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<document id="FBD5266BD6FD0CED0958846D2E1651F1" ID-DOI="10.5281/zenodo.5429666" ID-ISSN="0253-116X" ID-Zenodo-Dep="5429666" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630421338040" checkinUser="valdenar" docAuthor="Irikov, A." docDate="2008" docId="03F05744FFA2AE5122915DD88D63FD46" docLanguage="en" docName="LinzerbiolBeitr.2008.40.1.785-811.pdf" docOrigin="Linzer biologische Beiträge 40 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Monacha (Monacha) ocellata" docType="treatment" docVersion="4" lastPageNumber="793" masterDocId="FFC92F3CFFA5AE5922235E6A8C73FFC3" masterDocTitle="Genus Monacha FITZINGER 1833 in Bulgaria (Gastropoda, Pulmonata, Hygromiidae)" masterLastPageNumber="811" masterPageNumber="785" pageNumber="792" updateTime="1699264283420" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7081AA9D4D5D7ED7C5E66D7E3051E97A">Genus Monacha FITZINGER 1833 in Bulgaria (Gastropoda, Pulmonata, Hygromiidae)</mods:title>
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<mods:namePart id="018A2648DD5385F466727C39AE312C8E">Irikov, A.</mods:namePart>
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<mods:title id="178990BDE16FAE7BE5BDAE279F3CED58">Linzer biologische Beiträge</mods:title>
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<mods:date id="3070CEDC7A970B3A96AAA9A7688DFCBB">2008</mods:date>
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<mods:number id="D1ECEEBA6875C3F5A5999FF790D6152F">2008-07-10</mods:number>
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<treatment id="03F05744FFA2AE5122915DD88D63FD46" ID-DOI="http://doi.org/10.5281/zenodo.5485318" ID-Zenodo-Dep="5485318" LSID="urn:lsid:plazi:treatment:03F05744FFA2AE5122915DD88D63FD46" httpUri="http://treatment.plazi.org/id/03F05744FFA2AE5122915DD88D63FD46" lastPageId="8" lastPageNumber="793" pageId="7" pageNumber="792">
<subSubSection id="C343B5D9FFA2AE5E22915DD88F46FC2E" pageId="7" pageNumber="792" type="nomenclature">
<paragraph id="8BE6E652FFA2AE5E22915DD88E04FC0B" blockId="7.[178,631,946,968]" box="[178,631,946,968]" pageId="7" pageNumber="792">
<heading id="D0AE513EFFA2AE5E22915DD88E04FC0B" box="[178,631,946,968]" fontSize="8" level="2" pageId="7" pageNumber="792" reason="2">
<taxonomicName id="4C599DD1FFA2AE5E22915DD88E04FC0B" ID-CoL="43ZHT" authority="(ROTH 1839)" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[178,631,946,968]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="ocellata" subGenus="Monacha">
<emphasis id="B92D3A40FFA2AE5E22915DD88E04FC0B" bold="true" box="[178,631,946,968]" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915DD88D65FC0B" bold="true" box="[178,278,946,968]" italics="true" pageId="7" pageNumber="792">Monacha</emphasis>
(
<emphasis id="B92D3A40FFA2AE5E23065DD88DFAFC0B" bold="true" box="[293,393,946,968]" italics="true" pageId="7" pageNumber="792">Monacha</emphasis>
)
<emphasis id="B92D3A40FFA2AE5E23B45DD88D9BFC0B" bold="true" box="[407,488,946,968]" italics="true" pageId="7" pageNumber="792">ocellata</emphasis>
(ROTH 1839)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE6E652FFA2AE5E22915DB38F46FC2E" blockId="7.[178,892,984,1031]" box="[178,821,984,1005]" pageId="7" pageNumber="792">
1821
<emphasis id="B92D3A40FFA2AE5E22C65DB28D65FC2E" box="[229,278,984,1005]" italics="true" pageId="7" pageNumber="792">Helix</emphasis>
(
<emphasis id="B92D3A40FFA2AE5E23075DB28D04FC2E" box="[292,375,984,1005]" italics="true" pageId="7" pageNumber="792">Helicella</emphasis>
)
<emphasis id="B92D3A40FFA2AE5E23A75DB28DB6FC2E" box="[388,453,984,1005]" italics="true" pageId="7" pageNumber="792">olivieri</emphasis>
FÉRRUSAC: 47 [partim, nomen nudum].
</paragraph>
</subSubSection>
<subSubSection id="C343B5D9FFA2AE5E22915D998F1AFB90" pageId="7" pageNumber="792" type="description">
<paragraph id="8BE6E652FFA2AE5E22915D998F0FFBC4" blockId="7.[178,892,984,1031]" box="[178,892,1010,1031]" pageId="7" pageNumber="792">
1839
<emphasis id="B92D3A40FFA2AE5E22C65D988D2EFBC4" box="[229,349,1010,1031]" italics="true" pageId="7" pageNumber="792">Helix olivieri</emphasis>
<taxonomicName id="4C599DD1FFA2AE5E23405D998E63FBC5" authority="ROTH" authorityName="ROTH" box="[355,528,1010,1031]" pageId="7" pageNumber="792" rank="variety" variety="ocellata">
var.
<emphasis id="B92D3A40FFA2AE5E23AF5D988DA6FBC4" box="[396,469,1010,1031]" italics="true" pageId="7" pageNumber="792">ocellata</emphasis>
ROTH
</taxonomicName>
: 14. Locus typicus: &quot;Byzantii&quot;,
<collectingCountry id="F34EA6C2FFA2AE5E21175D998F0BFBC4" box="[820,888,1011,1031]" name="Turkey" pageId="7" pageNumber="792">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8BE6E652FFA2AE5E22915A728E19FBED" blockId="7.[178,618,1048,1070]" box="[178,618,1048,1070]" pageId="7" pageNumber="792">
Published for
<collectingCountry id="F34EA6C2FFA2AE5E23635A728DE4FBED" box="[320,407,1048,1070]" name="Bulgaria" pageId="7" pageNumber="792">Bulgaria</collectingCountry>
from: present study.
</paragraph>
<paragraph id="8BE6E652FFA2AE5E22915A578F1AFB90" blockId="7.[178,873,1085,1107]" box="[178,873,1085,1107]" pageId="7" pageNumber="792">
The species is registered in
<collectingCountry id="F34EA6C2FFA2AE5E23E85A578E51FB90" box="[459,546,1085,1107]" name="Bulgaria" pageId="7" pageNumber="792">Bulgaria</collectingCountry>
from one locality only (
<figureCitation id="1362FAD7FFA2AE5E21345A578F2EFB90" box="[791,861,1085,1107]" captionStart="Map 4" captionStartId="24.[178,223,1538,1558]" captionTargetBox="[196,1041,926,1508]" captionTargetId="figure-35@24.[178,1055,896,1535]" captionTargetPageId="24" captionText="Map 4. Distribution of Monacha (Monacha) ocellata (ROTH 1839) (UTM-grid, 10 km-squares)." figureDoi="http://doi.org/10.5281/zenodo.5429676" httpUri="https://zenodo.org/record/5429676/files/figure.png" pageId="7" pageNumber="792">Map 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343B5D9FFA2AE5E22915A098EF6FB36" pageId="7" pageNumber="792" type="materials_examined">
<paragraph id="8BE6E652FFA2AE5E22915A098F7CFB70" blockId="7.[178,1086,1123,1203]" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915A098E0DFBBA" bold="true" box="[178,638,1123,1145]" pageId="7" pageNumber="792">
Researched material and locality (
<figureCitation id="1362FAD7FFA2AE5E200F5A098E05FBBA" box="[556,630,1123,1145]" captionStart="Map 4" captionStartId="24.[178,223,1538,1558]" captionTargetBox="[196,1041,926,1508]" captionTargetId="figure-35@24.[178,1055,896,1535]" captionTargetPageId="24" captionText="Map 4. Distribution of Monacha (Monacha) ocellata (ROTH 1839) (UTM-grid, 10 km-squares)." figureDoi="http://doi.org/10.5281/zenodo.5429676" httpUri="https://zenodo.org/record/5429676/files/figure.png" pageId="7" pageNumber="792">Map 4</figureCitation>
)
</emphasis>
: South Black Sea Coast: sand coast Arkutino, near by marsh-reserve Arkutino, between Sozopol and Primorsko (
<collectionCode id="ED487E97FFA2AE5E21845AEA8FBCFB55" box="[935,975,1152,1174]" pageId="7" pageNumber="792">NG</collectionCode>
58),
<quantity id="4CA14BB7FFA2AE5E21DC5AEA884DFB55" box="[1023,1086,1152,1174]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="792" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2-3 m</quantity>
alt.,
<specimenCount id="9D5F2DDBFFA2AE5E22FD5AF78D19FB70" box="[222,362,1181,1203]" pageId="7" pageNumber="792" type="generic">72 specimens</specimenCount>
,
<date id="FFE7C092FFA2AE5E23575AF78D95FB70" box="[372,486,1181,1203]" pageId="7" pageNumber="792" value="2005-12-09">12.09.2005</date>
and
<date id="FFE7C092FFA2AE5E20345AF78EF8FB70" box="[535,651,1181,1203]" pageId="7" pageNumber="792" value="2006-06-23">23.06.2006</date>
, leg.
<collectionCode id="ED487E97FFA2AE5E20E15AF48F7AFB71" box="[706,777,1182,1203]" pageId="7" pageNumber="792">IRIKOV</collectionCode>
.
</paragraph>
<paragraph id="8BE6E652FFA2AE5E22915AA98EF6FB36" blockId="7.[178,1085,1219,1270]" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915AA98D4EFB1A" bold="true" box="[178,317,1219,1241]" pageId="7" pageNumber="792">Shell (Fig. 9)</emphasis>
: Measurements (n = 72), D:
<quantity id="4CA14BB7FFA2AE5E20455AA98E84FB1B" box="[614,759,1219,1241]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" metricValueMax="1.6199999999999999" metricValueMin="1.05" pageId="7" pageNumber="792" unit="mm" value="13.35" valueMax="16.2" valueMin="10.5">10.5-16.2 mm</quantity>
, x =
<quantity id="4CA14BB7FFA2AE5E21085AA98FFAFB1B" box="[811,905,1219,1241]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="792" unit="mm" value="12.4">12.4 mm</quantity>
; H: 7.7-12.0 mm, x = 9.0 mm; D/H =
<quantity id="4CA14BB7FFA2AE5E23585A8A8D81FB36" box="[379,498,1248,1270]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="7" pageNumber="792" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
, x =
<quantity id="4CA14BB7FFA2AE5E20075A8A8EF2FB36" box="[548,641,1248,1270]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="7" pageNumber="792" unit="mm" value="1.38">1.38 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C343B5D9FFA2AE5122915B6F8D63FD46" lastPageId="8" lastPageNumber="793" pageId="7" pageNumber="792" type="description">
<paragraph id="8BE6E652FFA2AE5E22915B6F8D6CFA95" blockId="7.[178,1086,1285,1366]" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915B6F8D1DFAD8" bold="true" box="[178,366,1285,1307]" pageId="7" pageNumber="792">Genitalia (Fig. 3)</emphasis>
: The measurements of some sections of the genitalia are compiled in
<tableCitation id="C6DBD3E9FFA2AE5E22915B488D73FAFB" box="[178,256,1314,1336]" captionStart="Table 2" captionStartId="18.[178,233,800,820]" captionText="Table 2. Measurements of some sections of the genitalia od some Monacha (Monacha) species (in mm). Abbreviations: apa = apical section of the appendicula; apb = basal cection of the appendicula; ep = epiphalus; fl = flagelum; p = penis; vap = vagina up to the base of the appendicula; vgm = vagina up to the glandulae mucosae; vt = total length of the vagina." pageId="7" pageNumber="792">Table 2</tableCitation>
and the variation of the proportions of some sections of the genitalia is represent in
<tableCitation id="C6DBD3E9FFA2AE5E22EF5B2A8D68FA95" box="[204,283,1344,1366]" captionStart="Table 3" captionStartId="19.[178,233,215,235]" captionText="Table 3. Comparison of the ratios of some sections of the genitalia of the Monacha (Monacha) species. Abbreviations;ap = appendicula; apa = apical section of the appendicula; apb = basal cection of the appendicula; ep = epiphalus; fl = flagelum; p = penis; vap = vagina up to the base of the appendicula; vgm = vagina up to the glandulae mucosae; vt = total length of the vagina." pageId="7" pageNumber="792">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BE6E652FFA2AE5E22915B0F8855FAB8" blockId="7.[178,1062,1381,1403]" box="[178,1062,1381,1403]" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915B0F8DE3FAB8" bold="true" box="[178,400,1381,1403]" pageId="7" pageNumber="792">Vertical distribution</emphasis>
: In
<collectingCountry id="F34EA6C2FFA2AE5E239B5B0F8E7CFAB8" box="[440,527,1381,1403]" name="Bulgaria" pageId="7" pageNumber="792">Bulgaria</collectingCountry>
on the Black Sea Coast on several metres of the sea.
</paragraph>
<paragraph id="8BE6E652FFA2AE5122915BE08865FEA5" blockId="7.[178,1086,1418,1587]" lastBlockId="8.[178,1085,219,358]" lastPageId="8" lastPageNumber="793" pageId="7" pageNumber="792">
<emphasis id="B92D3A40FFA2AE5E22915BE08D2AFA63" bold="true" box="[178,345,1418,1440]" pageId="7" pageNumber="792">Ecological data</emphasis>
: The species live on Black Sea Coast at xerothermal ecological conditions in the summer and comparatively mild and humid winter. There in the same habitat occur
<taxonomicName id="4C599DD1FFA2AE5E22D25BAC8D11FA19" baseAuthorityName="MOUSSON" baseAuthorityYear="1863" box="[241,354,1477,1499]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="solidior">
<emphasis id="B92D3A40FFA2AE5E22D25BAC8D11FA19" box="[241,354,1477,1499]" italics="true" pageId="7" pageNumber="792">M. solidior</emphasis>
</taxonomicName>
too, but in difference of her
<taxonomicName id="4C599DD1FFA2AE5E20A55BAC8E89FA19" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[646,762,1477,1499]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="ocellata">
<emphasis id="B92D3A40FFA2AE5E20A55BAC8E89FA19" box="[646,762,1477,1499]" italics="true" pageId="7" pageNumber="792">M. ocellata</emphasis>
</taxonomicName>
inhabit area more distant of the gritty dunes, where the psamophytic grassy vegetation is a quite abundant. Here perennial plants almost entirely cover the gritty coast. In difference of
<taxonomicName id="4C599DD1FFA2AE5E21A1586A8F80F9D6" baseAuthorityName="MOUSSON" baseAuthorityYear="1863" box="[898,1011,1536,1557]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="solidior">
<emphasis id="B92D3A40FFA2AE5E21A1586A8F80F9D6" box="[898,1011,1536,1557]" italics="true" pageId="7" pageNumber="792">M. solidior</emphasis>
</taxonomicName>
, which rarely may be found to be active on the plants in summertime (see below
<taxonomicName id="4C599DD1FFA2AE5E21B85877887FF9F1" baseAuthorityName="MOUSSON" baseAuthorityYear="1863" box="[923,1036,1565,1586]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="solidior">
<emphasis id="B92D3A40FFA2AE5E21B85877887FF9F1" box="[923,1036,1565,1586]" italics="true" pageId="7" pageNumber="792">M. solidior</emphasis>
</taxonomicName>
),
<taxonomicName id="4C599DD1FFA2AE51260158778D70FF33" baseAuthorityName="ROTH" baseAuthorityYear="1839" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" lastPageId="8" lastPageNumber="793" order="Stylommatophora" pageId="7" pageNumber="792" phylum="Mollusca" rank="species" species="ocellata">
<emphasis id="B92D3A40FFA2AE51260158778D70FF33" italics="true" lastPageId="8" lastPageNumber="793" pageId="7" pageNumber="792">M. ocellata</emphasis>
</taxonomicName>
make huge aggregates among the psamophytic vegetation and in determinate periods the animals are active. In the same locality
<taxonomicName id="4C599DD1FFADAE5121215E938F0DFECE" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[770,894,248,270]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="793" phylum="Mollusca" rank="species" species="ocellata">
<emphasis id="B92D3A40FFADAE5121215E938F0DFECE" box="[770,894,248,270]" italics="true" pageId="8" pageNumber="793">M. ocellata</emphasis>
</taxonomicName>
form the largest aggregates on artificial planted introduced plants of Genus
<emphasis id="B92D3A40FFADAE51213F5F7C8F00FEE8" box="[796,883,278,299]" italics="true" pageId="8" pageNumber="793">Opuncia</emphasis>
sp. These
<emphasis id="B92D3A40FFADAE5121C55F7C884EFEE8" box="[998,1085,278,299]" italics="true" pageId="8" pageNumber="793">Opuncia</emphasis>
occurs in large numbers in nearly located island &quot;Zmiyski Ostrov&quot;, where are introduced in the beginning of XX century, and subsequently are carried on the Black Sea Coast.
</paragraph>
<paragraph id="8BE6E652FFADAE5122915F1C886EFDDE" blockId="8.[178,1086,373,541]" pageId="8" pageNumber="793">
We don’t find
<taxonomicName id="4C599DD1FFADAE51236E5F1C8DB0FE49" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[333,451,373,395]" class="Gastropoda" family="Hygromiidae" genus="Monacha" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="793" phylum="Mollusca" rank="species" species="ocellata">
<emphasis id="B92D3A40FFADAE51236E5F1C8DB0FE49" box="[333,451,373,395]" italics="true" pageId="8" pageNumber="793">M. ocellata</emphasis>
</taxonomicName>
nowhere elsewhere in the southern Black Sea Coast. We assume that it is possible the species was carried there from the southern areas together with the plants of Genus
<emphasis id="B92D3A40FFADAE51239E5FDA8E67FE06" box="[445,532,432,453]" italics="true" pageId="8" pageNumber="793">Opuncia</emphasis>
. In the same locality already exist one introduced in the Bulgarian malacofauna species –
<emphasis id="B92D3A40FFADAE51201D5FA78E86FE20" box="[574,757,461,483]" italics="true" pageId="8" pageNumber="793">Cochlicella acuta</emphasis>
, reported at first by HUDEC &amp; VASATKO (1973). The manner of introduction is still vague and probably transferring was made accidentally, but the species is survivor and formed small stable population.
</paragraph>
<paragraph id="8BE6E652FFADAE5122915C478EC8FD80" blockId="8.[178,699,557,579]" box="[178,699,557,579]" pageId="8" pageNumber="793">Xerophilic, thermophilic, drought-resistant species.</paragraph>
<paragraph id="8BE6E652FFADAE5122915C388D63FD46" blockId="8.[178,1086,594,645]" pageId="8" pageNumber="793">
<emphasis id="B92D3A40FFADAE5122915C388D3DFDAB" bold="true" box="[178,334,594,616]" pageId="8" pageNumber="793">Zoogeography</emphasis>
:
<collectingCountry id="F34EA6C2FFADAE5123475C388D92FDAB" box="[356,481,594,616]" name="Turkey" pageId="8" pageNumber="793">Asia Minor</collectingCountry>
subelement, Subiranian element, Southwestern Asiatic complex.
</paragraph>
</subSubSection>
</treatment>
</document>
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