FFE3FFEE9" blockId="11.[264,1325,852,1878]" lastBlockId="12.[264,442,284,310]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<typeStatus id="54C236F7FFF1FFABFEC4BA5FFE20FC49" box="[312,421,892,918]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
male lost. Both the
<typeStatus id="54C236F7FFF1FFABFD5DBA5FFC8DFC49" box="[673,776,892,918]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C236F7FFF1FFABFCBFBA5FFC24FC49" box="[835,929,892,918]" pageId="11" pageNumber="12" type="allotype">allotype</typeStatus>
of
<taxonomicName id="4C79F3D6FFF1FFABFC37BA5FFB09FC4A" authorityName="Martinez" authorityYear="1981" box="[971,1164,892,917]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFC37BA5FFB09FC4A" box="[971,1164,892,917]" italics="true" pageId="11" pageNumber="12">C. argentinensis</emphasis>
</taxonomicName>
were present in the Martínez collection in 1994 as evidenced by a photograph taken at that time by Gerardo Arriagada (Las Cruces,
<collectingCountry id="F36EC8C5FFF1FFABFD65BAEFFD5BFC39" box="[665,734,972,998]" name="Chile" pageId="11" pageNumber="12">Chile</collectingCountry>
). Approximately one week after the photo was taken, Henry Howden purchased the Martínez collection, including non­primary types. The specimens purchased by Howden were subsequently donated to the CMNC, including the
<typeStatus id="54C236F7FFF1FFABFEC9BD67FE16FB81" box="[309,403,1092,1118]" pageId="11" pageNumber="12" type="allotype">allotype</typeStatus>
of
<taxonomicName id="4C79F3D6FFF1FFABFE42BD67FDFAFB82" authorityName="Martinez" authorityYear="1981" box="[446,639,1092,1117]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFE42BD67FDFAFB82" box="[446,639,1092,1117]" italics="true" pageId="11" pageNumber="12">C. argentinensis</emphasis>
</taxonomicName>
. The
<typeStatus id="54C236F7FFF1FFABFD2CBD67FCB3FB81" box="[720,822,1092,1118]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
was deposited in the collection of Museo Argentino de Ciencias Naturales, Buenos Aires, the depository of the majority of Martínez primary types. The first of three boxes of Martínez types was later stolen from the MACN (F. Ocampo, UNSM, personal communication). The specimens in each box were arranged alphabetically by genus, and as the second box begins with “
<taxonomicName id="4C79F3D6FFF1FFABFC20BDC7FBDCFB22" box="[988,1113,1252,1277]" class="Insecta" family="Hybosoridae" genus="Chaetodus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B90D5447FFF1FFABFC20BDC7FBDCFB22" box="[988,1113,1252,1277]" italics="true" pageId="11" pageNumber="12">Chaetodus</emphasis>
</taxonomicName>
”, the
<typeStatus id="54C236F7FFF1FFABFB5FBDC7FA8CFB21" box="[1187,1289,1252,1278]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C79F3D6FFF1FFABFEF4BC2FFDDEFAFA" authorityName="Martinez" authorityYear="1981" box="[264,603,1292,1317]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFEF4BC2FFDDEFAFA" box="[264,603,1292,1317]" italics="true" pageId="11" pageNumber="12">Ceratognathus argentinensis</emphasis>
</taxonomicName>
was in the box that was stolen and is lost. We designate a
<typeStatus id="54C236F7FFF1FFABFEF4BC17FEE0FA91" box="[264,357,1332,1358]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
to maintain stability in the nomenclature. Because the
<typeStatus id="54C236F7FFF1FFABFBC2BC17FB18FA91" box="[1086,1181,1332,1358]" pageId="11" pageNumber="12" type="allotype">allotype</typeStatus>
is in poor condition with the abdomen missing we are choosing a recently collected male as the
<typeStatus id="54C236F7FFF1FFABFEF4BCA7FEEDFA41" box="[264,360,1412,1438]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
. The specimen chosen matches the original description of the
<typeStatus id="54C236F7FFF1FFABFB8BBCA7FB58FA41" box="[1143,1245,1412,1438]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
in all characters and is from approximately
<quantity id="4C8125B0FFF1FFABFD1EBC8FFCB7FA19" box="[738,818,1452,1478]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="km" value="45.0">45 km</quantity>
northeast of the type locality in similar habitat.
<emphasis id="B90D5447FFF1FFABFE8EBCF7FCA0FA31" bold="true" box="[370,805,1492,1518]" pageId="11" pageNumber="12">
<typeStatus id="54C236F7FFF1FFABFE8EBCF7FD85FA31" box="[370,512,1492,1518]" pageId="11" pageNumber="12" type="neotype">NEOTYPE</typeStatus>
HERE DESIGNATED
</emphasis>
: deposited at MNNC, labeled: a) “
<collectingCountry id="F36EC8C5FFF1FFABFB35BCF7FAA3FA31" box="[1225,1318,1492,1518]" name="Chile" pageId="11" pageNumber="12">CHILE</collectingCountry>
: Región IX (
<collectingRegion id="49BD46B7FFF1FFABFE69BCDFFD94F9C9" box="[405,529,1532,1558]" country="Chile" name="Araucania" pageId="11" pageNumber="12">Araucania</collectingRegion>
) / P.N. Villarrica; Puesco,
<quantity id="4C8125B0FFF1FFABFCADBCDFFC19F9C9" box="[849,924,1532,1558]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="11" pageNumber="12" unit="m" value="732.0">732 m</quantity>
/
<geoCoordinate id="EE4DEE92FFF1FFABFC4ABCDFFBDBF9C9" box="[950,1118,1532,1558]" degrees="39" direction="south" minutes="31" orientation="latitude" pageId="11" pageNumber="12" precision="1" seconds="59.6" value="-39.533222">S 39º31’59.6”</geoCoordinate>
<geoCoordinate id="EE4DEE92FFF1FFABFB9BBCDFFA9CF9C9" box="[1127,1305,1532,1558]" degrees="71" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="1" seconds="19.6" value="-71.55545">W 71º33’19.6”</geoCoordinate>
/ BL/MV light; M.J. Paulsen / &amp; J. Mondaca;
<date id="FFC7AE95FFF1FFABFC87BF07FBA8F9E1" box="[891,1069,1572,1598]" pageId="11" pageNumber="12" value="2004-12-08">8 DEC 2004</date>
”; b) on red paper “
<emphasis id="B90D5447FFF1FFABFEE9BF6FFE57F9BA" box="[277,466,1612,1637]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C79F3D6FFF1FFABFEE9BF6FFE46F9BA" baseAuthorityName="sensu Holloway" baseAuthorityYear="1998" box="[277,451,1612,1637]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Ceratognathus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C79F3D6FFF1FFABFE25BF6FFD62F9B9" authority="Martinez" authorityName="Martinez" authorityYear="1981" box="[473,743,1612,1638]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFE25BF6FFDF3F9BA" box="[473,630,1612,1637]" italics="true" pageId="11" pageNumber="12">argentinensis</emphasis>
Martínez
</taxonomicName>
/
<typeStatus id="54C236F7FFF1FFABFD01BF6FFC06F9B9" box="[765,899,1612,1638]" pageId="11" pageNumber="12" type="neotype">NEOTYPE</typeStatus>
/ det. Paulsen &amp; Mondaca 2006”; c) “
<emphasis id="B90D5447FFF1FFABFEE9BF57FE37F952" box="[277,434,1652,1677]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C79F3D6FFF1FFABFEE9BF57FE1BF952" authorityName="Paulsen and Mondaca" authorityYear="2006" box="[277,414,1652,1677]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Hilophyllus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C79F3D6FFF1FFABFE43BF57FD63F951" authority="(Martinez)" baseAuthorityName="Martinez" box="[447,742,1652,1678]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFE43BF57FDD9F952" box="[447,604,1652,1677]" italics="true" pageId="11" pageNumber="12">argentinensis</emphasis>
(Martínez)
</taxonomicName>
/ det. M.J.
<bibRefCitation id="EFE8F5A4FFF1FFABFC83BF57FBA2F951" author="Paulsen, M. J. &amp; A. D. Smith" box="[895,1063,1652,1678]" pageId="11" pageNumber="12" pagination="45 - 60" refId="ref8260" refString="Paulsen, M. J. &amp; A. D. Smith (2005) A new species of stag beetle from sand dunes in west Texas, and a synopsis of the genus Nicagus (Coleoptera: Lucanidae: Aesalinae: Nicagini). Zootaxa, 1050, 45 - 60." type="journal article" year="2005">Paulsen 2005</bibRefCitation>
”.
<typeStatus id="54C236F7FFF1FFABFBB3BF57FB30F951" box="[1103,1205,1652,1678]" pageId="11" pageNumber="12" type="allotype">Allotype</typeStatus>
(CMNC) labeled: a) “
<collectingCountry id="F36EC8C5FFF1FFABFE68BFBFFDC7F969" box="[404,578,1692,1718]" name="Argentina" pageId="11" pageNumber="12">ARGENTINA</collectingCountry>
/
<collectingRegion id="49BD46B7FFF1FFABFDA6BFBFFD6DF969" box="[602,744,1692,1718]" country="Argentina" name="Neuquen" pageId="11" pageNumber="12">NEUQUEN</collectingRegion>
/ Lago Curruhe / Grande, s/tronco / J.P.R., H., A. y A. Martínez / Coll. Martínez / Ene. “967”; b) on pink paper, “
<typeStatus id="54C236F7FFF1FFABFC04BFE7FB28F901" box="[1016,1197,1732,1758]" pageId="11" pageNumber="12" type="allotype">ALLOTYPUS</typeStatus>
”; c) on red paper: “
<emphasis id="B90D5447FFF1FFABFE8CBFCFFD63F8DA" box="[368,742,1772,1797]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C79F3D6FFF1FFABFE8CBFCFFD9BF8DA" baseAuthorityName="sensu Holloway" baseAuthorityYear="1998" box="[368,542,1772,1797]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Ceratognathus</taxonomicName>
/
<taxonomicName id="4C79F3D6FFF1FFABFDB5BFCFFD63F8DA" box="[585,742,1772,1797]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">argentinensis</taxonomicName>
</emphasis>
(female symbol)
<emphasis id="B90D5447FFF1FFABFC28BFCFFC59F8DA" box="[980,988,1772,1797]" italics="true" pageId="11" pageNumber="12">/</emphasis>
sp. nov.”
<emphasis id="B90D5447FFF1FFABFB8CBFCFFBFDF8DA" box="[1136,1144,1772,1797]" italics="true" pageId="11" pageNumber="12">/</emphasis>
(typeset) “A. MARTÍNEZ–DET. 19__” (handwritten year illegible); d) “H. &amp; A. HOWDEN / COLLE CTION / ex. A. Martinez coll”; e) “
<emphasis id="B90D5447FFF1FFABFD32BE1FFCEEF88A" box="[718,875,1852,1877]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C79F3D6FFF1FFABFD32BE1FFCD2F88A" authorityName="Paulsen and Mondaca" authorityYear="2006" box="[718,855,1852,1877]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Hilophyllus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C79F3D6FFF1FFABFC84BE1FFB1BF889" authority="(Martinez)" baseAuthorityName="Martinez" box="[888,1182,1852,1878]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF1FFABFC84BE1FFB90F88A" box="[888,1045,1852,1877]" italics="true" pageId="11" pageNumber="12">argentinensis</emphasis>
(Martínez)
</taxonomicName>
/ det. M.J.
<bibRefCitation id="EFE8F5A4FFF6FFACFEF4B83FFE2FFEE9" author="Paulsen, M. J. &amp; A. D. Smith" box="[264,426,284,310]" pageId="12" pageNumber="13" pagination="45 - 60" refId="ref8260" refString="Paulsen, M. J. &amp; A. D. Smith (2005) A new species of stag beetle from sand dunes in west Texas, and a synopsis of the genus Nicagus (Coleoptera: Lucanidae: Aesalinae: Nicagini). Zootaxa, 1050, 45 - 60." type="journal article" year="2005">Paulsen 2005</bibRefCitation>
”.
</paragraph>
<paragraph id="8BC68855FFF6FFACFEF4B84FFE2FFE5A" blockId="12.[264,1325,364,1310]" box="[264,426,364,389]" pageId="12" pageNumber="13">
<emphasis id="B90D5447FFF6FFACFEF4B84FFE2FFE5A" box="[264,426,364,389]" italics="true" pageId="12" pageNumber="13">Redescription</emphasis>
</paragraph>
<paragraph id="8BC68855FFF6FFACFEC4B8B7FD60FBD9" blockId="12.[264,1325,364,1310]" pageId="12" pageNumber="13">
Male (n = 9).
<emphasis id="B90D5447FFF6FFACFE10B8B7FDCEFE71" bold="true" box="[492,587,404,430]" italics="true" pageId="12" pageNumber="13">Length:</emphasis>
<quantity id="4C8125B0FFF6FFACFDA9B8B7FD6DFE71" box="[597,744,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="12" pageNumber="13" unit="mm" value="8.3" valueMax="8.5" valueMin="8.1">8.1–8.5 mm</quantity>
.
<emphasis id="B90D5447FFF6FFACFD02B8B7FCD5FE72" box="[766,848,404,430]" italics="true" pageId="12" pageNumber="13">
<emphasis id="B90D5447FFF6FFACFD02B8B7FCC3FE71" bold="true" box="[766,838,404,430]" italics="true" pageId="12" pageNumber="13">Width</emphasis>
:
</emphasis>
<quantity id="4C8125B0FFF6FFACFCA5B8B7FC69FE71" box="[857,1004,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="12" pageNumber="13" unit="mm" value="3.3" valueMax="3.4" valueMin="3.2">3.2–3.4 mm</quantity>
.
<emphasis id="B90D5447FFF6FFACFBFFB8B7FB32FE71" bold="true" box="[1027,1207,404,430]" italics="true" pageId="12" pageNumber="13">Color/Surface:</emphasis>
Reddishbrown, weakly shining. Dorsum (
<figureCitation id="134294D0FFF6FFACFD63B89FFD62FE09" box="[671,743,444,470]" captionStart="FIGURES 1–6" captionStartId="8.[264,382,1611,1635]" captionTargetBox="[272,1285,617,1590]" captionTargetId="figure-176@8.[264,1323,609,1604]" captionTargetPageId="8" captionText="FIGURES 1–6. 1, dorsal habitus of male Hilophyllus penai. 2, dorsal habitus of male H. argentinensis. 3, dorsal habitus of male holotype H. martinezi, n.sp. 4, dorsal view of head of female H. penai showing smaller antennal club, eyes, and mandibles. 5, dorsal view of head of male H. penai showing eyes and mandibles. 6, dorsal view of head of male H. argentinensis showing large eyes and bent, interlocking mandibles. Scale bar = 2 mm (Figs. 1–3) or 1 mm (Figs. 4–6)." pageId="12" pageNumber="13">Fig. 2</figureCitation>
) almost uniformly covered with large punctures; most with a single elongate, apically acute scale.
<emphasis id="B90D5447FFF6FFACFCB1B8C7FC12FE21" bold="true" box="[845,919,484,510]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
Eyes large, width of eyes together in dorsal view greater than narrowest interocular width. Mandibles (
<figureCitation id="134294D0FFF6FFACFBDABB2FFBE9FDF9" box="[1062,1132,524,550]" captionStart="FIGURES 1–6" captionStartId="8.[264,382,1611,1635]" captionTargetBox="[272,1285,617,1590]" captionTargetId="figure-176@8.[264,1323,609,1604]" captionTargetPageId="8" captionText="FIGURES 1–6. 1, dorsal habitus of male Hilophyllus penai. 2, dorsal habitus of male H. argentinensis. 3, dorsal habitus of male holotype H. martinezi, n.sp. 4, dorsal view of head of female H. penai showing smaller antennal club, eyes, and mandibles. 5, dorsal view of head of male H. penai showing eyes and mandibles. 6, dorsal view of head of male H. argentinensis showing large eyes and bent, interlocking mandibles. Scale bar = 2 mm (Figs. 1–3) or 1 mm (Figs. 4–6)." pageId="12" pageNumber="13">Fig. 6</figureCitation>
) small, abruptly curved inward at middle; dorsal tooth acute; ventral tooth of right mandible acute or reduced to angulation. Antennal club segments elongate, longer than 1.5 times length of head (
<figureCitation id="134294D0FFF6FFACFEACBBA7FE2DFD41" box="[336,424,644,670]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutely­wrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="12" pageNumber="13">Fig. 15</figureCitation>
).
<emphasis id="B90D5447FFF6FFACFE3ABBA7FDCFFD41" bold="true" box="[454,586,644,670]" italics="true" pageId="12" pageNumber="13">Pronotum:</emphasis>
Sides strongly explanate, explanate margin subequal to eye in width. Anterior angle not produced anteriorly (
<figureCitation id="134294D0FFF6FFACFC9EBB8FFC45FD19" box="[866,960,684,710]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="12" pageNumber="13">Fig. 23</figureCitation>
), broadly rounded, appearing nearly contiguous and parallel with anterior margin. Lateral margin broadly rounded, widest at middle.
<emphasis id="B90D5447FFF6FFACFE04BBDFFDC9FCC9" bold="true" box="[504,588,764,790]" italics="true" pageId="12" pageNumber="13">Elytra:</emphasis>
Shape elongate. Surface almost uniformly punctate, most punctures with scales, punctures without scales forming longitudinal rows (not patches). Scales usually of two colors, brown and light testaceous (scales of greased specimens appearing unicolored). Sutural interval and lateral margin elevated at apex. Surface between punctures smooth (
<figureCitation id="134294D0FFF6FFACFDA6BABFFD36FC69" box="[602,691,924,950]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutely­wrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="12" pageNumber="13">Fig. 13</figureCitation>
), notably shining at 40x.
<emphasis id="B90D5447FFF6FFACFC06BABFFBB8FC69" bold="true" box="[1018,1085,924,950]" italics="true" pageId="12" pageNumber="13">Legs:</emphasis>
Mesotibiae lacking external tooth.
<emphasis id="B90D5447FFF6FFACFE3DBAE7FDBBFC01" bold="true" box="[449,574,964,990]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
Apex of last ventrite truncate.
<emphasis id="B90D5447FFF6FFACFC48BAE7FBE8FC01" bold="true" box="[948,1133,964,990]" italics="true" pageId="12" pageNumber="13">Male genitalia:</emphasis>
Parameres with exteriorly curved tooth at apex (
<figureCitation id="134294D0FFF6FFACFD83BACFFD5DFBD9" box="[639,728,1004,1030]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="12" pageNumber="13">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BC68855FFF6FFACFEC4BD37FE05FAC1" blockId="12.[264,1325,364,1310]" pageId="12" pageNumber="13">
Female (n = 5). Differing from male in following characters:
<emphasis id="B90D5447FFF6FFACFBD6BD37FB0CFBF1" bold="true" box="[1066,1161,1044,1070]" italics="true" pageId="12" pageNumber="13">Length:</emphasis>
<quantity id="4C8125B0FFF6FFACFB68BD37FAA2FBF1" box="[1172,1319,1044,1070]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.1" metricValueMin="8.9" pageId="12" pageNumber="13" unit="mm" value="9.0" valueMax="9.1" valueMin="8.9">8.9–9.1 mm</quantity>
.
<emphasis id="B90D5447FFF6FFACFEF4BD1FFEDFFB8A" box="[264,346,1084,1110]" italics="true" pageId="12" pageNumber="13">
<emphasis id="B90D5447FFF6FFACFEF4BD1FFED5FB89" bold="true" box="[264,336,1084,1110]" italics="true" pageId="12" pageNumber="13">Width</emphasis>
:
</emphasis>
<quantity id="4C8125B0FFF6FFACFE94BD1FFE7AFB89" box="[360,511,1084,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="12" pageNumber="13" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6–3.9 mm</quantity>
.
<emphasis id="B90D5447FFF6FFACFDDCBD1FFDEFFB89" bold="true" box="[544,618,1084,1110]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
Width of eyes together in dorsal view subequal to 1/3 interocular width. Mandibles small, not strongly curved inward at middle; dorsal tooth of each mandible reduced, ventral tooth of right mandible acute. Antennal club segments short, as long as 1/2 length of head.
<emphasis id="B90D5447FFF6FFACFD45BD97FCB9FB11" bold="true" box="[697,828,1204,1230]" italics="true" pageId="12" pageNumber="13">Pronotum:</emphasis>
Sides more narrowly explanate. Anterior angles constricted toward head. Lateral margin not uniformly rounded, pinched inward anteriorly.
</paragraph>
<paragraph id="8BC68855FFF6FFACFEF4BC77FEFAFAB2" blockId="12.[264,1323,1364,1710]" box="[264,383,1364,1389]" pageId="12" pageNumber="13">
<emphasis id="B90D5447FFF6FFACFEF4BC77FEFAFAB2" box="[264,383,1364,1389]" italics="true" pageId="12" pageNumber="13">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC68855FFF6FFACFEC4BC5FFE35F971" blockId="12.[264,1323,1364,1710]" pageId="12" pageNumber="13">
This species is distinguished from
<taxonomicName id="4C79F3D6FFF6FFACFD15BC5FFCD1FA4A" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" baseAuthorityName="Paulsen &amp; Mondaca" baseAuthorityYear="2006" box="[745,852,1404,1429]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B90D5447FFF6FFACFD15BC5FFCD1FA4A" box="[745,852,1404,1429]" italics="true" pageId="12" pageNumber="13">H. penai</emphasis>
</taxonomicName>
by the unarmed mesotibiae. Also, it differs from
<taxonomicName id="4C79F3D6FFF6FFACFE57BC87FD92FA62" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" baseAuthorityName="Paulsen &amp; Mondaca" baseAuthorityYear="2006" box="[427,535,1444,1469]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B90D5447FFF6FFACFE57BC87FD92FA62" box="[427,535,1444,1469]" italics="true" pageId="12" pageNumber="13">H. penai</emphasis>
</taxonomicName>
by the elytral scale pattern (
<taxonomicName id="4C79F3D6FFF6FFACFC6EBC87FBDFFA62" box="[914,1114,1444,1469]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF6FFACFC6EBC87FBDFFA62" box="[914,1114,1444,1469]" italics="true" pageId="12" pageNumber="13">H. argentinensis</emphasis>
</taxonomicName>
with patches of contrasting lighter and darker scales, whereas
<taxonomicName id="4C79F3D6FFF6FFACFCD5BCEFFC0BFA3A" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" baseAuthorityName="Paulsen &amp; Mondaca" baseAuthorityYear="2006" box="[809,910,1484,1509]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B90D5447FFF6FFACFCD5BCEFFC0BFA3A" box="[809,910,1484,1509]" italics="true" pageId="12" pageNumber="13">H. penai</emphasis>
</taxonomicName>
has patches of light­colored scales contrasting with glabrous areas). Males of
<taxonomicName id="4C79F3D6FFF6FFACFCE3BCD7FC61F9D2" box="[799,996,1524,1549]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF6FFACFCE3BCD7FC61F9D2" box="[799,996,1524,1549]" italics="true" pageId="12" pageNumber="13">H. argentinensis</emphasis>
</taxonomicName>
can be distinguished from either of the remaining species because they possess exceptionally long antennal club segments that are much longer than the head and mandibles together. In addition, they have a distinct genitalic form with hooked parameres and a laterally compressed median lobe (
<figureCitation id="134294D0FFF6FFACFEB6BFB7FE27F971" box="[330,418,1684,1710]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="12" pageNumber="13">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BC68855FFF6FFACFEF4BFC7FD83F921" blockId="12.[264,1323,1764,1870]" box="[264,518,1764,1790]" pageId="12" pageNumber="13">
<emphasis id="B90D5447FFF6FFACFEF4BFC7FE12F922" box="[264,407,1764,1789]" italics="true" pageId="12" pageNumber="13">Distribution</emphasis>
(
<figureCitation id="134294D0FFF6FFACFE5BBFC7FE79F921" box="[423,508,1764,1790]" captionStart="FIGURE 25" captionStartId="15.[264,368,1372,1396]" captionTargetBox="[296,1310,326,1355]" captionTargetId="figure-88@15.[264,1323,326,1367]" captionTargetPageId="15" captionText="FIGURE 25. Localities from which Hilophyllus specimens were examined: H. penai (asterisks), H. argentinensis (squares), and H. martinezi (triangle)." pageId="12" pageNumber="13">Fig. 25</figureCitation>
)
</paragraph>
<paragraph id="8BC68855FFF6FFACFEC4BE2FFCF7F891" blockId="12.[264,1323,1764,1870]" pageId="12" pageNumber="13">
<collectingCountry id="F36EC8C5FFF6FFACFEC4BE2FFE35F8F9" box="[312,432,1804,1830]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
,
<collectingCountry id="F36EC8C5FFF6FFACFE41BE2FFD85F8F9" box="[445,512,1804,1830]" name="Chile" pageId="12" pageNumber="13">Chile</collectingCountry>
. The species is distributed widely in central
<collectingCountry id="F36EC8C5FFF6FFACFBE3BE2FFBDBF8F9" box="[1055,1118,1804,1830]" name="Chile" pageId="12" pageNumber="13">Chile</collectingCountry>
and extends into neighboring areas with similar habitats in
<collectingCountry id="F36EC8C5FFF6FFACFD09BE17FCE8F891" box="[757,877,1844,1870]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8BC68855FFF7FFADFEC4B83FFAA8FE59" blockId="13.[264,1325,284,510]" pageId="13" pageNumber="14">
<materialsCitation id="3B118208FFF7FFADFEC4B83FFD8AFE81" country="Argentina" location="ARGENTINA" pageId="13" pageNumber="14" specimenCount="14">
Locality data:
<specimenCount id="9D7F43DCFFF7FFADFE1DB83FFD05FEE9" box="[481,640,284,310]" pageId="13" pageNumber="14" type="generic">14 specimens</specimenCount>
examined.
<collectingCountry id="F36EC8C5FFF7FFADFEC4B867FE66FE81" box="[312,483,324,350]" name="Argentina" pageId="13" pageNumber="14">ARGENTINA</collectingCountry>
(1)
</materialsCitation>
.
<materialsCitation id="3B118208FFF7FFADFDE7B867FE4EFE59" country="Chile" location="Neuquen" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Neuquen">
<collectingRegion id="49BD46B7FFF7FFADFDE7B867FD00FE81" box="[539,645,324,350]" country="Argentina" name="Neuquen" pageId="13" pageNumber="14">Neuquén</collectingRegion>
(1): Lago Curruhué Grande.
<collectingCountry id="F36EC8C5FFF7FFADFEC4B84FFE0BFE59" box="[312,398,364,390]" name="Chile" pageId="13" pageNumber="14">CHILE</collectingCountry>
(13)
</materialsCitation>
. VIII Región (1): W. of P.N. Nahuelbuta. IX Región (2): P.N. Villarrica
</paragraph>
<paragraph id="8BC68855FFF7FFADFEF4B8B7FAAEFE71" blockId="13.[264,1325,284,510]" box="[264,1323,404,430]" pageId="13" pageNumber="14">(Puesco, Vol. Villarrica). X Región (10): Cordillera Pelada, M. N. Alerce Costero, P. N.</paragraph>
<paragraph id="8BC68855FFF7FFADFEF4B89FFBA6FE21" blockId="13.[264,1325,284,510]" pageId="13" pageNumber="14">Mocho­Choshuenco, R. N. Valdiviana. Temporal distribution. December (2), January (8), February (4).</paragraph>
<caption id="DF06D8DDFFF7FFADFEF4BCE0FB52F948" pageId="13" pageNumber="14" startId="13.[264,383,1475,1499]" targetBox="[264,1323,567,1451]" targetPageId="13">
<paragraph id="8BC68855FFF7FFADFEF4BCE0FB52F948" blockId="13.[264,1323,1475,1687]" pageId="13" pageNumber="14">
<emphasis id="B90D5447FFF7FFADFEF4BCE0FE54FA04" bold="true" box="[264,465,1475,1499]" pageId="13" pageNumber="14">FIGURES 13–18.</emphasis>
13, detail of elytral surface of
<taxonomicName id="4C79F3D6FFF7FFADFCC8BCE7FC63FA04" box="[820,998,1476,1499]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF7FFADFCC8BCE7FC63FA04" box="[820,998,1476,1499]" italics="true" pageId="13" pageNumber="14">H. argentinensis</emphasis>
</taxonomicName>
showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of
<taxonomicName id="4C79F3D6FFF7FFADFBB9BCCAFB4FF9DF" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" box="[1093,1226,1513,1536]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="B90D5447FFF7FFADFBB9BCCAFB4FF9DF" box="[1093,1226,1513,1536]" italics="true" pageId="13" pageNumber="14">H. martinezi</emphasis>
</taxonomicName>
showing scale, puncture, and minutely­wrinkled surface structure. Scale bar = 100 m. 15, right antenna of male
<emphasis id="B90D5447FFF7FFADFEB8BF17FE79F994" box="[324,508,1588,1611]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C79F3D6FFF7FFADFEB8BF17FE7DF994" box="[324,504,1588,1611]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="argentinensis">H. argentinensis</taxonomicName>
.
</emphasis>
Scale bar = 200 m. 16, right antenna of male
<emphasis id="B90D5447FFF7FFADFBEEBF17FB25F994" box="[1042,1184,1588,1611]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C79F3D6FFF7FFADFBEEBF17FB19F994" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" box="[1042,1180,1588,1611]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martinezi">H. martinezi</taxonomicName>
.
</emphasis>
Scale bar = 200 m. 17, scape of right antenna of male
<emphasis id="B90D5447FFF7FFADFD29BF7AFCE7F9AF" box="[725,866,1625,1648]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C79F3D6FFF7FFADFD29BF7AFCDBF9AF" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" box="[725,862,1625,1648]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martinezi">H. martinezi</taxonomicName>
.
</emphasis>
Scale bar = 50 m. 18, external surface of apex of right paramere of male
<taxonomicName id="4C79F3D6FFF7FFADFDA4BFA3FD58F948" authorityName="Paulsen &amp; Mondaca" authorityYear="2006" box="[600,733,1664,1687]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="B90D5447FFF7FFADFDA4BFA3FD58F948" box="[600,733,1664,1687]" italics="true" pageId="13" pageNumber="14">H. martinezi</emphasis>
</taxonomicName>
showing two minute teeth. Scale bar = 50 m.
</paragraph>
</caption>
<paragraph id="8BC68855FFF7FFADFEF4BFE4FEEBF93F" blockId="13.[264,1324,1735,1880]" box="[264,366,1735,1760]" pageId="13" pageNumber="14">
<emphasis id="B90D5447FFF7FFADFEF4BFE4FEEBF93F" box="[264,366,1735,1760]" italics="true" pageId="13" pageNumber="14">Remarks</emphasis>
</paragraph>
<paragraph id="8BC68855FFF7FFAEFEC4BFCDFB7DFDA9" blockId="13.[264,1324,1735,1880]" lastBlockId="14.[264,1323,284,910]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<bibRefCitation id="EFE8F5A4FFF7FFADFEC4BFCDFE73F8D7" author="Martinez, A." box="[312,502,1774,1800]" pageId="13" pageNumber="14" pagination="13 - 19" refId="ref8179" refString="Martinez, A. (1981) Un nuevo Ceratognathus neotropical (Coleoptera: Lucanidae). Boletin de Entomologia Venezolana N. S., 2 (2), 13 - 19." type="journal article" year="1981">Martínez (1981)</bibRefCitation>
indicated that the
<typeStatus id="54C236F7FFF7FFADFD30BFCDFD7BF8D7" box="[716,766,1774,1800]" pageId="13" pageNumber="14">type</typeStatus>
series was collected on a tree trunk, possibly of
<taxonomicName id="4C79F3D6FFF7FFADFEF4BE34FE44F8EF" box="[264,449,1814,1840]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90D5447FFF7FFADFEF4BE34FE0AF8EF" box="[264,399,1815,1840]" italics="true" pageId="13" pageNumber="14">Nothofagus</emphasis>
sp.
</taxonomicName>
, in an advanced state of decomposition in the province of
<collectingRegion id="49BD46B7FFF7FFADFB47BE35FAADF8EF" box="[1211,1320,1814,1840]" country="Argentina" name="Neuquen" pageId="13" pageNumber="14">Neuquén</collectingRegion>
,
<collectingCountry id="F36EC8C5FFF7FFADFEF4BE1DFE04F887" box="[264,385,1854,1880]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
. More recently, adult and immature specimens of
<taxonomicName id="4C79F3D6FFF7FFADFC27BE1CFB18F887" box="[987,1181,1855,1880]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF7FFADFC27BE1CFB18F887" box="[987,1181,1855,1880]" italics="true" pageId="13" pageNumber="14">H. argentinensis</emphasis>
</taxonomicName>
were found in the lower branches of a decomposing coigüe,
<taxonomicName id="4C79F3D6FFF4FFAEFC95B83FFDD4FE81" authority="(Mirbel) Blume (Mondaca &amp; Paulsen 2005)" authorityName="Blume (Mondaca &amp; Paulsen" authorityYear="2005" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="dombeyi">
<emphasis id="B90D5447FFF4FFAEFC95B83FFBE5FEEA" box="[873,1120,284,309]" italics="true" pageId="14" pageNumber="15">Nothofagus dombeyi</emphasis>
(Mirbel) Blume (
<bibRefCitation id="EFE8F5A4FFF4FFAEFEECB867FDCCFE81" author="Mondaca, J. &amp; M. J. Paulsen" box="[272,585,324,350]" pageId="14" pageNumber="15" pagination="21 - 26" refId="ref8214" refString="Mondaca, J. &amp; M. J. Paulsen (2005) Ceratognathus argentinensis Martinez, nuevo registro de Aesalinae (Coleoptera: Lucanidae) para Chile. Revista Chilena de Entomologia, 30 (2), 21 - 26. Dated 2004." type="journal article" year="2005">Mondaca &amp; Paulsen 2005</bibRefCitation>
)
</taxonomicName>
. However, if this species is termitophilous, it is probable that its distribution is limited more by the presence or absence of termites than the range of a specific host tree. In 2006, MJP collected an additional specimen flying in late afternoon at a height of approximately three meters around the trunk of a partially decomposed lenga,
<taxonomicName id="4C79F3D6FFF4FFAEFEA2B8C7FBB4FE21" authority="(Poeppig &amp; Endlicher) Krasser. Furthermore" authorityName="Krasser. Furthermore" baseAuthorityName="Poeppig &amp; Endlicher" box="[350,1073,484,510]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pumilio">
<emphasis id="B90D5447FFF4FFAEFEA2B8C7FE66FE22" box="[350,483,484,509]" italics="true" pageId="14" pageNumber="15">N. pumilio</emphasis>
(Poeppig &amp; Endlicher) Krasser. Furthermore
</taxonomicName>
,
<specimenCount id="9D7F43DCFFF4FFAEFBB9B8C7FB6BFE21" box="[1093,1262,484,510]" pageId="14" pageNumber="15" type="generic">one specimen</specimenCount>
was collected by Dr. Elizabeth Arias (University of California–Berkeley) during insecticidal fogging of
<taxonomicName id="4C79F3D6FFF4FFAEFE61BB17FBDDFD91" authority="(Molina) K. Koch" authorityName="K. Koch" baseAuthorityName="Molina" box="[413,1112,564,590]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="araucana">
<emphasis id="B90D5447FFF4FFAEFE61BB17FD13FD92" box="[413,662,564,589]" italics="true" pageId="14" pageNumber="15">Araucaria araucana</emphasis>
(Molina) K. Koch (Araucariaceae)
</taxonomicName>
west of Parque Nacional Nahuelbuta, suggesting that the species may not be restricted to
<taxonomicName id="4C79F3D6FFF4FFAEFB96BB7FFB74FDAA" box="[1130,1265,604,629]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B90D5447FFF4FFAEFB96BB7FFB74FDAA" box="[1130,1265,604,629]" italics="true" pageId="14" pageNumber="15">Nothofagus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC68855FFF4FFAEFEC4BBA7FCD3FC51" blockId="14.[264,1323,284,910]" pageId="14" pageNumber="15">
In 2004, we collected a single male specimen of
<taxonomicName id="4C79F3D6FFF4FFAEFC72BBA7FBD7FD42" box="[910,1106,644,669]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="argentinensis">
<emphasis id="B90D5447FFF4FFAEFC72BBA7FBD7FD42" box="[910,1106,644,669]" italics="true" pageId="14" pageNumber="15">H. argentinensis</emphasis>
</taxonomicName>
at mercury vapor light in Parque Nacional Villarrica. In 2006, we collected
<specimenCount id="9D7F43DCFFF4FFAEFC53BB8FFBACFD19" box="[943,1065,684,710]" pageId="14" pageNumber="15" type="male">three male</specimenCount>
specimens at Reserva Nacional Valdiviana and
<specimenCount id="9D7F43DCFFF4FFAEFDC9BBF7FD27FD31" box="[565,674,724,750]" pageId="14" pageNumber="15" type="male">one male</specimenCount>
specimen at Parque Nacional Mocho­Choshuenco, all at mercury vapor lights. Previously, there was only anecdotal evidence that this species was attracted to light. It may be relevant that males of this species possess proportionately the largest eyes in the genus; large eyes suggest nocturnal activity, making collection at light traps less surprising although still infrequent.
</paragraph>
</subSubSection>
</treatment>
</document>
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