treatments-xml/data/03/86/F5/0386F515424C845E6C945255B226F90E.xml

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<document id="155A7F657EE022554C41CB6ED9FECA00" ID-DOI="10.26107/RBZ-2024-0010" ID-ISSN="2345-7600" ID-Zenodo-Dep="12171984" ID-ZooBank="FD896355-CE24-4BEE-AAE7-08F774430617" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718828511093" checkinUser="felipe" docAuthor="Amaliah, Rizki, Fahri, F. &amp; Nguyen, Anh D." docDate="2024" docId="0386F515424C845E6C945255B226F90E" docLanguage="en" docName="RafflesBZool.72.116-126.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Amynthas castaneus" docType="treatment" docVersion="2" lastPageNumber="123" masterDocId="FFBF8D6D424A84596C5B5533B75AFFD0" masterDocTitle="New species and new record of the genus Amynthas Kinberg, 1867 (Oligochaeta: Megascolecidae) from Sulawesi, Indonesia" masterLastPageNumber="126" masterPageNumber="116" pageNumber="122" updateTime="1718829009531" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2505F0DC09B9D4938CE39F91086B6E82">New species and new record of the genus Amynthas Kinberg, 1867 (Oligochaeta: Megascolecidae) from Sulawesi, Indonesia</mods:title>
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<mods:namePart id="51CE22D0C32FDD6A8C83C7484E8E5C54">Amaliah, Rizki</mods:namePart>
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<mods:namePart id="A4BA0E53C913A2890E7F78E12DF5A121">Fahri, F.</mods:namePart>
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<mods:namePart id="A4BD2FA5DF5A8A9BFC0718AAD32C4452">Nguyen, Anh D.</mods:namePart>
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<mods:title id="21F7ACE436A0B9DCE342B2904F09C073">Raffles Bulletin of Zoology</mods:title>
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<treatment id="0386F515424C845E6C945255B226F90E" LSID="urn:lsid:plazi:treatment:0386F515424C845E6C945255B226F90E" httpUri="http://treatment.plazi.org/id/0386F515424C845E6C945255B226F90E" lastPageId="7" lastPageNumber="123" pageId="6" pageNumber="122">
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<paragraph id="8B904403424C845F6C945255B5C8F8AE" blockId="6.[207,658,1894,1951]" box="[207,658,1894,1918]" pageId="6" pageNumber="122">
<heading id="D0D8F36F424C845F6C945255B5C8F8AE" bold="true" box="[207,658,1894,1918]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="122" reason="2">
<taxonomicName id="4C2F3F80424C845F6C945255B5C8F8AE" ID-CoL="87NQX" authority="(Michaelsen, 1899)" baseAuthorityName="Michaelsen" baseAuthorityYear="1899" box="[207,658,1894,1918]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="122" phylum="Annelida" rank="species" species="castaneus">
<emphasis id="B95B9811424C845F6C945255B5C8F8AE" bold="true" box="[207,658,1894,1918]" pageId="6" pageNumber="122">
<emphasis id="B95B9811424C845F6C945255B6E9F8AE" bold="true" box="[207,435,1894,1918]" italics="true" pageId="6" pageNumber="122">Amynthas castaneus</emphasis>
(
<bibRefCitation id="EFBE39F2424C845F6D985255B5D6F8AE" author="Michaelsen W" box="[451,652,1894,1918]" pageId="6" pageNumber="122" pagination="1 - 122" refId="ref7313" refString="Michaelsen W (1899) Terricolen von verschiedened Gebieten der Erde. Mittheilungen aus dem Naturhistorischen Museum in Hamburg, 16: 1 - 122." type="journal article" year="1899">Michaelsen, 1899</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="8B904403424C845F6D0C52B5B551F84F" blockId="6.[207,658,1894,1951]" box="[343,523,1926,1951]" pageId="6" pageNumber="122">
(
<tableCitation id="C6AD71B8424C845F6D0552B5B6E8F84F" box="[350,434,1926,1951]" captionStart="Table 1" captionStartId="1.[111,166,511,534]" captionText="Table 1. Earthworms reported from Sulawesi, Indonesia" pageId="6" pageNumber="122">Table 1</tableCitation>
,
<figureCitation id="13145886424C845F6D9952B5B558F84F" box="[450,514,1926,1951]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="6" pageNumber="122">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3351788424C845F6C2A52F6B682F828" pageId="6" pageNumber="122" type="reference_group">
<paragraph id="8B904403424C845F6C2A52F6B682F828" blockId="6.[113,753,1989,2040]" pageId="6" pageNumber="122">
<treatmentCitationGroup id="AB3F632D424C845F6C2A52F6B682F828" pageId="6" pageNumber="122">
<treatmentCitation id="0A8E6212424C845F6C2A52F6B505F80B" author="Michaelsen W" box="[113,607,1989,2011]" page="58" pageId="6" pageNumber="122" year="1899">
<taxonomicName id="4C2F3F80424C845F6C2A52F6B505F80B" ID-CoL="8G36S" authority="Michaelsen, 1899: 56 - 58" authorityName="Michaelsen" authorityPageNumber="56 - 58" authorityYear="1899" box="[113,607,1989,2011]" class="Clitellata" family="Megascolecidae" genus="Amyntas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="122" phylum="Annelida" rank="species" species="castaneus">
<emphasis id="B95B9811424C845F6C2A52F6B79EF80A" box="[113,196,1989,2010]" italics="true" pageId="6" pageNumber="122">Amyntas</emphasis>
[
<materialsCitation id="3B474E5E424C845F6C8852F6B505F80B" box="[211,607,1989,2011]" location="North" municipality="Matinang-Kette" pageId="6" pageNumber="122" specimenCount="1" stateProvince="Sulawesi Utara">
sic]
<emphasis id="B95B9811424C845F6D5B52F5B605F80B" box="[256,351,1990,2011]" italics="true" pageId="6" pageNumber="122">castaneus</emphasis>
<bibRefCitation id="EFBE39F2424C845F6D3C52F6B505F80B" author="Michaelsen W" box="[359,607,1989,2011]" pageId="6" pageNumber="122" pagination="1 - 122" refId="ref7313" refString="Michaelsen W (1899) Terricolen von verschiedened Gebieten der Erde. Mittheilungen aus dem Naturhistorischen Museum in Hamburg, 16: 1 - 122." type="journal article" year="1899">Michaelsen, 1899: 5658</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="5494FAA1424C845F6E3552F6B5C3F80A" box="[622,665,1989,2010]" pageId="6" pageNumber="122">type</typeStatus>
locality:
<collectingMunicipality id="6BF4DE79424C845F6CCC52D1B668F828" box="[151,306,2018,2040]" pageId="6" pageNumber="122">Matinang-Kette</collectingMunicipality>
,
<location id="8EF012D8424C845F6D6652D1B62FF828" LSID="urn:lsid:plazi:treatment:0386F515424C845E6C945255B226F90E:8EF012D8424C845F6D6652D1B62FF828" box="[317,373,2018,2040]" municipality="Matinang-Kette" name="North" pageId="6" pageNumber="122" stateProvince="Sulawesi Utara">North</location>
<collectingRegion id="49EB8AE1424C845F6D2652D1B697F828" box="[381,461,2018,2040]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="122">Celebes</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3351788424C845F6F765095B24FF9EE" pageId="6" pageNumber="122" type="materials_examined">
<paragraph id="8B904403424C845F6F765095B24FF9EE" blockId="6.[813,1452,1446,1598]" pageId="6" pageNumber="122">
<emphasis id="B95B9811424C845F6F765095B341FA6E" bold="true" box="[813,1051,1446,1470]" pageId="6" pageNumber="122">Material examined.</emphasis>
<materialsCitation id="3B474E5E424C845F687D5095B24BF9EE" collectingDate="2016-11-03" collectionCode="UNTAD" collectorName="Rizki Amaliah" country="Indonesia" county="South Kulawi" elevation="2" latitude="-1.3264638" location="Lake Kalimpa'a" longLatPrecision="1" longitude="120.3102" municipality="Lore Lindu National Park" pageId="6" pageNumber="122" specimenCount="1" stateProvince="Sulawesi Tengah">
2 clitellates (
<collectionCode id="ED3EDCC6424C845F68E45095B27FFA6E" box="[1215,1317,1446,1470]" pageId="6" pageNumber="122">UNTAD</collectionCode>
. Oli 0013), primary forests, near
<location id="8EF012D8424C845F684850F5B399FA0E" LSID="urn:lsid:plazi:treatment:0386F515424C845E6C945255B226F90E:8EF012D8424C845F684850F5B399FA0E" box="[1043,1219,1478,1502]" country="Indonesia" county="South Kulawi" latitude="-1.3264638" longLatPrecision="1" longitude="120.3102" municipality="Lore Lindu National Park" name="Lake Kalimpa'a" pageId="6" pageNumber="122" stateProvince="Sulawesi Tengah">Lake Kalimpaa</location>
,
<collectingMunicipality id="6BF4DE79424C845F689450F5B404FA2E" pageId="6" pageNumber="122">Lore Lindu National Park</collectingMunicipality>
(
<geoCoordinate id="EE1B22C4424C845F6F2B50D6B34AFA2E" box="[880,1040,1509,1534]" degrees="01" direction="south" minutes="19" orientation="latitude" pageId="6" pageNumber="122" precision="1" seconds="35.27" value="-1.3264638">01°1935.27″S</geoCoordinate>
,
<geoCoordinate id="EE1B22C4424C845F684450D6B394FA2E" box="[1055,1230,1509,1534]" degrees="120" direction="east" minutes="18" orientation="longitude" pageId="6" pageNumber="122" precision="1" seconds="36.71" value="120.3102">120°1836.71″E</geoCoordinate>
), elevation of
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<quantity id="4CD7E9E6424C845F692B50D5B418F9CD" metricMagnitude="0" metricUnit="m" metricValue="1.674" pageId="6" pageNumber="122" unit="m" value="1.674">1,674 m</quantity>
asl
</elevation>
,
<collectingCounty id="62F13C8F424C845F6F2F5335B350F9CE" box="[884,1034,1542,1566]" pageId="6" pageNumber="122">South Kulawi</collectingCounty>
,
<collectingRegion id="49EB8AE1424C845F684E5335B268F9CE" box="[1045,1330,1542,1566]" country="Indonesia" name="Sulawesi Tengah" pageId="6" pageNumber="122">Central Sulawesi province</collectingRegion>
,
<collectingCountry id="F3380493424C845F69665335B2FDF9CE" box="[1341,1447,1542,1566]" name="Indonesia" pageId="6" pageNumber="122">Indonesia</collectingCountry>
, leg.
<collectorName id="26DA21D5424C845F6F065315B4A1F9EE" box="[861,1019,1574,1598]" pageId="6" pageNumber="122">Rizki Amaliah</collectorName>
et al.,
<date id="FF9162C3424C845F68125315B24BF9EE" box="[1097,1297,1574,1598]" pageId="6" pageNumber="122" value="2016-11-03">
<collectingDate id="EFD59B2B424C845F68125315B24BF9EE" box="[1097,1297,1574,1598]" pageId="6" pageNumber="122" value="2016-11-03">3 November 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3351788424C845E6F765355B5C5F7CB" lastPageId="7" lastPageNumber="123" pageId="6" pageNumber="122" type="description">
<paragraph id="8B904403424C845E6F765355B664F88C" blockId="6.[811,1452,1638,2078]" lastBlockId="7.[135,775,1603,1884]" lastPageId="7" lastPageNumber="123" pageId="6" pageNumber="122">
<emphasis id="B95B9811424C845F6F765355B4E2F9AE" bold="true" box="[813,952,1638,1662]" pageId="6" pageNumber="122">Description.</emphasis>
Small-sized worm, length
<quantity id="4CD7E9E6424C845F68B35355B205F9AD" box="[1256,1375,1638,1662]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.4" metricValueMin="5.6" pageId="6" pageNumber="122" unit="mm" value="60.0" valueMax="64.0" valueMin="56.0">5664 mm</quantity>
(n=2), diameter 1.52.0 mm, and segments 7283. Prostomium epilobous. First dorsal pore in 12/13. Setal numbers:
<quantity id="4CD7E9E6424C845F69325395B418F90E" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="7.874" metricValueMin="7.112" pageId="6" pageNumber="122" unit="in" value="29.5" valueMax="31.0" valueMin="28.0">2831 in</quantity>
v,
<quantity id="4CD7E9E6424C845F6F3C53F5B492F90E" box="[871,968,1734,1758]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.652" metricValueMin="8.128" pageId="6" pageNumber="122" unit="in" value="35.0" valueMax="38.0" valueMin="32.0">3238 in</quantity>
vii,
<quantity id="4CD7E9E6424C845F6FA753F5B307F90E" box="[1020,1117,1734,1758]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="9.144" metricValueMin="7.366" pageId="6" pageNumber="122" unit="in" value="32.5" valueMax="36.0" valueMin="29.0">2936 in</quantity>
viii,
<quantity id="4CD7E9E6424C845F68C353F5B3A3F90E" box="[1176,1273,1734,1758]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" metricValueMax="9.652" metricValueMin="8.636" pageId="6" pageNumber="122" unit="in" value="36.0" valueMax="38.0" valueMin="34.0">3438 in</quantity>
xxv, and 1012 between male porophores in ventral xviii. Three pairs of spermathecal pores in lateroventral 6/7/8/9, distance between spermathecal pores about 0.35x body circumference ventrally (
<figureCitation id="13145886424C845F6F685275B4D0F88E" box="[819,906,1862,1886]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="6" pageNumber="122">Fig. 3A</figureCitation>
). Male pores simple, distance between male pores about 0.2x body circumference ventrally. Genital markings present, small, widely paired in pre- and post-setal xvii, and pre-setal xviiixix (
<figureCitation id="13145886424C845F68135295B3FDF86E" box="[1096,1191,1958,1982]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="6" pageNumber="122">Fig. 3D</figureCitation>
). Septa 8/9/10 absent. Spermathecae large, paired in viiix (
<figureCitation id="13145886424C845F68EC52F5B253F80E" box="[1207,1289,1990,2014]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="6" pageNumber="122">Fig. 3B</figureCitation>
). Spermathecal ampulla rounded, duct enlarged, about 2/5 as long as ampulla. Diverticulum cylindrical, about 1/2 as long as ampulla, attached to ducts. Holandric. Prostatic duct C-shaped. Accessory gland present, stalked. Oesophageal gizzard large in viiix. Oesophageal hearts paired in xixiii. Intestine beginning in xv; caeca originating from xxvii extending anteriorly to xxv (
<figureCitation id="13145886424D845E6D0053F0B6EEF90C" box="[347,436,1731,1756]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="7" pageNumber="123">Fig. 3E</figureCitation>
), paired laterally. Typhlosole simple. Testes sacs yellowish, small, ventrally paired but separated in xxi. Seminal vesicles in xixii. Ovaries paired in xiii; ovisacs paired in xiii. Prostate glands racemose, paired (
<figureCitation id="13145886424D845E6C805270B66BF88C" box="[219,305,1859,1884]" captionStart="Fig" captionStartId="7.[136,172,1469,1491]" captionTargetBox="[136,1475,189,1453]" captionTargetId="figure-61@7.[121,1483,172,1463]" captionTargetPageId="7" captionText="Fig. 3. Amynthas castaneus from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="7" pageNumber="123">Fig. 3C</figureCitation>
).
</paragraph>
<caption id="DF50148B424D845E6CD3508EB595F9DC" ID-DOI="http://doi.org/10.5281/zenodo.12171990" ID-Zenodo-Dep="12171990" httpUri="https://zenodo.org/record/12171990/files/figure.png" pageId="7" pageNumber="123" startId="7.[136,172,1469,1491]" targetBox="[136,1475,189,1453]" targetPageId="7" targetType="figure">
<paragraph id="8B904403424D845E6CD3508EB595F9DC" blockId="7.[136,1474,1469,1548]" pageId="7" pageNumber="123">
Fig. 3.
<taxonomicName id="4C2F3F80424D845E6C8E508EB6FAFA02" baseAuthorityName="Michaelsen" baseAuthorityYear="1899" box="[213,416,1469,1490]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="123" phylum="Annelida" rank="species" species="castaneus">
<emphasis id="B95B9811424D845E6C8E508EB6FAFA02" box="[213,416,1469,1490]" italics="true" pageId="7" pageNumber="123">Amynthas castaneus</emphasis>
</taxonomicName>
from Lore Lindu National Park, Central Sulawesi (UNTAD. Oli 0014). A, spermathecal pores (sp) B, spermathecae, right side in viiix (amp = ampulla; dv = diverticulum) C, prostate gland D, male pore region (mp = male pore, gm = genital markings) E, intestinal caecum. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B904403424D845E6CD352B0B5C5F7CB" blockId="7.[135,776,1923,2075]" pageId="7" pageNumber="123">
<emphasis id="B95B9811424D845E6CD352B0B62EF84C" bold="true" box="[136,372,1923,1948]" pageId="7" pageNumber="123">Habitat and ecology.</emphasis>
Specimens were collected in primary forest near Lake Kalimpaa, Lore Lindu National Park, South Kulawi,
<collectingRegion id="49EB8AE1424D845E6CBD52F0B6C4F80B" box="[230,414,1987,2011]" country="Indonesia" name="Sulawesi Tengah" pageId="7" pageNumber="123">Central Sulawesi</collectingRegion>
at the elevation of
<quantity id="4CD7E9E6424D845E6E2752F0B58DF80B" box="[636,727,1987,2011]" metricMagnitude="0" metricUnit="m" metricValue="1.674" pageId="7" pageNumber="123" unit="m" value="1.674">1,674 m</quantity>
asl. This species was found at the depth of
<quantity id="4CD7E9E6424D845E6E6252D0B5FFF82B" box="[569,677,2019,2043]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="7" pageNumber="123" unit="cm" value="17.5" valueMax="20.0" valueMin="15.0">1520 cm</quantity>
in moist and sandy soils around rocks and decayed wood.
</paragraph>
</subSubSection>
<subSubSection id="C3351788424D845E6F1E5375B226F90E" pageId="7" pageNumber="123" type="discussion">
<paragraph id="8B904403424D845E6F1E5375B226F90E" blockId="7.[834,1475,1606,1758]" pageId="7" pageNumber="123">
<emphasis id="B95B9811424D845E6F1E5375B4E8F98E" bold="true" box="[837,946,1606,1630]" pageId="7" pageNumber="123">Remarks.</emphasis>
This is the second record since it was first described by
<bibRefCitation id="EFBE39F2424D845E6F385355B373F9AE" author="Michaelsen W" box="[867,1065,1638,1662]" pageId="7" pageNumber="123" pagination="1 - 122" refId="ref7313" refString="Michaelsen W (1899) Terricolen von verschiedened Gebieten der Erde. Mittheilungen aus dem Naturhistorischen Museum in Hamburg, 16: 1 - 122." type="journal article" year="1899">Michaelsen (1899)</bibRefCitation>
from Matinang-Kette,
<collectingRegion id="49EB8AE1424D845E69425355B2E5F9AE" box="[1305,1471,1638,1662]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="123">North Sulawesi</collectingRegion>
. Specimens described by Michaelsen are slightly different from the present material in having diverticulum with slender stalk and an elongated rod-shaped seminal chamber.
</paragraph>
</subSubSection>
</treatment>
</document>