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304 lines
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<document id="E5F90E5D0CA0E680F51AFDB589E30F60" ID-DOI="10.5281/zenodo.276675" ID-GBIF-Dataset="7ba68779-c4ac-4f7d-989d-dd24a9e033d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="276675" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227523469" checkinUser="plazi" docAuthor="Molineri, Carlos, Cruz, Paulo Vilela & Emmerich, Daniel" docDate="2011" docId="03942C161E79FFC4FF370E910309E18B" docLanguage="en" docName="zt02750p038.pdf" docOrigin="Zootaxa 2750" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Asthenopus crenulatus Molineri, Cruz & Emmerich, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFAD546E1E78FFC0FFA00F17024FE560" masterDocTitle="A new species of Asthenopus (Ephemeroptera: Polymitarcyidae: Asthenopodinae) from Brazil and Colombia" masterLastPageNumber="38" masterPageNumber="33" pageNumber="34" updateTime="1698238683989" updateUser="plazi">
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<mods:title id="3B9EABB611DC07407247B83CFD5ED097">A new species of Asthenopus (Ephemeroptera: Polymitarcyidae: Asthenopodinae) from Brazil and Colombia</mods:title>
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<mods:namePart id="A92A113FB4E76AA343DFDDCE5A1573CB">Molineri, Carlos</mods:namePart>
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<mods:namePart id="05035640F367B9CC4E9D8EAE9DC3E04A">Cruz, Paulo Vilela</mods:namePart>
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<mods:namePart id="F9EC6B8885D488A65EDFEBC2DD30144D">Emmerich, Daniel</mods:namePart>
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<mods:title id="9D1D1C22EBDDDFE9944977157509C0F2">Zootaxa</mods:title>
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<mods:date id="908F0C8C85B28FF3ECE8DF73906FDDC9">2011</mods:date>
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<mods:number id="66E042EB6790E9FA09FBD0854DAAA348">2750</mods:number>
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<mods:classification id="80253120D588B8CF23CCB376CBE9864A">journal article</mods:classification>
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<mods:identifier id="C58D35BA0173BEC295F8E76080965C31" type="DOI">10.5281/zenodo.276675</mods:identifier>
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<treatment id="03942C161E79FFC4FF370E910309E18B" ID-DOI="http://doi.org/10.5281/zenodo.6183725" ID-GBIF-Taxon="119386777" ID-Zenodo-Dep="6183725" LSID="urn:lsid:plazi:treatment:03942C161E79FFC4FF370E910309E18B" httpUri="http://treatment.plazi.org/id/03942C161E79FFC4FF370E910309E18B" lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34">
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<subSubSection id="C327CE8B1E79FFC1FF370E910388E0ED" pageId="1" pageNumber="34" type="nomenclature">
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<paragraph id="8B829D001E79FFC1FF370E91005EE4C0" blockId="1.[151,529,390,416]" box="[151,529,390,416]" pageId="1" pageNumber="34">
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<heading id="D0CA2A6C1E79FFC1FF370E91005EE4C0" bold="true" box="[151,529,390,416]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
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<emphasis id="B94941121E79FFC1FF370E91005EE4C0" bold="true" box="[151,529,390,416]" pageId="1" pageNumber="34">
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<taxonomicName id="4C3DE6831E79FFC1FF370E9103E2E4C0" ID-CoL="5VW9X" box="[151,429,390,416]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
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<emphasis id="B94941121E79FFC1FF370E9103E2E4C0" bold="true" box="[151,429,390,416]" italics="true" pageId="1" pageNumber="34">Asthenopus crenulatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27AFC691E79FFC1FE140E91005EE4C0" box="[436,529,390,416]" pageId="1" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B829D001E79FFC1FF370EDA0388E0ED" blockId="1.[151,1437,461,2037]" pageId="1" pageNumber="34">
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Male imago (
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<figureCitation id="130681851E79FFC1FE8E0EDA03D4E485" box="[302,411,461,485]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 1–7</figureCitation>
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). Length (mm): body, 7.0–7.8; fore wing, 7.5–8.6; hind wing, 3.1–3.7; fore leg (from coxa to apex of claw), 6.2–6.9; cerci, 21.6. General coloration yellowish white. Head (
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<figureCitation id="130681851E79FFC1FBEE0EE606F4E769" box="[1102,1211,497,521]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 1–2</figureCitation>
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) brownish dorsally except small pale median mark on hind margin, tinged black at base of antennae and on a triangle between ocelli; ventrally much paler, with blackish markings. Antennae yellowish shaded slightly with gray on apex of scape and pedicel, flagellum hyaline; scape and pedicel long and slender, scape (
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<quantity id="4CC530E51E79FFC1FC100D4A0658E715" box="[944,1047,605,629]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="34" unit="mm" value="0.23">0.23 mm</quantity>
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) slightly longer than pedicel (
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<quantity id="4CC530E51E79FFC1FAC90D4A0288E7F9" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="1" pageNumber="34" unit="mm" value="0.18">0.18 mm</quantity>
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), flagellum long, reaching hind margin of pronotum. Thorax (
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<figureCitation id="130681851E79FFC1FCDC0D9601A7E7F9" box="[892,1000,641,665]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 1–2</figureCitation>
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). Pronotum and propleurae translucent yellowish completely shaded with brown, darker on anterior ring, and lateral and hind margins of posterior ring; a narrow pale sinuous strip separates anterior and posterior rings; presternum pale. Mesonotum yellowish shaded gray on sutures mainly between posteroscutal protuberances; mesopleurae and sternum slightly paler with gray shading on anterior half. Metanotum yellowish shaded gray medially, pleurae and sternum paler, translucent. Legs. Forelegs similar in length to body, 0.7–0.9 times the length of the forewing; coxa, trochanter and femur yellowish brown, rest translucent yellowish, shaded gray almost completely but lighter apically. Length (mm) of foreleg segments (mean of two males): femur (0.79), tibia (1.59), tarsite 1 (0.1), tarsite 2 (1.22), tarsite 3 (0.92), tarsite 4 (0.81), tarsite 5 (0.66), longer claw (0.38), shorter claw (0.22). Middle and hind legs with long coxae and trochanters; femora, tibiae and tarsi very reduced and membranous; both legs yellowish, shaded with gray mainly on a basal spot of hind coxa and distal mark on hind femur. Wings (
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<figureCitation id="130681851E79FFC1FCFB0CFE0186E161" box="[859,969,1001,1025]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 3–4</figureCitation>
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). Membrane hyaline except on costal and subcostal sector of fore and hind wings, grayish, sometimes whitish apical half; all veins yellowish brown; short marginal intercalaries (mi in
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<figureCitation id="130681851E79FFC1FE490B260061E129" box="[489,558,1073,1097]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 3</figureCitation>
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) attached along entire hind margin of both wings. Three to four crossveins between R and M sectors basally to R stem (arrows in
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<figureCitation id="130681851E79FFC1FCA80B420103E10D" box="[776,844,1109,1133]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 3</figureCitation>
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). Ratio Rs length from fork to margin (B in
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<figureCitation id="130681851E79FFC1FAE90B4207C5E10D" box="[1353,1418,1109,1133]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 3</figureCitation>
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)/ stem length (A in
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<figureCitation id="130681851E79FFC1FEC10B6E03ECE1F1" box="[353,419,1145,1169]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 3</figureCitation>
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) = 4. Ratio MA length from fork to margin/stem length, range 9 to 11. Abdomen (
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<figureCitation id="130681851E79FFC1FAE10B6E02E9E1D5" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 1– 2</figureCitation>
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) whitish translucent shaded very slightly with brownish gray on dorsum, sterna paler; tergum X shaded stronger on three longitudinal marks (one median, two sublateral) and margins. Genitalia (
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<figureCitation id="130681851E79FFC1FB870BD606DAE1B9" box="[1063,1173,1217,1241]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 5–7</figureCitation>
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): forceps whitish translucent, penes yellowish; pedestals (p in
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<figureCitation id="130681851E79FFC1FDF60BF200D8E19D" box="[598,663,1253,1277]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 5</figureCitation>
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) relatively short, well separated from each other by a subrectangular median plate (mp in
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<figureCitation id="130681851E79FFC1FE230A1E0387E041" box="[387,456,1289,1313]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 5</figureCitation>
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); forceps relatively long and slender, with setae on inner margin increasing in number distally, apex curved medially (
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<figureCitation id="130681851E79FFC1FE5F0A3A000AE025" box="[511,581,1325,1349]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Fig. 5</figureCitation>
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); base of penes well developed, penean arms cylindrical, curved medially, and with many spines subapically on outer margin (
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<figureCitation id="130681851E79FFC1FD590A460125E009" box="[761,874,1361,1385]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="1" pageNumber="34">Figs. 6–7</figureCitation>
|
|||
|
|
). Cerci whitish; terminal filament very reduced, straight, with 6 thin annuli.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C327CE8B1E79FFC1FF670A8E03E5E2CC" pageId="1" pageNumber="34" type="description">
|
|||
|
|
<paragraph id="8B829D001E79FFC1FF670A8E034AE220" blockId="1.[151,1437,461,2037]" pageId="1" pageNumber="34">
|
|||
|
|
Female subimago (
|
|||
|
|
<figureCitation id="130681851E79FFC1FE3C0A8E0058E0D1" box="[412,535,1433,1457]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Figs. 8–13</figureCitation>
|
|||
|
|
). Length (mm): body, 10.2–10.8; fore wing, 11.1; hind wing, 4.3. General coloration yellowish light brown. Head (
|
|||
|
|
<figureCitation id="130681851E79FFC1FD890AAA00DBE0B5" box="[553,660,1469,1493]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Figs. 8–9</figureCitation>
|
|||
|
|
) dorsally blackish except on median zone, paler; venter of head yellowish white. Antennae light yellowish, shaded gray on apex of scape and pedicel. Thorax (
|
|||
|
|
<figureCitation id="130681851E79FFC1FB120AF60750E099" box="[1202,1311,1505,1529]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Figs. 8–9</figureCitation>
|
|||
|
|
) yellowish brown with blackish membranes, shaded with brownish gray on pronotum and with black on posteromedian marks on meso- and metanotum. Legs whitish yellow shaded brownish on dorsum. Wings (
|
|||
|
|
<figureCitation id="130681851E79FFC1FB21093E0742E321" box="[1153,1293,1577,1601]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Figs. 10–11</figureCitation>
|
|||
|
|
), membrane slightly tinged with light brown, veins brownish; costal and subcostal sector of fore and hind wings, grayish, sometimes apical half of costal and subcostal sector of forewing whitish; forewing with four longitudinal intercalary veins between R2 and R3. Abdomen (
|
|||
|
|
<figureCitation id="130681851E79FFC1FD94098200EEE3CD" box="[564,673,1685,1709]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Figs. 8–9</figureCitation>
|
|||
|
|
). Terga yellowish shaded with brownish gray; sterna whitish yellow with anteromedial longitudinal brownish line and two reddish brown spots medially; sternum VIII (
|
|||
|
|
<figureCitation id="130681851E79FFC1FAA709AB0716E3B4" box="[1287,1369,1724,1748]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Fig. 12</figureCitation>
|
|||
|
|
) with paired anteromedian sockets, sockets small, shallow and contiguous (so in
|
|||
|
|
<figureCitation id="130681851E79FFC1FC5E09F7061BE398" box="[1022,1108,1760,1784]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="1" pageNumber="34">Fig. 13</figureCitation>
|
|||
|
|
). Terminal filament whitish, shorter than tergum VIII, with 6 thin annuli; cerci yellowish brown paler apically, 0.5–0.6 times the length of the abdomen.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B829D001E79FFC1FF67085A064EE2E8" blockId="1.[151,1437,461,2037]" pageId="1" pageNumber="34">Eggs. Length, 221–266 μm; width, 143–152μm. Slightly oval; yellow with two polar caps on apices, each formed by a long coiled filament; no chorion microsculptures visible at 100X.</paragraph>
|
|||
|
|
<paragraph id="8B829D001E79FFC1FF67088303E5E2CC" blockId="1.[151,1437,461,2037]" box="[199,426,1940,1964]" pageId="1" pageNumber="34">Nymphs. Unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C327CE8B1E79FFC1FF6708AF06F8E295" pageId="1" pageNumber="34" type="discussion">
|
|||
|
|
<paragraph id="8B829D001E79FFC1FF6708AF06F8E295" blockId="1.[151,1437,461,2037]" pageId="1" pageNumber="34">
|
|||
|
|
<emphasis id="B94941121E79FFC1FF6708AF0374E2B0" bold="true" box="[199,315,1976,2000]" pageId="1" pageNumber="34">Remarks.</emphasis>
|
|||
|
|
The differences between male imago and subimago are limited to length of forelegs, forceps and cerci. The penes of imago and subimago are similar, allowing the identification in both stages.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF42CD881E7AFFC2FF37084F01A6E2CC" httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="2" pageNumber="35" targetBox="[156,1431,200,1851]" targetPageId="2">
|
|||
|
|
<paragraph id="8B829D001E7AFFC2FF37084F01A6E2CC" blockId="2.[151,1436,1880,1964]" pageId="2" pageNumber="35">
|
|||
|
|
<emphasis id="B94941121E7AFFC2FF37084F0373E20E" bold="true" box="[151,316,1880,1903]" pageId="2" pageNumber="35">FIGURES 1–7.</emphasis>
|
|||
|
|
<taxonomicName id="4C3DE6831E7AFFC2FEE4084F0064E20E" box="[324,555,1880,1902]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
|
|||
|
|
<emphasis id="B94941121E7AFFC2FEE4084F0064E20E" box="[324,555,1880,1902]" italics="true" pageId="2" pageNumber="35">Asthenopus crenulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A27AFC691E7AFFC2FD93084E00CAE20E" box="[563,645,1881,1902]" pageId="2" pageNumber="35" rank="species">
|
|||
|
|
<emphasis id="B94941121E7AFFC2FD93084E00CAE20E" bold="true" box="[563,645,1881,1902]" pageId="2" pageNumber="35">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Male imago. 1, body, d.v. (= dorsal view); 2, body, v.v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v.v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF42CD881E7BFFC3FF37086C03CCE2B6" httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="3" pageNumber="36" targetBox="[158,1429,199,1887]" targetPageId="3">
|
|||
|
|
<paragraph id="8B829D001E7BFFC3FF37086C03CCE2B6" blockId="3.[151,1436,1915,2006]" pageId="3" pageNumber="36">
|
|||
|
|
<emphasis id="B94941121E7BFFC3FF37086C0308E2F1" bold="true" box="[151,327,1915,1938]" pageId="3" pageNumber="36">FIGURES 8–13.</emphasis>
|
|||
|
|
<taxonomicName id="4C3DE6831E7BFFC3FEEC086B007EE2F2" box="[332,561,1916,1938]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
|
|||
|
|
<emphasis id="B94941121E7BFFC3FEEC086B007EE2F2" box="[332,561,1916,1938]" italics="true" pageId="3" pageNumber="36">Asthenopus crenulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A27AFC691E7BFFC3FD96086B00C9E2F1" box="[566,646,1916,1937]" pageId="3" pageNumber="36" rank="species">
|
|||
|
|
<emphasis id="B94941121E7BFFC3FD96086B00C9E2F1" bold="true" box="[566,646,1916,1937]" pageId="3" pageNumber="36">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Female subimago. 8, body, d.v. (= dorsal view); 9, body, v.v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII–IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d.v.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C327CE8B1E7CFFC4FF670F8F0356E5B4" pageId="4" pageNumber="37" type="etymology">
|
|||
|
|
<paragraph id="8B829D001E7CFFC4FF670F8F0356E5B4" blockId="4.[151,1437,151,1259]" pageId="4" pageNumber="37">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FF670F8F0301E5D0" bold="true" box="[199,334,152,176]" pageId="4" pageNumber="37">Etymology.</emphasis>
|
|||
|
|
From Latin
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FE400F8F001AE5D0" box="[480,597,152,176]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="crenulatus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FE400F8F001AE5D0" box="[480,597,152,176]" italics="true" pageId="4" pageNumber="37">crenulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
meaning "minutely crenated", because of the remarkable penile structure (
|
|||
|
|
<figureCitation id="130681851E7CFFC4FF3F0FAB0342E5B4" box="[159,269,188,212]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="4" pageNumber="37">Figs. 5–7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C327CE8B1E7CFFC4FF670FC80181E497" pageId="4" pageNumber="37" type="diagnosis">
|
|||
|
|
<paragraph id="8B829D001E7CFFC4FF670FC80181E497" blockId="4.[151,1437,151,1259]" pageId="4" pageNumber="37">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FF670FC803B7E598" bold="true" box="[199,504,223,248]" pageId="4" pageNumber="37">Diagnosis and discussion.</emphasis>
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FE5F0FF700B1E598" box="[511,766,224,248]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FE5F0FF700B1E598" box="[511,766,224,248]" italics="true" pageId="4" pageNumber="37">Asthenopus crenulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A27AFC691E7CFFC4FCA60FF7012FE598" box="[774,864,224,248]" pageId="4" pageNumber="37" rank="species">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FCA60FF7012FE598" bold="true" box="[774,864,224,248]" pageId="4" pageNumber="37">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
can be distinguished from the other species of the genus by the following combination of characters: 1) male foreleg with large and apically widening tibia, first tarsite partially fused to second tarsite, foreleg length 0.7–0.9 times the length of forewing; 2) marginal intercalary veins present on the entire margin of fore and hind wings (
|
|||
|
|
<figureCitation id="130681851E7CFFC4FCF70E5B0188E404" box="[855,967,332,356]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="4" pageNumber="37">Figs. 3–4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="130681851E7CFFC4FC770E5B0650E404" box="[983,1055,332,356]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="4" pageNumber="37">10–11</figureCitation>
|
|||
|
|
), generally shorter than distance between longitudinal veins in male (
|
|||
|
|
<figureCitation id="130681851E7CFFC4FD900E7800D3E4E7" box="[560,668,367,391]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="4" pageNumber="37">Figs. 3–4</figureCitation>
|
|||
|
|
), but longer and anastomosed in female (
|
|||
|
|
<figureCitation id="130681851E7CFFC4FBCC0E7806BBE4E7" box="[1132,1268,367,391]" captionStart="FIGURES 8 – 13" captionStartId="3.[151,264,1915,1938]" captionTargetBox="[158,1429,199,1887]" captionTargetId="figure@3.[151,1436,193,1894]" captionTargetPageId="3" captionText="FIGURES 8 – 13. Asthenopus crenulatus sp. nov. Female subimago. 8, body, d. v. (= dorsal view); 9, body, v. v. (= ventral view); 10, forewing; 11, hind wing; 12, abdominal sterna VIII – IX; 13, detail of sockets. Abbreviations: so = sockets; s VIII, s IX = sterna VIII and IX. d. v." httpUri="https://zenodo.org/record/276677/files/figure.png" pageId="4" pageNumber="37">Figs. 10–11</figureCitation>
|
|||
|
|
); 3) basal relation of forewing veins IMP-MP1 variable (IMP joined to MP1, or basally free); 4) forewing with 3 to 4 crossveins between Rs and MA basal to Rs fork; 5) ratio total length/basal width of forceps 8.5–9.0 (tl/bw in
|
|||
|
|
<figureCitation id="130681851E7CFFC4FB440EAD0768E4B2" box="[1252,1319,442,466]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="4" pageNumber="37">Fig. 5</figureCitation>
|
|||
|
|
); 6) penes long and slender, outer apical margin with many small spines (
|
|||
|
|
<figureCitation id="130681851E7CFFC4FCF40EC8018EE497" box="[852,961,479,503]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[156,1431,200,1851]" captionTargetId="figure@2.[151,1436,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Asthenopus crenulatus sp. nov. Male imago. 1, body, d. v. (= dorsal view); 2, body, v. v. (= ventral view); 3, forewing; 4, hind wing; 5, genitalia, v. v.; 6, penes; 7, detail of penes. Abbreviations: bw = basal width; mi = marginal intercalaries; mp = median plate of styliger; p = pedestal; s IX = sternum IX; tl = total length." httpUri="https://zenodo.org/record/276676/files/figure.png" pageId="4" pageNumber="37">Figs. 6–7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C327CE8B1E7CFFC4FF670D140376E6E2" pageId="4" pageNumber="37" type="reference_group">
|
|||
|
|
<paragraph id="8B829D001E7CFFC4FF670D14064DE7AF" blockId="4.[151,1437,151,1259]" pageId="4" pageNumber="37">
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FF670D14038BE77B" ID-CoL="5VW9X" box="[199,452,515,539]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="crenulatus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FF670D14038BE77B" box="[199,452,515,539]" italics="true" pageId="4" pageNumber="37">Asthenopus crenulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is more closely related with
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FCA70D1401A8E77A" ID-CoL="67VXX" authority="Hubbard" authorityName="Hubbard" box="[775,999,514,539]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="picteti">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FCA70D140123E77B" box="[775,876,515,539]" italics="true" pageId="4" pageNumber="37">A. picteti</emphasis>
|
|||
|
|
(Hubbard)
|
|||
|
|
</taxonomicName>
|
|||
|
|
than to other species of the genus. The characters shared by both species include the relatively slender forceps and penes and the relatively long male forelegs. Both species can be easily distinguished by the larger size (forewing length,
|
|||
|
|
<quantity id="4CC530E51E7CFFC4FB820D5D063BE702" box="[1058,1140,586,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="37" unit="mm" value="12.0">12 mm</quantity>
|
|||
|
|
), and absence of subapical penile spines in
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FEED0D7803FDE7E7" ID-CoL="67VXX" box="[333,434,623,647]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="picteti">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FEED0D7803FDE7E7" box="[333,434,623,647]" italics="true" pageId="4" pageNumber="37">A. picteti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Furthermore, the extremely specialized foretibial-tarsal joint and apically enlarged foretarsal claws of
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FEE00D840007E7CA" ID-CoL="67VXX" authority="Traver 1956" authorityName="Traver" authorityYear="1956" box="[320,584,658,683]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="picteti">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FEE00D8403E9E7CB" box="[320,422,659,683]" italics="true" pageId="4" pageNumber="37">A. picteti</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFACE0F11E7CFFC4FE140D85000FE7CA" author="Traver" box="[436,576,658,682]" pageId="4" pageNumber="38" refString="Traver, J. R. (1956) The genus Asthenopodes (Ephemeroptera). Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 75, 1 - 15." type="journal article" year="1956">Traver 1956</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
are absent in the new species, that shows the common states for these characters (tarsite 1 not fused with tibia, and claws widening slightly on the apex).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B829D001E7CFFC4FF670DCD0376E6E2" blockId="4.[151,1437,151,1259]" pageId="4" pageNumber="37">
|
|||
|
|
Female adults are only known for
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FDE40DCC0103E792" ID-CoL="5W7Y9" authority="Berner 1978" authorityName="Berner" authorityYear="1978" box="[580,844,730,755]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="curtus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FDE40DCC00E7E793" box="[580,680,731,755]" italics="true" pageId="4" pageNumber="37">A. curtus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFACE0F11E7CFFC4FD160DCD010BE792" author="Berner" box="[694,836,730,754]" pageId="4" pageNumber="38" refString="Berner, L. (1978) The status of Asthenopus curtus (Hagen) (Ephemeroptera: Polymitarcyidae). Acta Amazonica, 8, 103 - 105." type="journal article" year="1978">Berner 1978</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and an undetermined species from
|
|||
|
|
<collectingCountry id="F32ADD901E7CFFC4FB770DCC0776E793" box="[1239,1337,731,755]" name="Uruguay" pageId="4" pageNumber="37">Uruguay</collectingCountry>
|
|||
|
|
("
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FAF40DCC02BAE677" class="Insecta" family="Polymitarcyidae" genus="Asthenopodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FAF40DCC02BAE677" italics="true" pageId="4" pageNumber="37">Asthenopodes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp." in
|
|||
|
|
<bibRefCitation id="EFACE0F11E7CFFC4FEF20C170391E677" author="Traver" box="[338,478,767,791]" pageId="4" pageNumber="38" refString="Traver, J. R. (1956) The genus Asthenopodes (Ephemeroptera). Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 75, 1 - 15." type="journal article" year="1956">Traver 1956</bibRefCitation>
|
|||
|
|
) but sockets on the VIII sternum were not known to occur in the genus until now. These structures were studied by
|
|||
|
|
<bibRefCitation id="EFACE0F11E7CFFC4FD8E0C350187E65A" author="McCafferty" box="[558,968,802,826]" pageId="4" pageNumber="38" refString="McCafferty, W. P. & Bloodgood, D. W. (1989) The female and male coupling apparatus in Tortopus mayflies. Aquatic Insects, 11, 141 - 146." type="journal article" year="1989">McCafferty and Bloodgood (1989)</bibRefCitation>
|
|||
|
|
in a species of
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FB370C3306B3E65B" ID-CoL="7XX2" box="[1175,1276,804,827]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FB370C3306B3E65B" box="[1175,1276,804,827]" italics="true" pageId="4" pageNumber="37">Tortopsis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and recently described for females of
|
|||
|
|
<taxonomicName id="4C3DE6831E7CFFC4FE0C0C5F0082E63F" ID-CoL="7XX3" authority="Molineri 2010" authorityName="Molineri" authorityYear="2010" box="[428,717,839,863]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FE0C0C5F0040E63F" box="[428,527,840,863]" italics="true" pageId="4" pageNumber="37">Tortopus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFACE0F11E7CFFC4FDBD0C50008AE63F" author="Molineri" box="[541,709,839,863]" pageId="4" pageNumber="38" refString="Molineri, C. (2010) A cladistic revision of Tortopus Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae). Zootaxa, 2481, 1 - 36." type="journal article" year="2010">Molineri 2010</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
. They are supposed to function as part of the coupling apparatus during copula.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C327CE8B1E7CFFC4FF670C990309E18B" pageId="4" pageNumber="37" type="materials_examined">
|
|||
|
|
<paragraph id="8B829D001E7CFFC4FF670C990309E18B" blockId="4.[151,1437,151,1259]" pageId="4" pageNumber="37">
|
|||
|
|
<emphasis id="B94941121E7CFFC4FF670C990379E6C7" bold="true" box="[199,310,910,935]" pageId="4" pageNumber="37">Material.</emphasis>
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FE9D0C9803EAE6C7" box="[317,421,911,935]" pageId="4" pageNumber="37" type="holotype">Holotype</typeStatus>
|
|||
|
|
male from
|
|||
|
|
<collectingCountry id="F32ADD901E7CFFC4FD890C9800DDE6C7" box="[553,658,911,935]" name="Brazil" pageId="4" pageNumber="37">BRAZIL</collectingCountry>
|
|||
|
|
: AM 240, Ramal Paulo Freire, Sitio os Pioneiros, Km-05, Presidente Figueiredo, Amazonas,
|
|||
|
|
<quantity id="4CC530E51E7CFFC4FE010CA50393E6AA" box="[417,476,946,970]" metricMagnitude="1" metricUnit="m" metricValue="9.4" pageId="4" pageNumber="37" unit="m" value="94.0">94 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FF83BBC01E7CFFC4FE470CA5002FE6AA" box="[487,608,946,970]" pageId="4" pageNumber="37" value="2008-04-24">24.iv.2008</date>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FDCA0CA5009CE6AA" box="[618,723,946,970]" direction="south" orientation="latitude" pageId="4" pageNumber="37" precision="15" value="-2.0680554">S 2° 4' 5"</geoCoordinate>
|
|||
|
|
–
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FD4E0CA401D3E6AA" box="[750,924,946,970]" direction="west" orientation="longitude" pageId="4" pageNumber="37" precision="1" value="-59.908">W 59° 54'28.8"</geoCoordinate>
|
|||
|
|
, light trap, N. Hamada col..
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FB760CA40708E6AB" box="[1238,1351,947,971]" pageId="4" pageNumber="37" type="paratype">Paratypes</typeStatus>
|
|||
|
|
:
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FAF10CA507D4E6AA" box="[1361,1435,946,970]" pageId="4" pageNumber="37" type="male">1 male</specimenCount>
|
|||
|
|
imago and
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FEB40CC00310E68F" box="[276,351,983,1007]" pageId="4" pageNumber="37" type="male">3 male</specimenCount>
|
|||
|
|
subimagos same data as
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FDDC0CC00091E68F" box="[636,734,983,1007]" pageId="4" pageNumber="37" type="holotype">holotype</typeStatus>
|
|||
|
|
;
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FD540CC00170E68F" box="[756,831,983,1007]" pageId="4" pageNumber="37" type="male">1 male</specimenCount>
|
|||
|
|
imago (wings and genitalia on slide),
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FB4F0CC00775E68F" box="[1263,1338,983,1007]" pageId="4" pageNumber="37" type="male">9 male</specimenCount>
|
|||
|
|
subimagos, and
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FEA30CED0327E172" box="[259,360,1018,1042]" pageId="4" pageNumber="37" type="female">4 female</specimenCount>
|
|||
|
|
subimagos from
|
|||
|
|
<collectingCountry id="F32ADD901E7CFFC4FD980CEC00EDE173" box="[568,674,1019,1043]" name="Brazil" pageId="4" pageNumber="37">BRAZIL</collectingCountry>
|
|||
|
|
: AM 240 - Km 21: Sitio da Corredeira Santo Amaro, Presidente Figueiredo, Amazonas,
|
|||
|
|
<quantity id="4CC530E51E7CFFC4FE030B0803A4E157" box="[419,491,1055,1079]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" pageId="4" pageNumber="37" unit="m" value="112.0">112 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FF83BBC01E7CFFC4FE580B08003FE157" box="[504,624,1055,1079]" pageId="4" pageNumber="37" value="2008-04-21">21.iv.2008</date>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FDDC0B080144E157" box="[636,779,1055,1079]" direction="south" orientation="latitude" pageId="4" pageNumber="37" precision="1" value="-2.0392778">S 2° 2' 21.4"</geoCoordinate>
|
|||
|
|
–
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FC880B3701AFE157" box="[808,992,1055,1079]" direction="west" orientation="longitude" pageId="4" pageNumber="37" precision="1" value="-59.845528">W 59° 50' 43.9"</geoCoordinate>
|
|||
|
|
, light trap, N. Hamada col.; and
|
|||
|
|
<specimenCount id="9D3B56891E7CFFC4FAF00B0807D3E157" box="[1360,1436,1055,1079]" pageId="4" pageNumber="37" type="male">1 male</specimenCount>
|
|||
|
|
imago (wings and legs on slide) from COLOMBIA-BRAZIL: Reserva Natural Palmarí, río Yavarí, Muelle Centro Administrativo,
|
|||
|
|
<quantity id="4CC530E51E7CFFC4FEEF0B7003D7E11F" box="[335,408,1127,1151]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="37" unit="m" value="120.0">120 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FF83BBC01E7CFFC4FE030B70005BE11F" box="[419,532,1127,1151]" pageId="4" pageNumber="37" value="2002-05-29">29.v.2002</date>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FDBF0B7000E9E11F" box="[543,678,1127,1151]" direction="south" orientation="latitude" pageId="4" pageNumber="37" precision="15" value="-4.286111">S 4° 17' 10"</geoCoordinate>
|
|||
|
|
–
|
|||
|
|
<geoCoordinate id="EE09FBC71E7CFFC4FD610B7F012CE11F" box="[705,867,1127,1151]" direction="west" orientation="longitude" pageId="4" pageNumber="37" precision="15" value="-70.29694">W 70° 17' 49"</geoCoordinate>
|
|||
|
|
, light trap, M. C. Zúñiga, D. Emmerich, R. Zúñiga and A. Zúñiga col. The
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FE0A0B9C0045E1C3" box="[426,522,1163,1187]" pageId="4" pageNumber="37" type="paratype">paratype</typeStatus>
|
|||
|
|
male imago from river Yavarí is deposited in MUSENUV, one male imago and one female subimago (
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FE3D0BB80043E1A7" box="[413,524,1199,1223]" pageId="4" pageNumber="37" type="paratype">paratypes</typeStatus>
|
|||
|
|
) from
|
|||
|
|
<collectingCountry id="F32ADD901E7CFFC4FDFD0BB800EEE1A7" box="[605,673,1199,1223]" name="Brazil" pageId="4" pageNumber="37">Brazil</collectingCountry>
|
|||
|
|
are deposited in
|
|||
|
|
<collectionCode id="ED2C05C51E7CFFC4FCCB0BA701EEE1A7" box="[875,929,1200,1223]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="4" pageNumber="37">IML</collectionCode>
|
|||
|
|
, the remaining material (
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FB630BB80768E1A7" box="[1219,1319,1199,1223]" pageId="4" pageNumber="37" type="holotype">holotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<typeStatus id="548623A21E7CFFC4FAC20BA7029BE18A" pageId="4" pageNumber="37" type="paratype">paratypes</typeStatus>
|
|||
|
|
) in
|
|||
|
|
<collectionCode id="ED2C05C51E7CFFC4FF5F0BC4030DE18B" box="[255,322,1235,1259]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="4" pageNumber="37">INPA</collectionCode>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|