treatments-xml/data/03/95/87/039587B4FFB3FFBBFF5CF899DD29FB6C.xml

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<document id="73F201555C05C2CC6A5B240F6D7F389F" ID-CLB-Dataset="56069" ID-DOI="10.11646/zootaxa.5125.3.5" ID-GBIF-Dataset="da274829-479f-4ea1-8f2e-1e598de786b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6443876" ID-ZooBank="4645A035-17E6-488C-8F10-DA66E146487D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649661642485" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2022" docId="039587B4FFB3FFBBFF5CF899DD29FB6C" docLanguage="en" docName="zootaxa.5125.3.5.pdf" docOrigin="Zootaxa 5125 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudomicracis pennata" docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FFACFFCCFFBEFFABFFCBFFAED841FF85" masterDocTitle="The strongly dimorphic bark beetle genus Pseudomicracis (Coleoptera, Scolytinae) in Madagascar-an integrated taxonomic revision" masterLastPageNumber="343" masterPageNumber="325" pageNumber="338" updateTime="1699319771094" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5AAD7419112424409ADC4374C96C90BC">The strongly dimorphic bark beetle genus Pseudomicracis (Coleoptera, Scolytinae) in Madagascar-an integrated taxonomic revision</mods:title>
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<mods:namePart id="F94429F4B4C57172E321828C39FE529F">Jordal, Bjarte H.</mods:namePart>
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<treatment id="039587B4FFB3FFBBFF5CF899DD29FB6C" ID-DOI="http://doi.org/10.5281/zenodo.6448646" ID-GBIF-Taxon="194565926" ID-Zenodo-Dep="6448646" LSID="urn:lsid:plazi:treatment:039587B4FFB3FFBBFF5CF899DD29FB6C" httpUri="http://treatment.plazi.org/id/039587B4FFB3FFBBFF5CF899DD29FB6C" lastPageId="16" lastPageNumber="341" pageId="13" pageNumber="338">
<subSubSection id="C3266529FFB3FFA6FF5CF899DA37F8D4" box="[151,630,1847,1874]" pageId="13" pageNumber="338" type="nomenclature">
<paragraph id="8B8336A2FFB3FFA6FF5CF899DA37F8D4" blockId="13.[151,630,1847,1910]" box="[151,630,1847,1874]" pageId="13" pageNumber="338">
<heading id="D0CB81CEFFB3FFA6FF5CF899DA37F8D4" bold="true" box="[151,630,1847,1874]" fontSize="11" level="1" pageId="13" pageNumber="338" reason="1">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CF899DA37F8D4" authority="(Schedl, 1965)" baseAuthorityName="Schedl" baseAuthorityYear="1965" box="[151,630,1847,1874]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="pennata">
<emphasis id="B948EAB0FFB3FFA6FF5CF899DA37F8D4" bold="true" box="[151,630,1847,1874]" pageId="13" pageNumber="338">
<emphasis id="B948EAB0FFB3FFA6FF5CF899D9FEF8D4" bold="true" box="[151,447,1847,1874]" italics="true" pageId="13" pageNumber="338">Pseudomicracis pennata</emphasis>
(
<bibRefCitation id="EFAD4B53FFB3FFA6FE05F899DA2FF8D4" author="Schedl, K. E." box="[462,622,1847,1873]" pageId="13" pageNumber="338" pagination="51 - 85" refId="ref10566" refString="Schedl, K. E. (1965) Fauna Madagascariensis, VI. 232 Beitrag. Reichenbachia, Dresden, 5, 51 - 85." type="journal article" year="1965">Schedl, 1965</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF5CF8F5D98EF8F3" box="[151,463,1883,1910]" pageId="13" pageNumber="338" type="description">
<paragraph id="8B8336A2FFB3FFA6FF5CF8F5D98EF8F3" blockId="13.[151,630,1847,1910]" box="[151,463,1883,1910]" pageId="13" pageNumber="338">
(
<figureCitation id="13072A27FFB3FFA6FF54F8F5D90AF8F0" box="[159,331,1883,1909]" captionStart="FIGURES 3644" captionStartId="14.[151,264,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-25@14.[151,1436,181,1890]" captionTargetPageId="14" captionText="FIGURES 3644. Dorsal, lateral and front view of Pseudomicracis tomicoides male holotype (36, 39, 43); Pseudomicracis dispar male (37, 40, 43); Pseudomicracis pennata male holotype (38, 41, 44)." figureDoi="http://doi.org/10.5281/zenodo.6443900" httpUri="https://zenodo.org/record/6443900/files/figure.png" pageId="13" pageNumber="338">Figs 38, 41, 44</figureCitation>
,
<figureCitation id="13072A27FFB3FFA6FE9DF8F5D986F8F3" box="[342,455,1883,1910]" captionStart="FIGURES 4553" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[178,1418,185,1919]" captionTargetId="figure-15@15.[170,1418,181,1919]" captionTargetPageId="15" captionText="FIGURES 4553. Dorsal, lateral and front view of Pseudomicracis tomicoides female allotype (45, 48, 51); Pseudomicracis dispar female (46, 49, 52); Pseudomicracis pennata female allotype (47, 50, 53)." figureDoi="http://doi.org/10.5281/zenodo.6443904" httpUri="https://zenodo.org/record/6443904/files/figure.png" pageId="13" pageNumber="338">47, 50, 53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF5CF80EDB2CF853" pageId="13" pageNumber="338" type="reference_group">
<paragraph id="8B8336A2FFB3FFA6FF5CF80EDAD6F83D" blockId="13.[151,877,1951,2006]" box="[151,663,1951,1976]" pageId="13" pageNumber="338">
<treatmentCitationGroup id="AB2C118CFFB3FFA6FF5CF80EDAD6F83D" box="[151,663,1951,1976]" pageId="13" pageNumber="338">
<treatmentCitation id="0A9D10B3FFB3FFA6FF5CF80EDA47F832" author="Schedl, K. E." box="[151,518,1951,1976]" page="63" pageId="13" pageNumber="338" year="1965">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CF80EDA47F832" ID-CoL="42PXM" authority="Schedl, 1965: 63" authorityName="Schedl" authorityPageNumber="63" authorityYear="1965" box="[151,518,1951,1976]" class="Insecta" family="Curculionidae" genus="Micracis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="pennatus">
<emphasis id="B948EAB0FFB3FFA6FF5CF80ED912F832" box="[151,339,1952,1975]" italics="true" pageId="13" pageNumber="338">Micracis pennatus</emphasis>
<bibRefCitation id="EFAD4B53FFB3FFA6FE92F80EDA47F832" author="Schedl, K. E." box="[345,518,1951,1976]" pageId="13" pageNumber="338" pagination="51 - 85" refId="ref10566" refString="Schedl, K. E. (1965) Fauna Madagascariensis, VI. 232 Beitrag. Reichenbachia, Dresden, 5, 51 - 85." type="journal article" year="1965">Schedl, 1965: 63</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, orig. spelling
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8336A2FFB3FFA6FF5CF810DB2CF853" blockId="13.[151,877,1951,2006]" box="[151,877,1982,2006]" pageId="13" pageNumber="338">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CF810DA78F853" ID-CoL="78G5S" authority="(Schedl, 1965)" baseAuthorityName="Schedl" baseAuthorityYear="1965" box="[151,569,1982,2006]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="pennatus">
<emphasis id="B948EAB0FFB3FFA6FF5CF810D9DBF853" box="[151,410,1982,2006]" italics="true" pageId="13" pageNumber="338">Pseudomicracis pennatus</emphasis>
(
<bibRefCitation id="EFAD4B53FFB3FFA6FE63F810DA70F853" author="Schedl, K. E." box="[424,561,1982,2006]" pageId="13" pageNumber="338" pagination="51 - 85" refId="ref10566" refString="Schedl, K. E. (1965) Fauna Madagascariensis, VI. 232 Beitrag. Reichenbachia, Dresden, 5, 51 - 85." type="journal article" year="1965">Schedl, 1965</bibRefCitation>
)
</taxonomicName>
, combination by
<bibRefCitation id="EFAD4B53FFB3FFA6FD2CF810DB2CF853" author="Wood, S. L." box="[743,877,1982,2006]" pageId="13" pageNumber="338" pagination="223 - 230" refId="ref10686" refString="Wood, S. L. (1984) New generic synonomy and new genera of Scolytidae (Coleoptera). Great Basin Naturalist, 44, 223 - 230." type="journal article" year="1984">Wood (1984)</bibRefCitation>
</paragraph>
</subSubSection>
<caption id="DF43662AFFB0FFA5FF5CF8D4DBECF833" ID-DOI="http://doi.org/10.5281/zenodo.6443900" ID-Zenodo-Dep="6443900" httpUri="https://zenodo.org/record/6443900/files/figure.png" pageId="14" pageNumber="339" startId="14.[151,264,1914,1938]" targetBox="[152,1436,181,1890]" targetPageId="14">
<paragraph id="8B8336A2FFB0FFA5FF5CF8D4DBECF833" blockId="14.[151,1436,1914,1974]" pageId="14" pageNumber="339">
<emphasis id="B948EAB0FFB0FFA5FF5CF8D4D917F817" bold="true" box="[151,342,1914,1938]" pageId="14" pageNumber="339">FIGURES 3644.</emphasis>
Dorsal, lateral and front view of
<taxonomicName id="4C3C4D21FFB0FFA5FD7DF8D4DB8FF817" baseAuthorityName="Schedl" baseAuthorityYear="1961" box="[694,974,1914,1938]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB0FFA5FD7DF8D4DB8FF817" box="[694,974,1914,1938]" italics="true" pageId="14" pageNumber="339">Pseudomicracis tomicoides</emphasis>
</taxonomicName>
male holotype (36, 39, 43);
<taxonomicName id="4C3C4D21FFB0FFA5FB31F8D4D899F833" baseAuthorityName="Schedl" baseAuthorityYear="1961" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B948EAB0FFB0FFA5FB31F8D4D899F833" italics="true" pageId="14" pageNumber="339">Pseudomicracis dispar</emphasis>
</taxonomicName>
male (37, 40, 43);
<taxonomicName id="4C3C4D21FFB0FFA5FE52F830DAD5F833" baseAuthorityName="Schedl" baseAuthorityYear="1965" box="[409,660,1950,1974]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="339" phylum="Arthropoda" rank="species" species="pennata">
<emphasis id="B948EAB0FFB0FFA5FE52F830DAD5F833" box="[409,660,1950,1974]" italics="true" pageId="14" pageNumber="339">Pseudomicracis pennata</emphasis>
</taxonomicName>
male holotype (38, 41, 44).
</paragraph>
</caption>
<caption id="DF43662AFFB1FFA4FF5CF839DB8CF856" ID-DOI="http://doi.org/10.5281/zenodo.6443904" ID-Zenodo-Dep="6443904" httpUri="https://zenodo.org/record/6443904/files/figure.png" pageId="15" pageNumber="340" startId="15.[151,264,1943,1967]" targetBox="[178,1418,185,1919]" targetPageId="15">
<paragraph id="8B8336A2FFB1FFA4FF5CF839DB8CF856" blockId="15.[151,1436,1943,2004]" pageId="15" pageNumber="340">
<emphasis id="B948EAB0FFB1FFA4FF5CF839D914F82A" bold="true" box="[151,341,1943,1967]" pageId="15" pageNumber="340">FIGURES 4553.</emphasis>
Dorsal, lateral and front view of
<taxonomicName id="4C3C4D21FFB1FFA4FD64F836DB87F835" baseAuthorityName="Schedl" baseAuthorityYear="1961" box="[687,966,1944,1968]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB1FFA4FD64F836DB87F835" box="[687,966,1944,1968]" italics="true" pageId="15" pageNumber="340">Pseudomicracis tomicoides</emphasis>
</taxonomicName>
female allotype (45, 48, 51);
<taxonomicName id="4C3C4D21FFB1FFA4FB31F836D899F851" baseAuthorityName="Schedl" baseAuthorityYear="1961" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B948EAB0FFB1FFA4FB31F836D899F851" italics="true" pageId="15" pageNumber="340">Pseudomicracis dispar</emphasis>
</taxonomicName>
female (46, 49, 52);
<taxonomicName id="4C3C4D21FFB1FFA4FE66F812DAE6F856" baseAuthorityName="Schedl" baseAuthorityYear="1965" box="[429,679,1980,2004]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="340" phylum="Arthropoda" rank="species" species="pennata">
<emphasis id="B948EAB0FFB1FFA4FE66F812DAE6F856" box="[429,679,1980,2004]" italics="true" pageId="15" pageNumber="340">Pseudomicracis pennata</emphasis>
</taxonomicName>
female allotype (47, 50, 53).
</paragraph>
</caption>
<subSubSection id="C3266529FFAEFFBBFF5CFF38DA4AFF50" pageId="16" pageNumber="341" type="materials_examined">
<paragraph id="8B8336A2FFAEFFBBFF5CFF38DA4AFF50" blockId="16.[151,1437,150,428]" pageId="16" pageNumber="341">
<emphasis id="B948EAB0FFAEFFBBFF5CFF38D904FF35" bold="true" box="[151,325,150,176]" pageId="16" pageNumber="341">Type material.</emphasis>
<materialsCitation id="3B543CFFFFAEFFBBFE87FF38DCA4FF34" ID-GBIF-Occurrence="3750353326" box="[332,1253,150,177]" collectionCode="MNHN" country="Madagascar" location="Sandrangato" pageId="16" pageNumber="341" specimenCount="1" typeStatus="holotype">
<typeStatus id="54878800FFAEFFBBFE87FF38D9F6FF35" box="[332,439,150,176]" pageId="16" pageNumber="341" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="F32B7632FFAEFFBBFDC3FF38DAD0FF35" box="[520,657,150,176]" name="Madagascar" pageId="16" pageNumber="341">Madagascar</collectingCountry>
,
<location id="8EE36079FFAEFFBBFD57FF38DB69FF35" LSID="urn:lsid:plazi:treatment:039587B4FFB3FFBBFF5CF899DD29FB6C:8EE36079FFAEFFBBFD57FF38DB69FF35" box="[668,808,150,176]" country="Madagascar" name="Sandrangato" pageId="16" pageNumber="341">Sandrangato</location>
[likely south of Moramanga] [
<collectionCode id="ED2DAE67FFAEFFBBFB4BFF38DC9EFF34" box="[1152,1247,150,177]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="341">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B543CFFFFAEFFBBFB25FF38DA46FF50" ID-GBIF-Occurrence="3750353328" collectionCode="NHMW" country="Madagascar" location="Sandrangato" pageId="16" pageNumber="341" specimenCount="1" typeStatus="allotype">
<typeStatus id="54878800FFAEFFBBFB25FF38DD12FF35" box="[1262,1363,150,176]" pageId="16" pageNumber="341" type="allotype">Allotype</typeStatus>
, male: same data as holotype [
<collectionCode id="ED2DAE67FFAEFFBBFE50FF14DA41FF50" box="[411,512,186,213]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="341" type="Museum">NHMW</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFAEFFBBFF0CFF70DD3DFEC5" pageId="16" pageNumber="341" type="diagnosis">
<paragraph id="8B8336A2FFAEFFBBFF0CFF70DD3DFEC5" blockId="16.[151,1437,150,428]" pageId="16" pageNumber="341">
<emphasis id="B948EAB0FFAEFFBBFF0CFF70D903FF7D" bold="true" box="[199,322,222,248]" pageId="16" pageNumber="341">Diagnosis.</emphasis>
Length
<quantity id="4CC49B47FFAEFFBBFE68FF70DA6EFF7C" box="[419,559,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="16" pageNumber="341" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
, 2.2 × as long as wide.
<emphasis id="B948EAB0FFAEFFBBFC8EFF70DBC3FF7D" bold="true" box="[837,898,222,248]" pageId="16" pageNumber="341">Male</emphasis>
declivity on interstriae 45 with a tall, narrow ridge, extending the total length of elytra by one-quarter; interstrial setae short, spatulate or truncated, densely placed, strongly confused.
<emphasis id="B948EAB0FFAEFFBBFE0FFE88DA5AFEC5" bold="true" box="[452,539,294,320]" pageId="16" pageNumber="341">Female</emphasis>
with rounded declivity, interstrial setae conical, in part confused on declivity
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFAEFFBBFF0CFEE4DB2DFEE1" box="[199,876,330,357]" pageId="16" pageNumber="341" type="distribution">
<paragraph id="8B8336A2FFAEFFBBFF0CFEE4DB2DFEE1" blockId="16.[151,1437,150,428]" box="[199,876,330,357]" pageId="16" pageNumber="341">
<emphasis id="B948EAB0FFAEFFBBFF0CFEE4D9B0FEE1" bold="true" box="[199,497,330,356]" pageId="16" pageNumber="341">Distribution and biology.</emphasis>
<collectingCountry id="F32B7632FFAEFFBBFE33FEE4DAC0FEE1" box="[504,641,330,356]" name="Madagascar" pageId="16" pageNumber="341">Madagascar</collectingCountry>
. Biology not known.
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFAEFFBBFF0CFEC0DD29FB6C" pageId="16" pageNumber="341" type="discussion">
<paragraph id="8B8336A2FFAEFFBBFF0CFEC0D960FE29" blockId="16.[151,1437,150,428]" pageId="16" pageNumber="341">
<emphasis id="B948EAB0FFAEFFBBFF0CFEC0D97AFE0D" bold="true" box="[199,315,366,392]" pageId="16" pageNumber="341">Remarks.</emphasis>
Gender of genus is feminine (
<bibRefCitation id="EFAD4B53FFAEFFBBFD57FEC0DBBCFE0D" author="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C." box="[668,1021,366,392]" pageId="16" pageNumber="341" pagination="1 - 134" refId="ref9733" refString="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C. (2009) A catalogue of family and genus group names in Scolytinae and Platypodinae with nomenclatural remarks (Coleoptera: Curculionidae). Zootaxa, 2258 (1), 1 - 134. https: // doi. org / 10.11646 / zootaxa. 2258.1.1" type="journal article" year="2009">Alonso-Zarazaga &amp; Lyal 2009</bibRefCitation>
) and the species epithet is modified accordingly.
</paragraph>
<paragraph id="8B8336A2FFAEFFBBFF5CFE50DBC9FD9D" blockId="16.[151,904,510,536]" box="[151,904,510,536]" pageId="16" pageNumber="341">
<emphasis id="B948EAB0FFAEFFBBFF5CFE50D9EBFD9D" bold="true" box="[151,426,510,536]" pageId="16" pageNumber="341">Sexual dimorphism in</emphasis>
<taxonomicName id="4C3C4D21FFAEFFBBFE79FE50DA2CFD9D" authorityName="Eggers" authorityYear="1920" box="[434,621,510,536]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAB0FFAEFFBBFE79FE50DA2CFD9D" box="[434,621,510,536]" italics="true" pageId="16" pageNumber="341">Pseudomicracis</emphasis>
</taxonomicName>
<emphasis id="B948EAB0FFAEFFBBFDBEFE50DBC9FD9D" bold="true" box="[629,904,510,536]" pageId="16" pageNumber="341">and other Micracidini</emphasis>
</paragraph>
<paragraph id="8B8336A2FFAEFFBBFF5CFDE8DD0FFCBD" blockId="16.[151,1437,582,1257]" pageId="16" pageNumber="341">
Almost all Micracidini species have some sort of sexual dimorphism in the head where females often have longer setae on a larger antennal scapus, and a variously modified frons that may be concave and textured, or flattened to lightly bulbous and shiny, with or without dense setae. It is presumed that this is due to females being the second arriving sex to a tunnel system, and that the male will control access (
<bibRefCitation id="EFAD4B53FFAEFFBBFC7AFD1CDCF0FD48" author="Kirkendall, L. R. &amp; Biedermann, P. H. W. &amp; Jordal, B. H." box="[945,1201,690,717]" pageId="16" pageNumber="341" pagination="85 - 156" refId="ref10298" refString="Kirkendall, L. R., Biedermann, P. H. W. &amp; Jordal, B. H. (2015) Diversity and evolution of bark beetles. In: Vega, F. &amp; Hofstetter, R. (Eds.), Bark beetles: biology and ecology of native and invasive species. Elsevier, London, pp. 85 - 156. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 417156 - 5.00003 - 4" type="book chapter" year="2015">
Kirkendall
<emphasis id="B948EAB0FFAEFFBBFBF8FD1DDC2DFD49" box="[1075,1132,690,716]" italics="true" pageId="16" pageNumber="341">et al.</emphasis>
2015
</bibRefCitation>
). Females have been observed to court males in the tunnel opening and apparently the tactile stimuli provided by the fluffy antennal and frontal apparatus is important for male acceptance. Typically a male will accept exactly
<specimenCount id="9D3AFD2BFFAEFFBBFBBBFD55DCB9FC91" box="[1136,1272,762,789]" pageId="16" pageNumber="341" type="female">two females</specimenCount>
to their tunnel entrance, although this is an average condition with many exceptions given the resource availability on site.
</paragraph>
<paragraph id="8B8336A2FFAEFFBBFF0CFCECDD29FB6C" blockId="16.[151,1437,582,1257]" pageId="16" pageNumber="341">
<taxonomicName id="4C3C4D21FFAEFFBBFF0CFCECD93BFCD9" authorityName="Eggers" authorityYear="1920" box="[199,378,834,860]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAB0FFAEFFBBFF0CFCECD93BFCD9" box="[199,378,834,860]" italics="true" pageId="16" pageNumber="341">Pseudomicracis</emphasis>
</taxonomicName>
is special among the Micracidini in that males often differ from the females not only in head structures, but also in the shape of the pronotum and the declivity. Some species have males with a distinct row of teeth along the anterior margin of the pronotum, a trait which occasionally appear in other micracidine taxa (
<bibRefCitation id="EFAD4B53FFAEFFBBFA9DFC24D8A0FC4D" author="Jordal, B. H." pageId="16" pageNumber="341" pagination="245 - 278" refId="ref10239" refString="Jordal, B. H. (2021 c) A phylogenetic and taxonomic assessment of Afrotropical Micracidini (Coleoptera, Scolytinae) reveals a strong diversifying role for Madagascar. Organisms Diversity &amp; Evolution, 21, 245 - 278. https: // doi. org / 10.1007 / s 13127 - 021 - 00481 - 4" type="journal article" year="2021">Jordal 2021c</bibRefCitation>
). Almost all
<taxonomicName id="4C3C4D21FFAEFFBBFEB3FC00DA6AFC4D" authorityName="Eggers" authorityYear="1920" box="[376,555,942,968]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAB0FFAEFFBBFEB3FC00DA6AFC4D" box="[376,555,942,968]" italics="true" pageId="16" pageNumber="341">Pseudomicracis</emphasis>
</taxonomicName>
have males with lightly to quite strongly granulated declivity and posterior elytral disc. Additionally, the interstrial setae are broader and truncated, compared to the narrowly spatulate setae in females. The most exceptional features are seen in species in the
<taxonomicName id="4C3C4D21FFAEFFBBFC5FFC59DC4AFB94" authorityName="Wood" authorityYear="1984" box="[916,1035,1015,1041]" class="Insecta" family="Curculionidae" genus="Saurotocis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAB0FFAEFFBBFC5FFC59DC4AFB94" box="[916,1035,1015,1041]" italics="true" pageId="16" pageNumber="341">Saurotocis</emphasis>
</taxonomicName>
group which have males with huge tubercles along the outer lateral part of the declivity and the overall elytral vestiture is dense and confused. Molecular analyses (
<figureCitation id="13072A27FFAEFFBBFEE6FB90D92EFBDD" box="[301,367,1086,1112]" captionStart="FIGURE 1" captionStartId="3.[151,250,1880,1904]" captionTargetId="figure-139@3.[215,1364,448,1676]" captionTargetPageId="3" captionText="FIGURE 1. Maximum likelihood tree topology based on nucleotides from five gene fragments. Bootstrap node support above nodes. Parsimony bootsrap support below (if different)." figureDoi="http://doi.org/10.5281/zenodo.6448648" httpUri="https://zenodo.org/record/6448648/files/figure.png" pageId="16" pageNumber="341">Fig. 1</figureCitation>
) nevertheless show that these features are derived within the typical
<taxonomicName id="4C3C4D21FFAEFFBBFB42FB90DD7DFBDD" authorityName="Eggers" authorityYear="1920" box="[1161,1340,1086,1112]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAB0FFAEFFBBFB42FB90DD7DFBDD" box="[1161,1340,1086,1112]" italics="true" pageId="16" pageNumber="341">Pseudomicracis</emphasis>
</taxonomicName>
lineage, which is also supported by female morphology in these species which is typical for the genus (
<figureCitation id="13072A27FFAEFFBBFB13FBCCDD1CFBF8" box="[1240,1373,1122,1149]" captionStart="FIGURES 4553" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[178,1418,185,1919]" captionTargetId="figure-15@15.[170,1418,181,1919]" captionTargetPageId="15" captionText="FIGURES 4553. Dorsal, lateral and front view of Pseudomicracis tomicoides female allotype (45, 48, 51); Pseudomicracis dispar female (46, 49, 52); Pseudomicracis pennata female allotype (47, 50, 53)." figureDoi="http://doi.org/10.5281/zenodo.6443904" httpUri="https://zenodo.org/record/6443904/files/figure.png" pageId="16" pageNumber="341">Figs 4553</figureCitation>
). The male inclination for declivital evolution is also seen in the peculiar
<taxonomicName id="4C3C4D21FFAEFFBBFC4DFB29DC51FB25" authority="Jordal, 2022" authorityName="Jordal" authorityYear="2022" box="[902,1040,1159,1184]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="341" phylum="Arthropoda" rank="species" species="coronatus" status="sp. nov.">
<emphasis id="B948EAB0FFAEFFBBFC4DFB29DC51FB25" box="[902,1040,1159,1184]" italics="true" pageId="16" pageNumber="341">P. coronatus</emphasis>
</taxonomicName>
<emphasis id="B948EAB0FFAEFFBBFBDCFB28DC31FB25" bold="true" box="[1047,1136,1158,1184]" pageId="16" pageNumber="341">
<taxonomicNameLabel id="A27B57CBFFAEFFBBFBDCFB28DC31FB25" box="[1047,1136,1158,1184]" pageId="16" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
which also has males with a granulated demarcation of the declivity. It is not clear what the purpose of rough declivities might be for the males, but similar traits are found throughout the subfamily and may even appear in females (
<bibRefCitation id="EFAD4B53FFAEFFBBFBABFB60DD1DFB6C" author="Kirkendall, L. R. &amp; Biedermann, P. H. W. &amp; Jordal, B. H." box="[1120,1372,1230,1257]" pageId="16" pageNumber="341" pagination="85 - 156" refId="ref10298" refString="Kirkendall, L. R., Biedermann, P. H. W. &amp; Jordal, B. H. (2015) Diversity and evolution of bark beetles. In: Vega, F. &amp; Hofstetter, R. (Eds.), Bark beetles: biology and ecology of native and invasive species. Elsevier, London, pp. 85 - 156. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 417156 - 5.00003 - 4" type="book chapter" year="2015">
Kirkendall
<emphasis id="B948EAB0FFAEFFBBFB2BFB61DD58FB6D" box="[1248,1305,1230,1256]" italics="true" pageId="16" pageNumber="341">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>