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<document id="F115AA9EEB84C8EAC2D43419A127FE60" ID-CLB-Dataset="198847" ID-DOI="10.11646/zootaxa.5213.5.1" ID-GBIF-Dataset="2a26dfdb-0e17-40a1-b5fd-14787882b9d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381607" ID-ZooBank="B16F6ED4-A8C2-4F3D-A5DD-20537DE0CF01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1669810135404" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="03FFB253777B7A63DD90FF796DAF5476" docLanguage="en" docName="zootaxa.5213.5.1.pdf" docOrigin="Zootaxa 5213 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus convexicollis Redtenbacher" docType="treatment" docVersion="5" lastPageNumber="465" masterDocId="FFC6CA2B77757A6DDD07FFEE68335217" masterDocTitle="The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="496" masterPageNumber="451" pageNumber="465" updateTime="1699412587891" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9EDD08702238A3586DC9039D9F376605">The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="80741DEF1D44046C24E8DC1F71F680AC">Shavrin, Alexey V.</mods:namePart>
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<treatment id="03FFB253777B7A63DD90FF796DAF5476" ID-DOI="http://doi.org/10.5281/zenodo.7386469" ID-GBIF-Taxon="205348427" ID-Zenodo-Dep="7386469" LSID="urn:lsid:plazi:treatment:03FFB253777B7A63DD90FF796DAF5476" httpUri="http://treatment.plazi.org/id/03FFB253777B7A63DD90FF796DAF5476" lastPageNumber="465" pageId="14" pageNumber="465">
<subSubSection id="C34C50CE777B7A63DD90FF796A4652A6" box="[151,629,151,178]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="8BE90345777B7A63DD90FF796A4652A6" blockId="14.[151,629,151,178]" box="[151,629,151,178]" pageId="14" pageNumber="465">
<heading id="D0A1B429777B7A63DD90FF796A4652A6" bold="true" box="[151,629,151,178]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<emphasis id="B922DF57777B7A63DD90FF796A4652A6" bold="true" box="[151,629,151,178]" pageId="14" pageNumber="465">
Redefinition of the
<taxonomicName id="4C5678C6777B7A63DC8FFF796A1752A5" ID-CoL="3FPGR" authorityName="Redtenbacher" box="[392,548,151,178]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63DC8FFF796A1752A5" bold="true" box="[392,548,151,178]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777B7A63DD90FF376B8D5393" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="8BE90345777B7A63DD90FF376B8D5393" blockId="14.[151,1436,217,856]" pageId="14" pageNumber="465">
<emphasis id="B922DF57777B7A63DD90FF37692152E4" bold="true" box="[151,274,217,243]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
Body medium-sized (
<quantity id="4CAEAEA0777B7A63DF14FF346A8D52E3" box="[531,702,218,245]" metricMagnitude="-3" metricUnit="m" metricValue="6.075" metricValueMax="7.9" metricValueMin="4.25" pageId="14" pageNumber="465" unit="mm" value="6.075" valueMax="7.9" valueMin="4.25">4.25–7.90 mm</quantity>
); pronotum convex, moderately narrow or distinctly transverse, from widest anterior third gradually or sharply narrowed toward subparallel-sided short or elongate laterobasal portion; elytra slightly longer than broad (
<taxonomicName id="4C5678C6777B7A63DF65FECC6B06532B" authorityName="Shavrin" authorityYear="2022" box="[610,821,290,316]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malcolmi">
<emphasis id="B922DF57777B7A63DF65FECC6A45532B" box="[610,630,290,316]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DFCAFECC6B06532B" box="[717,821,290,316]" italics="true" pageId="14" pageNumber="465">malcolmi</emphasis>
</taxonomicName>
) or distinctly broader than long, slightly or significantly broadened posteriad; aedeagus narrow or wide, with parameres slightly or significantly broadened anteriad, with weakly sclerotized internal sac and very narrow short or long flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777B7A63DDC0FE63690F53E7" pageId="14" pageNumber="465" type="discussion">
<paragraph id="8BE90345777B7A63DDC0FE63690F53E7" blockId="14.[151,1436,217,856]" pageId="14" pageNumber="465">
<emphasis id="B922DF57777B7A63DDC0FE6369AB53B0" bold="true" box="[199,408,397,423]" pageId="14" pageNumber="465">Species included.</emphasis>
<taxonomicName id="4C5678C6777B7A63DCA3FE606B7053BF" authority="Luze, 1903" authorityName="Luze" authorityYear="1903" box="[420,835,398,424]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63DCA3FE60698B53BF" box="[420,440,398,424]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DF19FE606A8353BF" box="[542,688,398,424]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
<bibRefCitation id="EFC77EB4777B7A63DFBBFE606B7053BF" author="Luze, G." box="[700,835,398,424]" pageId="14" pageNumber="465" pagination="103 - 117" refId="ref31553" refString="Luze, G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Geodromicus Redtenb. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 103 - 117." type="journal article" year="1903">Luze, 1903</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5678C6777B7A63DE54FE606D2C53BF" authority="Cameron, 1930" authorityName="Cameron" authorityYear="1930" box="[851,1311,398,424]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="kashmirensis">
<emphasis id="B922DF57777B7A63DE54FE606B5453BF" box="[851,871,398,424]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DECBFE606C6E53BF" box="[972,1117,398,424]" italics="true" pageId="14" pageNumber="465">kashmirensis</emphasis>
<bibRefCitation id="EFC77EB4777B7A63D96DFE606D2C53BF" author="Cameron, M." box="[1130,1311,398,424]" pageId="14" pageNumber="465" refId="ref30874" refString="Cameron, M. (1930) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 1. Taylor and Francis, London, xvii + 471 pp." type="book" year="1930">Cameron, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5678C6777B7A63D828FE6069F453DB" authority="Roubal, 1929" authorityName="Roubal" authorityYear="1929" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="lebedevianus">
<emphasis id="B922DF57777B7A63D828FE606D7053BF" box="[1327,1347,398,424]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DD90FE5C691A53DB" box="[151,297,434,460]" italics="true" pageId="14" pageNumber="465">lebedevianus</emphasis>
<bibRefCitation id="EFC77EB4777B7A63DC28FE5C69F453DB" author="Roubal, J." box="[303,455,434,460]" pageId="14" pageNumber="465" pagination="46 - 48" refId="ref31707" refString="Roubal, J. (1929) Vier neue Coleopteren aus SSSR. Entomologische Blatter, 25 (1), 46 - 48." type="journal article" year="1929">Roubal, 1929</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5678C6777B7A63DCD6FE5C6A9653DB" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[465,677,434,460]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malcolmi" status="sp. nov.">
<emphasis id="B922DF57777B7A63DCD6FE5C69D653DB" box="[465,485,434,460]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DF3AFE5C6A9653DB" box="[573,677,434,460]" italics="true" pageId="14" pageNumber="465">malcolmi</emphasis>
</taxonomicName>
<emphasis id="B922DF57777B7A63DFADFE5C6AD153DB" bold="true" box="[682,738,434,460]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="A211622C777B7A63DFADFE5C6AD153DB" box="[682,738,434,460]" pageId="14" pageNumber="465" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C5678C6777B7A63DFE8FE5C6C8253DB" authority="Kashcheev, 1999" authorityName="Kashcheev" authorityYear="1999" box="[751,1201,434,460]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macrothorax">
<emphasis id="B922DF57777B7A63DFE8FE5C6B3053DB" box="[751,771,434,460]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63DE5CFE5C6BD853DB" box="[859,1003,434,460]" italics="true" pageId="14" pageNumber="465">macrothorax</emphasis>
<bibRefCitation id="EFC77EB4777B7A63DEF7FE5C6C8253DB" author="Kashcheev, V. A." box="[1008,1201,434,460]" pageId="14" pageNumber="465" pagination="12 - 16" refId="ref31422" refString="Kashcheev, V. A. (1999) Novye vidy Oxytelinae (Coleoptera, Staphylinidae) fauny Kazakhstana [New species of Oxytelinae (Coleoptera, Staphylinidae) of the fauna of Kazakhstan]. Selevinia, 1996, 12 - 16. [in Russian]" type="journal article" year="1999">Kashcheev, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5678C6777B7A63D9BDFE5C690B53E7" authority="Shavrin, 2009" authorityName="Shavrin" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="B922DF57777B7A63D9BDFE5C6CFD53DB" box="[1210,1230,434,460]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63D821FE5C6DAF53DB" box="[1318,1436,434,460]" italics="true" pageId="14" pageNumber="465">medvedevi</emphasis>
<bibRefCitation id="EFC77EB4777B7A63DD90FE38690B53E7" author="Shavrin, A. V." box="[151,312,470,496]" pageId="14" pageNumber="465" pagination="183 - 184" refId="ref31913" refString="Shavrin, A. V. (2009) A new species and new records of Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from Pakistan. Caucasian Entomological Bulletin, 5 (2), 183 - 184." type="journal article" year="2009">Shavrin, 2009</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777B7A63DDC0FE176AB5502F" pageId="14" pageNumber="465" type="distribution">
<paragraph id="8BE90345777B7A63DDC0FE176AB5502F" blockId="14.[151,1436,217,856]" pageId="14" pageNumber="465">
<emphasis id="B922DF57777B7A63DDC0FE1769525004" bold="true" box="[199,353,505,531]" pageId="14" pageNumber="465">Distribution.</emphasis>
The species of the
<taxonomicName id="4C5678C6777B7A63DF3BFE146A8C5003" authorityName="Bernhauer" authorityYear="1902" box="[572,703,506,532]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B922DF57777B7A63DF3BFE146A8C5003" box="[572,703,506,532]" italics="true" pageId="14" pageNumber="465">bodemeyeri</emphasis>
</taxonomicName>
group are distributed in the Palaearctic Region, and known from the Central Asia and the Himalayan Region.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777B7A63DDC0FDAF69EC51AE" pageId="14" pageNumber="465" type="discussion">
<paragraph id="8BE90345777B7A63DDC0FDAF692750B3" blockId="14.[151,1436,217,856]" pageId="14" pageNumber="465">
<emphasis id="B922DF57777B7A63DDC0FDAF6978504C" bold="true" box="[199,331,577,603]" pageId="14" pageNumber="465">Bionomics.</emphasis>
The species of the
<taxonomicName id="4C5678C6777B7A63DF29FDAC6AF3504B" box="[558,704,578,604]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63DF29FDAC6AF3504B" box="[558,704,578,604]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
</taxonomicName>
group inhabit banks of rivers and streams, and can be found in gravel, under stones, mosses, drift, etc. Specimens were collected at elevations from
<quantity id="4CAEAEA0777B7A63D942FD886CD65097" box="[1093,1253,614,640]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="4.2" metricValueMin="0.7" pageId="14" pageNumber="465" unit="m" value="2450.0" valueMax="4200.0" valueMin="700.0">700 to 4200 m</quantity>
a.s.l. from April to October.
</paragraph>
<paragraph id="8BE90345777B7A63DDC0FD40691B514F" blockId="14.[151,1436,217,856]" pageId="14" pageNumber="465">
<emphasis id="B922DF57777B7A63DDC0FD40690850DF" bold="true" box="[199,315,686,712]" pageId="14" pageNumber="465">Remarks.</emphasis>
Previously,
<bibRefCitation id="EFC77EB4777B7A63DCC4FD406A5D50DF" author="Bordoni, A." box="[451,622,686,712]" pageId="14" pageNumber="465" pagination="19 - 59" refId="ref30793" refString="Bordoni, A. (1984) Appunti per una revisione dei Geodromicus Redt. della regione Palearctica occidentale (Coleoptera, Staphylinidae). Redia, 67, 19 - 59." type="journal article" year="1984">Bordoni (1984)</bibRefCitation>
included
<taxonomicName id="4C5678C6777B7A63DFDEFD406BCD50DF" authorityName="Bordoni" authorityYear="1984" box="[729,1022,686,712]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="hauserianus">
<emphasis id="B922DF57777B7A63DFDEFD406BCD50DF" box="[729,1022,686,712]" italics="true" pageId="14" pageNumber="465">Geodromicus hauserianus</emphasis>
</taxonomicName>
(synonym of
<taxonomicName id="4C5678C6777B7A63D992FD406DA050DF" box="[1173,1427,686,712]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63D992FD406C9A50DF" box="[1173,1193,686,712]" italics="true" pageId="14" pageNumber="465">G</emphasis>
. (s.str.)
<emphasis id="B922DF57777B7A63D806FD406DA050DF" box="[1281,1427,686,712]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
</taxonomicName>
) to the
<emphasis id="B922DF57777B7A63DDD8FD3D692B50FB" box="[223,280,723,748]" italics="true" pageId="14" pageNumber="465">rousi</emphasis>
group (
<bibRefCitation id="EFC77EB4777B7A63DC76FD3C6A3E50FB" author="Shavrin, A. V." box="[369,525,722,748]" pageId="14" pageNumber="465" pagination="191 - 202" refId="ref32174" refString="Shavrin, A. V. (2021) The rousi species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae). Caucasian Entomological Bulletin, 17 (1), 191 - 202. https: // doi. org / 10.23885 / 181433262021171 - 191202" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
). Based on the general shape of the apical portions of the parameres and weakly sclerotized internal sac with narrow flagellum, the
<taxonomicName id="4C5678C6777B7A63DFE5FD186B475107" box="[738,884,758,784]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63DFE5FD186B475107" box="[738,884,758,784]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
</taxonomicName>
group is similar to the
<taxonomicName id="4C5678C6777B7A63D98FFD186D385107" authorityName="Bernhauer" authorityYear="1902" box="[1160,1291,758,784]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="bodemeyeri">
<emphasis id="B922DF57777B7A63D98FFD186D385107" box="[1160,1291,758,784]" italics="true" pageId="14" pageNumber="465">bodemeyeri</emphasis>
</taxonomicName>
group, from which it can be distinguished by the absence of paired sclerotized structures in apical and middle portions of the median lobe.
</paragraph>
<paragraph id="8BE90345777B7A63DD90FC7169EC51AE" blockId="14.[151,479,927,954]" box="[151,479,927,954]" pageId="14" pageNumber="465">
<heading id="D0A1B429777B7A63DD90FC7169EC51AE" bold="true" box="[151,479,927,954]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<emphasis id="B922DF57777B7A63DD90FC7169EC51AE" bold="true" box="[151,479,927,954]" pageId="14" pageNumber="465">
Key to
<taxonomicName id="4C5678C6777B7A63DDF6FC7169BE51AD" box="[241,397,927,954]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63DDF6FC7169BE51AD" bold="true" box="[241,397,927,954]" italics="true" pageId="14" pageNumber="465">convexicollis</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE777B7A63DD90FC0A6DAF5476" pageId="14" pageNumber="465" type="key">
<keyStep id="30A21BE0777B7A63DD90FC0A6DAF5646" pageId="14" pageNumber="465">
<paragraph id="8BE90345777B7A63DD90FC0A6DAF5601" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FC0A6DAF5601" pageId="14" pageNumber="465">
1 Elytra slightly longer than broad, slightly more than twice as long as pronotum. Median lobe narrow, with very wide parameres (
<figureCitation id="136D1FC0777B7A63DDEEFBEF69025600" box="[233,305,1025,1047]" captionStart="FIGURES 64–69" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[173,1414,192,1907]" captionTargetId="figure-15@33.[159,1428,181,1919]" captionTargetPageId="33" captionText="FIGURES 64–69. Aedeagus of Geodromicus malcolmi (Figs. 64–65: holotype) and G. affinis (Fig. 66: holotype; Figs. 67–68: India, Sonamarg; Fig. 69: Pakistan, Utrar): 64, 66–67, 69—parameral view, 65, 68—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381650" httpUri="https://zenodo.org/record/7381650/files/figure.png" pageId="14" pageNumber="465">Fig. 64</figureCitation>
). Body length:
<quantity id="4CAEAEA0777B7A63DCC2FBEC6A2E5600" box="[453,541,1026,1047]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="465" unit="mm" value="4.9">4.90 mm</quantity>
. Habitus as in
<figureCitation id="136D1FC0777B7A63DFAEFBEF6AC35600" box="[681,752,1025,1047]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="14" pageNumber="465">Fig. 60</figureCitation>
.
<collectingCountry id="F34143D5777B7A63DFFEFBEF6B195600" box="[761,810,1025,1047]" name="India" pageId="14" pageNumber="465">India</collectingCountry>
.........................................
<taxonomicName id="4C5678C6777B7A63D821FBEC6DAF5601" authorityName="Shavrin" authorityYear="2022" box="[1318,1436,1025,1047]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malcolmi">
<emphasis id="B922DF57777B7A63D821FBEC6DAF5601" box="[1318,1436,1025,1047]" italics="true" pageId="14" pageNumber="465">G. malcolmi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE90345777B7A63DD90FBF16DAF5646" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FBF16DAF5646" pageId="14" pageNumber="465">- Elytra as broad as or broader than long, distinctly less than twice as long as pronotum. Median lobe narrow or wide, with moderately narrow or wide parameres.................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A21BE0777B7A63DD90FBB46DAF569B" pageId="14" pageNumber="465">
<paragraph id="8BE90345777B7A63DD90FBB46DAF5678" blockId="14.[151,1437,996,1634]" box="[151,1436,1113,1135]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FBB46DAF5678" box="[151,1436,1113,1135]" pageId="14" pageNumber="465">2 Pronotum slightly broader (1.1/1.2) than long.............................................................. 3</keyLead>
</paragraph>
<paragraph id="8BE90345777B7A63DD90FB996DAF569B" blockId="14.[151,1437,996,1634]" box="[151,1436,1143,1165]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FB996DAF569B" box="[151,1436,1143,1165]" pageId="14" pageNumber="465">- Pronotum distinctly transverse, 1.4/1.8 as broad as long...................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A21BE0777B7A63DD90FB7A6DAF572B" pageId="14" pageNumber="465">
<paragraph id="8BE90345777B7A63DD90FB7A6DAF56F3" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FB7A6DAF56F3" pageId="14" pageNumber="465">
3 Pronotum sharply narrowed toward long laterobasal margins. Parameres strongly broadened apically, distinctly longer than median lobe (
<figureCitation id="136D1FC0777B7A63DC6FFB5F698156D0" box="[360,434,1201,1223]" captionStart="FIGURES 52–56" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[287,1291,190,1901]" captionTargetId="figure-31@28.[281,1307,181,1919]" captionTargetPageId="28" captionText="FIGURES 52–56. Aedeagus of Geodromicus lebedevianus (Figs. 52–53: Kazakhstan, Chichkan Valley) and G. macrothorax (Figs. 54–55: Kazakhstan, Chelkydysu; Fig. 56: Kyrgyzstan, Grigoryevka): 52, 54, 56—parameral view, 53, 55—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381642" httpUri="https://zenodo.org/record/7381642/files/figure.png" pageId="14" pageNumber="465">Fig. 52</figureCitation>
). Body yellow-brown to black, usually with wide and long pale macula on each elytron. Body length:
<quantity id="4CAEAEA0777B7A63DDE5FB21694356F3" box="[226,368,1231,1253]" metricMagnitude="-3" metricUnit="m" metricValue="5.385" metricValueMax="6.05" metricValueMin="4.72" pageId="14" pageNumber="465" unit="mm" value="5.385" valueMax="6.05" valueMin="4.72">4.72–6.05 mm</quantity>
. Habitus as in
<figureCitation id="136D1FC0777B7A63DCFAFB216A5456F2" box="[509,615,1231,1253]" captionStart="FIGURES 40–45" captionStartId="25.[151,264,1900,1924]" captionTargetBox="[161,1419,567,1866]" captionTargetId="figure-201@25.[151,1436,561,1877]" captionTargetPageId="25" captionText="FIGURES 40–45. Habitus of Geodromicus: 40—G. kashmirensis (lectotype), 41—G. similis (lectoype), 42—G. consors (holotype), 43—G. karakorus (holotype), 44–45—G. lebedevianus (Fig. 44: Kyrgyzstan, Chichkan Vallley; Fig. 45: Kyrgyzstan, Kara-Balta). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381636" httpUri="https://zenodo.org/record/7381636/files/figure.png" pageId="14" pageNumber="465">Fig. 44–45</figureCitation>
.
<collectingCountry id="F34143D5777B7A63DF76FB216AEC56F3" box="[625,735,1231,1252]" name="Kyrgyzstan" pageId="14" pageNumber="465">Kyrgyzstan</collectingCountry>
............................................
<taxonomicName id="4C5678C6777B7A63D805FB216DAF56F3" ID-CoL="3FPHR" authorityName="Roubal" authorityYear="1929" box="[1282,1436,1231,1252]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="lebedevianus">
<emphasis id="B922DF57777B7A63D805FB216DAF56F3" box="[1282,1436,1231,1252]" italics="true" pageId="14" pageNumber="465">G. lebedevianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE90345777B7A63DD90FB036DAF572B" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FB036DAF572B" pageId="14" pageNumber="465">
- Pronotum from gradually narrowed toward moderately short laterobasal margins. Parameres slightly broadened apically, slightly exceeding apex of median lobe (
<bibRefCitation id="EFC77EB4777B7A63DF76FAE76ACA5708" author="Shavrin, A. V." box="[625,761,1289,1311]" pageId="14" pageNumber="465" pagination="183 - 184" refId="ref31913" refString="Shavrin, A. V. (2009) A new species and new records of Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from Pakistan. Caucasian Entomological Bulletin, 5 (2), 183 - 184." type="journal article" year="2009">Shavrin 2009</bibRefCitation>
: fig. 1). Body brown, without maculae. Body length:
<quantity id="4CAEAEA0777B7A63D80EFAE76DAA5708" box="[1289,1433,1289,1311]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="14" pageNumber="465" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.90–7.20 mm</quantity>
. Habitus as in
<figureCitation id="136D1FC0777B7A63DC62FAC9699F572A" box="[357,428,1319,1341]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="14" pageNumber="465">Fig. 61</figureCitation>
.
<collectingCountry id="F34143D5777B7A63DCB2FAC96A35572A" box="[437,518,1319,1341]" name="Pakistan" pageId="14" pageNumber="465">Pakistan</collectingCountry>
................................................................
<taxonomicName id="4C5678C6777B7A63D81DFAC96DAF572B" ID-CoL="3FPML" authorityName="Shavrin" authorityYear="2009" box="[1306,1436,1319,1340]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="B922DF57777B7A63D81DFAC96DAF572B" box="[1306,1436,1319,1340]" italics="true" pageId="14" pageNumber="465">G. medvedevi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A21BE0777B7A63DD90FAAA6DAF57D8" pageId="14" pageNumber="465">
<paragraph id="8BE90345777B7A63DD90FAAA6DAF5783" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FAAA6DAF5783" pageId="14" pageNumber="465">
4 Pronotum very convex, without or with indistinct longitudinal depression. Median lobe very narrow; parameres significantly exceeding apex of median lobe, with four long apical and three to four subapical setae (
<figureCitation id="136D1FC0777B7A63D936FA8F6C945760" box="[1073,1191,1377,1399]" captionStart="FIGURES 52–56" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[287,1291,190,1901]" captionTargetId="figure-31@28.[281,1307,181,1919]" captionTargetPageId="28" captionText="FIGURES 52–56. Aedeagus of Geodromicus lebedevianus (Figs. 52–53: Kazakhstan, Chichkan Valley) and G. macrothorax (Figs. 54–55: Kazakhstan, Chelkydysu; Fig. 56: Kyrgyzstan, Grigoryevka): 52, 54, 56—parameral view, 53, 55—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381642" httpUri="https://zenodo.org/record/7381642/files/figure.png" pageId="14" pageNumber="465">Figs. 54, 56</figureCitation>
). Body length:
<quantity id="4CAEAEA0777B7A63D83BFA8F69395783" metricMagnitude="-3" metricUnit="m" metricValue="6.375" metricValueMax="7.9" metricValueMin="4.85" pageId="14" pageNumber="465" unit="mm" value="6.375" valueMax="7.9" valueMin="4.85">4.85–7.90 mm</quantity>
. Body yellow-brown to black. Habitus as in
<figureCitation id="136D1FC0777B7A63DFB1FA916B125782" box="[694,801,1407,1429]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="14" pageNumber="465">Fig. 57–59</figureCitation>
.
<collectingCountry id="F34143D5777B7A63DE2DFA916BAF5782" box="[810,924,1407,1429]" name="Kazakhstan" pageId="14" pageNumber="465">Kazakhstan</collectingCountry>
,
<collectingCountry id="F34143D5777B7A63DEA1FA916C275783" box="[934,1044,1407,1428]" name="Kyrgyzstan" pageId="14" pageNumber="465">Kyrgyzstan</collectingCountry>
...................
<taxonomicName id="4C5678C6777B7A63D803FA916DAF5783" ID-CoL="3FPHZ" authorityName="Kashcheev" authorityYear="1999" box="[1284,1436,1407,1428]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macrothorax">
<emphasis id="B922DF57777B7A63D803FA916DAF5783" box="[1284,1436,1407,1428]" italics="true" pageId="14" pageNumber="465">G. macrothorax</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE90345777B7A63DD90FA736DAF57D8" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FA736DAF57D8" pageId="14" pageNumber="465">- Pronotum less convex, without longitudinal depression. Median lobe wide; parameres slightly exceeding apex of median lobe, with four moderately short apical setae.................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A21BE0777B7A63DD90FA396DAF5476" pageId="14" pageNumber="465">
<paragraph id="8BE90345777B7A63DD90FA396DAF541E" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90FA396DAF541E" pageId="14" pageNumber="465">
5 Median lobe very wide, from apical third strongly narrowed apicad; apical portions of parameres moderately narrow (
<figureCitation id="136D1FC0777B7A63D84FFA396911541D" captionStart="FIGURES 31–36" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[169,1412,203,1903]" captionTargetId="figure-31@18.[158,1430,181,1919]" captionTargetPageId="18" captionText="FIGURES 31–36. Aedeagus of Geodromicus convexicollis (Figs. 31–32: Kazakhstan, Kyrgyzsay Pass; Fig. 33: Kazakhstan, Kaskasu; Fig. 34: Kazakhstan, Ketmen Mnt.; Fig. 35: Tajikistan, Kulikulan; Fig. 36: Kyrgyzstan, Terskey-Alatoo Mts.): 31, 33–36—parameral view, 32—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381628" httpUri="https://zenodo.org/record/7381628/files/figure.png" pageId="14" pageNumber="465">Figs. 31, 33–36</figureCitation>
). Body length:
<quantity id="4CAEAEA0777B7A63DCB2FA1A6A70541D" box="[437,579,1524,1546]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="7.2" metricValueMin="4.3" pageId="14" pageNumber="465" unit="mm" value="5.75" valueMax="7.2" valueMin="4.3">4.30–7.20 mm</quantity>
. Body brown to black. Habitus as in
<figureCitation id="136D1FC0777B7A63DEA2FA1A6C23541E" box="[933,1040,1524,1546]" captionStart="FIGURE 38–39" captionStartId="20.[151,250,1974,1998]" captionTargetBox="[159,1414,184,1930]" captionTargetId="figure-31@20.[159,1428,181,1949]" captionTargetPageId="20" captionText="FIGURE 38–39. Habitats of Geodromicus convexicollis in Uzbekistan, Chatkal Mts., Iertash River." figureDoi="http://doi.org/10.5281/zenodo.7381634" httpUri="https://zenodo.org/record/7381634/files/figure.png" pageId="14" pageNumber="465">Fig. 38–39</figureCitation>
. Middle Asia........
<taxonomicName id="4C5678C6777B7A63D805FA1A6DAF541E" ID-CoL="3FPGR" box="[1282,1436,1524,1545]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57777B7A63D805FA1A6DAF541E" box="[1282,1436,1524,1545]" italics="true" pageId="14" pageNumber="465">G. convexicollis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE90345777B7A63DD90F9FC6DAF5476" blockId="14.[151,1437,996,1634]" pageId="14" pageNumber="465">
<keyLead id="30A7A070777B7A63DD90F9FC6DAF5476" pageId="14" pageNumber="465">
- Median lobe moderately narrow, from basal portion gradually narrowed apicad; apical portions of parameres distinctly broadened (
<figureCitation id="136D1FC0777B7A63DDEEF9C1696D5452" box="[233,350,1583,1605]" captionStart="FIGURES 46–50" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[238,1353,187,1907]" captionTargetId="figure-31@26.[219,1369,181,1919]" captionTargetPageId="26" captionText="FIGURES 46–50. Aedeagus of Geodromicus kashmirensis (Fig. 46: Afghanistan, Pagman; Fig. 47: India, Gulmarg; Fig. 48: India, Gya-Ladak; Figs. 49–50: India, Tangmarg Pir Panjal): 46–49—parameral view, 50—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381638" httpUri="https://zenodo.org/record/7381638/files/figure.png" pageId="14" pageNumber="465">Figs. 46–49</figureCitation>
). Body length:
<quantity id="4CAEAEA0777B7A63DCEBF9C16A4A5453" box="[492,633,1583,1605]" metricMagnitude="-3" metricUnit="m" metricValue="5.725" metricValueMax="7.2" metricValueMin="4.25" pageId="14" pageNumber="465" unit="mm" value="5.725" valueMax="7.2" valueMin="4.25">4.25–7.20 mm</quantity>
. Body reddish-brown to dark-brown Habitus as in
<figureCitation id="136D1FC0777B7A63D952F9C16C8D5453" box="[1109,1214,1583,1605]" captionStart="FIGURES 40–45" captionStartId="25.[151,264,1900,1924]" captionTargetBox="[161,1419,567,1866]" captionTargetId="figure-201@25.[151,1436,561,1877]" captionTargetPageId="25" captionText="FIGURES 40–45. Habitus of Geodromicus: 40—G. kashmirensis (lectotype), 41—G. similis (lectoype), 42—G. consors (holotype), 43—G. karakorus (holotype), 44–45—G. lebedevianus (Fig. 44: Kyrgyzstan, Chichkan Vallley; Fig. 45: Kyrgyzstan, Kara-Balta). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381636" httpUri="https://zenodo.org/record/7381636/files/figure.png" pageId="14" pageNumber="465">Fig. 40–43</figureCitation>
.
<collectingCountry id="F34143D5777B7A63D9C2F9C16D0E5452" box="[1221,1341,1583,1605]" name="Afghanistan" pageId="14" pageNumber="465">Afghanistan</collectingCountry>
,
<collectingCountry id="F34143D5777B7A63D842F9C16DAA5452" box="[1349,1433,1583,1605]" name="Pakistan" pageId="14" pageNumber="465">Pakistan</collectingCountry>
,
<collectingCountry id="F34143D5777B7A63DDE5F9A269205475" box="[226,275,1612,1634]" name="India" pageId="14" pageNumber="465">India</collectingCountry>
...................................................................................
<taxonomicName id="4C5678C6777B7A63D805F9A26DAF5476" ID-CoL="3FPHJ" authorityName="Cameron" authorityYear="1930" box="[1282,1436,1612,1633]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="kashmirensis">
<emphasis id="B922DF57777B7A63D805F9A26DAF5476" box="[1282,1436,1612,1633]" italics="true" pageId="14" pageNumber="465">G. kashmirensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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