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168 lines
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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin & Hamer, Michelle" docDate="2013" docId="039587C7FFB1FFAF66B9FBC962D2770D" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Harpagophora : Attems 1909" docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="285" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vohland, Katrin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hamer, Michelle</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>54</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>251</mods:start>
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<mods:end>251</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:039587C7FFB1FFAF66B9FBC962D2770D" httpUri="http://treatment.plazi.org/id/039587C7FFB1FFAF66B9FBC962D2770D" lastPageId="35" lastPageNumber="286" pageId="34" pageNumber="285">
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<subSubSection box="[422,825,1098,1124]" pageId="34" pageNumber="285" type="nomenclature">
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<paragraph blockId="34.[422,825,1098,1124]" box="[422,825,1098,1124]" pageId="34" pageNumber="285">
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<heading box="[422,825,1098,1124]" centered="true" fontSize="36" level="2" pageId="34" pageNumber="285" reason="6">
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Genus
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<taxonomicName ID-CoL="7PJHK" authority="Attems, 1909" authorityName=": Attems" authorityYear="1909" box="[501,825,1098,1124]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
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<emphasis box="[501,669,1098,1124]" italics="true" pageId="34" pageNumber="285">Harpagophora</emphasis>
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<bibRefCitation author="ATTEMS, C. G." box="[674,825,1098,1124]" pageId="34" pageNumber="285" pagination="1 - 52" refId="ref23416" refString="ATTEMS, C. G. 1909 a. Myriapoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1901 - 1903 von Dr. Leonhard Schultze. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="285" type="reference_group">
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<paragraph blockId="34.[144,1103,1138,1185]" pageId="34" pageNumber="285">
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<treatmentCitationGroup pageId="34" pageNumber="285">
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<treatmentCitation author="ATTEMS, C. G." box="[144,445,1139,1160]" page="41" pageId="34" pageNumber="285" year="1909">
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<taxonomicName ID-CoL="7PJHK" authority=": Attems 1909 a: 41" authorityName=": Attems" authorityPageNumber="41" authorityYear="1909" box="[144,445,1139,1160]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
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<emphasis box="[144,277,1139,1160]" italics="true" pageId="34" pageNumber="285">Harpagophora</emphasis>
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:
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<bibRefCitation author="ATTEMS, C. G." box="[286,445,1139,1160]" pageId="34" pageNumber="285" pagination="1 - 52" refId="ref23416" refString="ATTEMS, C. G. 1909 a. Myriapoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1901 - 1903 von Dr. Leonhard Schultze. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">
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Attems 1909
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<emphasis box="[398,409,1139,1160]" italics="true" pageId="34" pageNumber="285">a</emphasis>
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: 41
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</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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; 1914
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<emphasis box="[496,507,1139,1160]" italics="true" pageId="34" pageNumber="285">a</emphasis>
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: 162; 1928:372; Lawrence 1965: 50; Schubart 1966: 166; Demange 1983: 579; Hamer 1998: 54; 1999: 13; Redman
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<emphasis box="[675,722,1163,1184]" italics="true" pageId="34" pageNumber="285">et al.</emphasis>
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2003: 210.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="35" lastPageNumber="286" pageId="34" pageNumber="285" type="materials_examined">
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<paragraph blockId="34.[144,1092,1202,1228]" box="[144,1092,1202,1228]" pageId="34" pageNumber="285">
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<typeStatus box="[144,200,1202,1228]" pageId="34" pageNumber="285">Type</typeStatus>
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species:
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<taxonomicName authority="Attems, 1909" authorityName=": Attems" authorityYear="1909" box="[302,761,1202,1228]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="diplocrada">
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<emphasis box="[302,600,1202,1228]" italics="true" pageId="34" pageNumber="285">Harpagophora diplocrada</emphasis>
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<bibRefCitation author="ATTEMS, C. G." box="[607,761,1202,1228]" pageId="34" pageNumber="285" pagination="1 - 52" refId="ref23416" refString="ATTEMS, C. G. 1909 a. Myriapoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1901 - 1903 von Dr. Leonhard Schultze. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909</bibRefCitation>
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</taxonomicName>
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, designated by Jeekel (1970).
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</paragraph>
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<paragraph blockId="34.[144,1104,1244,1367]" pageId="34" pageNumber="285">
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Diagnosis: Two femoral processes on the telopodite, these unequal in length and width (
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<figureCitation box="[151,238,1276,1302]" captionStart="Figs 63, 64" captionStartId="36.[144,183,1343,1365]" captionTargetBox="[257,1031,212,1328]" captionTargetId="figure-108@36.[155,1094,204,1338]" captionTargetPageId="36" captionText="Figs 63, 64. Harpagophora diplocrada Attems, ♂: (63) telopodite of gonopod, lateral view; (64) gonopod, oral view.Abbreviations: (fp1) femoral process 1, (fp2) femoral process 2, (pe) pectinophore, (sb) spine branch. Scale bar = 1 mm." pageId="34" pageNumber="285">Figs 64</figureCitation>
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,
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<figureCitation box="[249,279,1276,1303]" captionStart="Figs 65–70" captionStartId="37.[181,220,1534,1556]" captionTargetBox="[193,1116,490,1513]" captionTargetId="figure-187@37.[181,1141,469,1516]" captionTargetPageId="37" captionText="Figs 65–70.Harpagophora aridaRedman, ♂: (65)telopodite,oral view;(66) telocoxite,oral view;(67) gnathochilarium; (68) head and first segments, lateral view; (69)epiproct, lateral view; (70) leg 2 ofmale. Abbreviations:(as) additionalsmall spine on telopodite,(c) cardo,(col) collum,(cx) coxa,(fp1) femoral process 1, (fp2) femoral process 2, (g) gonocoel, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of gonocoxite, (m) mentum, (ml) median lamella of gonocoxite, (p) penis, (pb) prebasilare, (pe) pectinophore, (sb) spine branch, (st) stipes, (t) telocoxite projection. Scale bar = 1 mm." pageId="34" pageNumber="285">65</figureCitation>
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,
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<emphasis box="[290,375,1276,1302]" italics="true" pageId="34" pageNumber="285">fp1, fp2</emphasis>
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); posterior limb of telopodite comprising two branches, the spine branch (
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<figureCitation box="[235,317,1308,1334]" captionStart="Figs 63, 64" captionStartId="36.[144,183,1343,1365]" captionTargetBox="[257,1031,212,1328]" captionTargetId="figure-108@36.[155,1094,204,1338]" captionTargetPageId="36" captionText="Figs 63, 64. Harpagophora diplocrada Attems, ♂: (63) telopodite of gonopod, lateral view; (64) gonopod, oral view.Abbreviations: (fp1) femoral process 1, (fp2) femoral process 2, (pe) pectinophore, (sb) spine branch. Scale bar = 1 mm." pageId="34" pageNumber="285">Fig. 64</figureCitation>
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,
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<emphasis box="[329,354,1308,1334]" italics="true" pageId="34" pageNumber="285">sb</emphasis>
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) and the pectinophore (
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<figureCitation box="[619,701,1308,1334]" captionStart="Figs 63, 64" captionStartId="36.[144,183,1343,1365]" captionTargetBox="[257,1031,212,1328]" captionTargetId="figure-108@36.[155,1094,204,1338]" captionTargetPageId="36" captionText="Figs 63, 64. Harpagophora diplocrada Attems, ♂: (63) telopodite of gonopod, lateral view; (64) gonopod, oral view.Abbreviations: (fp1) femoral process 1, (fp2) femoral process 2, (pe) pectinophore, (sb) spine branch. Scale bar = 1 mm." pageId="34" pageNumber="285">Fig. 64</figureCitation>
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,
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<emphasis box="[713,739,1309,1334]" italics="true" pageId="34" pageNumber="285">pe</emphasis>
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). Syncoxosternite of first pair of legs with suture or divided.
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</paragraph>
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<paragraph blockId="34.[144,1104,1383,1442]" pageId="34" pageNumber="285">
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Distribution: Four species occur in
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<collectingCountry box="[549,694,1383,1409]" name="South Africa" pageId="34" pageNumber="285">South Africa</collectingCountry>
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(
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<collectingRegion box="[709,869,1383,1409]" country="South Africa" name="Western Cape" pageId="34" pageNumber="285">Western Cape</collectingRegion>
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and
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<collectingRegion box="[927,1096,1383,1409]" country="South Africa" name="Northern Cape" pageId="34" pageNumber="285">Northern Cape</collectingRegion>
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) and two in
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<collectingCountry box="[269,369,1415,1441]" name="Namibia" pageId="34" pageNumber="285">Namibia</collectingCountry>
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.
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</paragraph>
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<paragraph blockId="34.[144,1104,1458,1676]" pageId="34" pageNumber="285">
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Remarks:
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<taxonomicName authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[257,843,1458,1484]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="spirobolinus" subGenus="Nodopyge">
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<emphasis box="[257,687,1458,1484]" italics="true" pageId="34" pageNumber="285">Spirostreptus (Nodopyge) spirobolinus</emphasis>
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Karsch, 1881
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</taxonomicName>
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, was the first
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<taxonomicName pageId="34" pageNumber="285">
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<emphasis italics="true" pageId="34" pageNumber="285">Harpagophora</emphasis>
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species
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</taxonomicName>
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to be named but the description was based on a female, and only later was a male described from Hantam in the
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<collectingRegion box="[630,798,1522,1548]" country="South Africa" name="Northern Cape" pageId="34" pageNumber="285">Northern Cape</collectingRegion>
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by Attems (1914
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<emphasis box="[997,1011,1523,1548]" italics="true" pageId="34" pageNumber="285">a</emphasis>
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). While
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<taxonomicName authorityName="Karsch" authorityYear="1881" box="[144,283,1554,1580]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="spirobolinus">
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<emphasis box="[144,283,1554,1580]" italics="true" pageId="34" pageNumber="285">spirobolinus</emphasis>
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</taxonomicName>
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was placed in
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<taxonomicName authorityName=": Attems" authorityYear="1909" box="[448,616,1554,1580]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
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<emphasis box="[448,616,1554,1580]" italics="true" pageId="34" pageNumber="285">Harpagophora</emphasis>
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</taxonomicName>
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by Attems (1914
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<emphasis box="[808,822,1555,1580]" italics="true" pageId="34" pageNumber="285">a</emphasis>
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, 1928), Demange (1983) followed Jeekel (1970) in allocating
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<taxonomicName authorityName=": Attems" authorityYear="1909" box="[565,724,1586,1612]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="diplocrada">
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<emphasis box="[565,724,1586,1612]" italics="true" pageId="34" pageNumber="285">H. diplocrada</emphasis>
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</taxonomicName>
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as the
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<typeStatus box="[809,857,1586,1612]" pageId="34" pageNumber="285">type</typeStatus>
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species of the genus. Redman
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<emphasis box="[241,286,1618,1644]" italics="true" pageId="34" pageNumber="285">et al</emphasis>
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. (2003) subsequently created a new monotypic genus,
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<taxonomicName authority="Redman, 2003" authorityName="Redman" authorityYear="2003" class="Diplopoda" family="Harpagophoridae" genus="Metaphora" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
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<emphasis box="[877,1000,1618,1644]" italics="true" pageId="34" pageNumber="285">Metaphora</emphasis>
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Redman, 2003
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</taxonomicName>
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, for
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<emphasis box="[253,381,1650,1676]" italics="true" pageId="34" pageNumber="285">spirobolina</emphasis>
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.
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</paragraph>
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<paragraph blockId="35.[181,1142,204,358]" pageId="35" pageNumber="286">
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Demange (1983) divided
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<taxonomicName authorityName=": Attems" authorityYear="1909" box="[489,657,204,230]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="35" pageNumber="286" phylum="Arthropoda" rank="genus">
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<emphasis box="[489,657,204,230]" italics="true" pageId="35" pageNumber="286">Harpagophora</emphasis>
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</taxonomicName>
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into two subgenera on the basis of the structure of the margin of the spine branch. Redman
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<emphasis box="[692,738,236,262]" italics="true" pageId="35" pageNumber="286">et al</emphasis>
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. (2003), however, suggested that this character does not show any relationship in its occurrence to the other major taxonomic characters in the genus, including the form of the femoral process and telocoxites, and so the two subgenera were synonymised.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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