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<document id="E469D82C45D0BE2467587BD2F96AE10F" ID-CLB-Dataset="21898" ID-DOI="10.5852/ejt.2016.216" ID-GBIF-Dataset="45c06988-c82e-40ce-8a48-234d4b9c1a99" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840149" ID-ZooBank="2F526459-E23A-458B-9829-31F4C1BC6C46" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590165220169" checkinUser="valdenar" docAuthor="Martens, Jochen &amp; Schönhofer, Axel L." docDate="2016" docId="BA4E87D3FFEBBB1C724EFD94C927FC2C" docLanguage="en" docName="ejt-2016-216.pdf.imf" docOrigin="European Journal of Taxonomy 216 (216)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leiobunum rupestre" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="4677FFABFFEFBB14700BFFBFC826FFB0" masterDocTitle="The Leiobunum rupestre species group: resolving the taxonomy of four widespread European taxa (Opiliones: Sclerosomatidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="5" updateTime="1698841016496" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6B4754F113654E02DF14DA129775D3E5">The Leiobunum rupestre species group: resolving the taxonomy of four widespread European taxa (Opiliones: Sclerosomatidae)</mods:title>
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<mods:namePart id="8FF0E3DC86726F32D596B99B88D5C3CA">Martens, Jochen</mods:namePart>
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<mods:affiliation id="0319AA3768D646B9BAC2F47C04C19887">Johannes Gutenberg-Universität Mainz, Institut für Zoologie, 55099 Mainz, Germany &amp; urn: lsid: zoobank. org: author: F 01 C 78 F 1 - 5 EBF- 48 E 8 - B 37 E- 634 DF 00 E 0237 &amp; Corresponding author: martens @ uni-mainz. de</mods:affiliation>
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<mods:namePart id="931715E48F7B85975A7B1E9D26FA4FF8">Schönhofer, Axel L.</mods:namePart>
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<mods:affiliation id="2365B09802EF109EABC755B1E02E9223">Johannes Gutenberg-Universität Mainz, Institut für Zoologie, 55099 Mainz, Germany &amp; Email: Axel. Schoenhofer @ gmx. net &amp; urn: lsid: zoobank. org: author: 91 B 2971 C-D 8 CE- 4 F 98 - 903 F-C 963 E 83 E 8 C 7 C</mods:affiliation>
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<treatment id="BA4E87D3FFEBBB1C724EFD94C927FC2C" ID-DOI="http://doi.org/10.5281/zenodo.3853162" ID-GBIF-Taxon="164250213" ID-Zenodo-Dep="3853162" LSID="urn:lsid:plazi:treatment:BA4E87D3FFEBBB1C724EFD94C927FC2C" httpUri="http://treatment.plazi.org/id/BA4E87D3FFEBBB1C724EFD94C927FC2C" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
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<paragraph id="325836C5FFEBBB10724EFD94CBC8FDF5" blockId="4.[552,1035,555,617]" box="[581,1006,555,582]" pageId="4" pageNumber="5">
<heading id="691081A9FFEBBB10724EFD94CBC8FDF5" box="[581,1006,555,582]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="F5E74D46FFEBBB10724EFD94CBC8FDF5" authority="(Herbst, 1799)" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[581,1006,555,582]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB10724EFD94CB1AFDF6" bold="true" box="[581,828,555,582]" italics="true" pageId="4" pageNumber="5">Leiobunum rupestre</emphasis>
(
<bibRefCitation id="56764B34FFEBBB107340FD94CBC0FDF5" author="Herbst J. F. W." box="[843,998,555,581]" pageId="4" pageNumber="5" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst, 1799</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7AFD654EFFEBBB107223FDF1CCBDFD41" pageId="4" pageNumber="5" type="description">
<paragraph id="325836C5FFEBBB107223FDF1CC2DFDD9" blockId="4.[552,1035,555,617]" box="[552,1035,590,617]" pageId="4" pageNumber="5">
<figureCitation id="AADC2A40FFEBBB107223FDF1CA4AFDD9" box="[552,620,590,617]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[189,1398,767,1853]" captionTargetId="figure@5.[189,1398,767,1853]" captionTargetPageId="5" captionText="Fig. 1. Habit of Leiobunum rupestre species group. AB. Leiobunum rupestre Herbst, 1799, Slovenia, Pohorje Mountains, resting at rock faces. A. ♁. B. ♀. CD. Leiobunum apenninicum (Martens, 1969), Italy, Monesi di Triora, at night. C. ♁. D. ♀. EF. Leiobunum gracile Thorell, 1876, Denmark. E. ♁. F. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field." figureDoi="http://doi.org/10.5281/zenodo.3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
AB, 2, 3AB, 4AB, HG, 5AC
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FD2FCCBDFD41" blockId="4.[189,1399,656,753]" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FD2FCA87FD1A" authority="Fabricius, 1793: 429" authorityName="Fabricius" authorityPageNumber="429" authorityYear="1793" box="[189,673,656,682]" class="Liliopsida" family="Asphodelaceae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="0093EAD7FFEBBB1070B6FD2FC981FD1A" box="[189,423,656,682]" italics="true" pageId="4" pageNumber="5">Phalangium bicolor</emphasis>
Fabricius, 1793: 429
</taxonomicName>
. The name was synonymized with
<taxonomicName id="F5E74D46FFEBBB107438FD2FCCCCFD1A" baseAuthorityName="Olivier" baseAuthorityYear="1791" box="[1075,1258,656,682]" class="Arachnida" family="Sclerosomatidae" genus="Gyas" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="0093EAD7FFEBBB107438FD2FCCCCFD1A" box="[1075,1258,656,682]" italics="true" pageId="4" pageNumber="5">Gyas annulatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="56764B34FFEBBB107518FD2FC928FD7D" author="Latreille P. A." pageId="4" pageNumber="5" pagination="314 - 329" refId="ref13050" refString="Latreille P. A. 1804. Huitieme genre - Dixieme genre et particuliere des Crustaces et des Insectes. In: Sonnini C. S. (ed.) Histoire naturelle, generale et particuliere des Crustaces et des Insectes 7: 314 - 329. F. Dufart, Paris. Available from http: // biodiversitylibrary. org / page / 24883291 [accessed 23 Jun. 2016]" type="journal article" year="1804">Latreille (1804)</bibRefCitation>
, which was missed by several subsequent authors; some of these used it as valid name, in synonymy of, or in relation to,
<taxonomicName id="F5E74D46FFEBBB10720AFD68CAA7FD40" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[513,641,727,752]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB10720AFD68CAA7FD40" box="[513,641,727,752]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
(e.g., C.L. Koch 1847, 16: 56; redescription).
</paragraph>
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<subSubSection id="7AFD654EFFEBBB1070B6FCA7C980FAEB" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="325836C5FFEBBB1070B6FCA7CAD9FCE6" blockId="4.[189,1399,792,889]" pageId="4" pageNumber="5">
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<treatmentCitation id="B34610D4FFEBBB1070B6FCA7CA14FC82" author="Herbst J. F. W." box="[189,562,792,818]" page="4" pageId="4" pageNumber="5" year="1799">
<taxonomicName id="F5E74D46FFEBBB1070B6FCA7CA14FC82" authority="Herbst, 1799: 4" authorityName="Herbst" authorityPageNumber="4" authorityYear="1799" box="[189,562,792,818]" class="Arachnida" family="Phalangiidae" genus="Opilio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestris">
<emphasis id="0093EAD7FFEBBB1070B6FCA7C953FC82" box="[189,373,792,818]" italics="true" pageId="4" pageNumber="5">Opilio rupestris</emphasis>
<bibRefCitation id="56764B34FFEBBB107177FCA7CA14FC82" author="Herbst J. F. W." box="[380,562,792,818]" pageId="4" pageNumber="5" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst, 1799: 4</bibRefCitation>
</taxonomicName>
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, fig. 1 (material from the
<typeStatus id="ED5C8867FFEBBB107350FCA7CBABFC82" box="[859,909,792,818]" pageId="4" pageNumber="5">type</typeStatus>
locality
<collectingRegion id="F023F827FFEBBB1073F9FCA7CC77FC82" box="[1010,1105,792,818]" country="Germany" name="Sachsen" pageId="4" pageNumber="5">Sachsen</collectingRegion>
, i.e.,
<collectingRegion id="F023F827FFEBBB10749AFCA7CCCFFC82" box="[1169,1257,792,818]" country="Germany" name="Sachsen" pageId="4" pageNumber="5">Saxony</collectingRegion>
, examined).
<treatmentCitation id="B34610D4FFEBBB1070B6FC84CAA5FCE6" author="Koch C. L." box="[189,643,827,854]" page="59" pageId="4" pageNumber="5" year="1848">
<taxonomicName id="F5E74D46FFEBBB1070B6FC84CAA5FCE6" authority="C. L. Koch, 1848: 59" authorityName="C. L. Koch" authorityPageNumber="59" authorityYear="1848" box="[189,643,827,854]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ovale">
<emphasis id="0093EAD7FFEBBB1070B6FC84C9AEFCE5" box="[189,392,827,853]" italics="true" pageId="4" pageNumber="5">Leiobunum ovale</emphasis>
C.L.
<bibRefCitation id="56764B34FFEBBB1071C0FC84CAA5FCE6" author="Koch C. L." box="[459,643,827,854]" pageId="4" pageNumber="5" refId="ref12141" refString="Koch C. L. 1848. Die Arachniden. Sechzehnter Band. J. L. Lotzbeck, Nurnberg. Available from http: // biodiversitylibrary. org / page / 16576103 [accessed 23 Jun. 2016]" type="url" year="1848">Koch, 1848: 59</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1540.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FCE1CA96FCC8" blockId="4.[189,1399,792,889]" box="[189,688,862,889]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FCE1CA96FCC8" box="[189,688,862,889]" pageId="4" pageNumber="5">
<treatmentCitation id="B34610D4FFEBBB1070B6FCE1CA56FCC8" author="Koch L." box="[189,624,862,889]" page="4" pageId="4" pageNumber="5" year="1869">
<taxonomicName id="F5E74D46FFEBBB1070B6FCE1CA56FCC8" authority="L. Koch, 1869: 4" authorityName="L. Koch" authorityPageNumber="4" authorityYear="1869" box="[189,624,862,889]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFEBBB1070B6FCE1C9B8FCC8" box="[189,414,862,888]" italics="true" pageId="4" pageNumber="5">Liobunum glabrum</emphasis>
L.
<bibRefCitation id="56764B34FFEBBB1071CCFCE1CA56FCC8" author="Koch L." box="[455,624,862,888]" pageId="4" pageNumber="5" pagination="149 - 206" refId="ref12226" refString="Koch L. 1869. Beitrag zur Kenntniss der Arachnidenfauna Tirols. Zeitschrift des Ferdinandeums fur Tirol und Vorarlberg, Innsbruck, series 3 (14): 149 - 206." type="journal article" year="1869">Koch, 1869: 4</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 67.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FC1FCD2EFC0A" blockId="4.[189,1399,926,1306]" box="[189,1288,926,954]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FC1FCD2EFC0A" box="[189,1288,926,954]" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FC1FC9AEFC0A" authorityName="C. L. Koch" authorityYear="1848" box="[189,392,928,954]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ovale">
<emphasis id="0093EAD7FFEBBB1070B6FC1FC9AEFC0A" box="[189,392,928,954]" italics="true" pageId="4" pageNumber="5">Leiobunum ovale</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB107184FC1FC9B8FC0A" bold="true" box="[399,414,928,954]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071AEFC1FCAA4FC0A" author="Roewer C. F." box="[421,642,928,954]" page="203" pageId="4" pageNumber="5" year="1910">
<bibRefCitation id="56764B34FFEBBB1071AEFC1FCAA4FC0A" author="Roewer C. F." box="[421,642,928,954]" pageId="4" pageNumber="5" pagination="1 - 294" refId="ref13921" refString="Roewer C. F. 1910. Revision der Opiliones Plagiostethi (= Opiliones Palpatores). I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Liobunini, Leptobunini.) Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg 19 (4): 1 - 294." type="journal article" year="1910">Roewer 1910: 203</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107286FC1FCA8CFC0A" bold="true" box="[653,682,928,954]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1072BAFC21CBADFC0A" author="Silhavy V." box="[689,907,926,954]" page="174" pageId="4" pageNumber="5" year="1956">
<bibRefCitation id="56764B34FFEBBB1072BAFC21CBADFC0A" author="Silhavy V." box="[689,907,926,954]" pageId="4" pageNumber="5" refId="ref14153" refString="Silhavy V. 1956. Sekaci - Opilionidea. Fauna CSR 7. Akademia Nauk, Praha. [in Czech]" type="book" year="1956">Šilhavý 1956: 174</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB10739CFC1FCB92FC0A" bold="true" box="[919,948,928,954]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1073B7FC1FCD22FC0A" author="Novak T. &amp; Gruber J." box="[956,1284,928,954]" page="299" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="56764B34FFEBBB1073B7FC1FCD22FC0A" author="Novak T. &amp; Gruber J." box="[956,1284,928,954]" pageId="4" pageNumber="5" pagination="281 - 308" refId="ref13819" refString="Novak T. &amp; Gruber J. 2000. Remarks on published data on harvestmen (Arachnida: Opiliones) from Slovenia. Annales, Series historia naturalis 10 (2 / 21): 281 - 308." type="journal article" year="2000">Novak &amp; Gruber 2000: 299</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="325836C5FFEBBB1070B6FC7CCA7BFBB0" blockId="4.[189,1399,926,1306]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FC7CCA7BFBB0" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FC7CC98EFC6D" ID-CoL="4FR7C" authorityName="Fabricius" authorityYear="1793" box="[189,424,963,989]" class="Liliopsida" family="Asphodelaceae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="0093EAD7FFEBBB1070B6FC7CC98EFC6D" box="[189,424,963,989]" italics="true" pageId="4" pageNumber="5">Phalangium bicolor</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB1071A5FC7CC99BFC6D" bold="true" box="[430,445,963,989]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071C8FC7CCAA8FC6D" author="Simon E." box="[451,654,963,989]" page="181" pageId="4" pageNumber="5" year="1879">
<bibRefCitation id="56764B34FFEBBB1071C8FC7CCAA8FC6D" author="Simon E." box="[451,654,963,989]" pageId="4" pageNumber="5" refId="ref14212" refString="Simon E. 1879. Les Arachnides de France. Tome 7. Contenant les ordres des Chernetes, Scorpiones et Opiliones. Librairie Encyclopedique de Roret, Paris." type="book" year="1879">Simon 1879: 181</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107292FC7CCA90FC6D" bold="true" box="[665,694,963,989]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1072B7FC7CCBB5FC6D" author="Roewer C. F." box="[700,915,963,989]" page="203" pageId="4" pageNumber="5" year="1910">
<bibRefCitation id="56764B34FFEBBB1072B7FC7CCBB5FC6D" author="Roewer C. F." box="[700,915,963,989]" pageId="4" pageNumber="5" pagination="1 - 294" refId="ref13921" refString="Roewer C. F. 1910. Revision der Opiliones Plagiostethi (= Opiliones Palpatores). I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Liobunini, Leptobunini.) Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg 19 (4): 1 - 294." type="journal article" year="1910">Roewer 1910: 203</bibRefCitation>
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(both also listing
<taxonomicName id="F5E74D46FFEBBB107468FC7CCD50FC6D" authorityName="Olivier" authorityYear="1791" box="[1123,1398,963,989]" class="Arachnida" family="Phalangiidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Plantae" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="annulatum">
<emphasis id="0093EAD7FFEBBB107468FC7CCD50FC6D" box="[1123,1398,963,989]" italics="true" pageId="4" pageNumber="5">Phalangium annulatum</emphasis>
</taxonomicName>
in synonymy).
<emphasis id="0093EAD7FFEBBB107165FC59C9ADFBB0" bold="true" box="[366,395,998,1024]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107199FC58CA7FFBB0" author="Lessert R. de" box="[402,601,998,1025]" page="16" pageId="4" pageNumber="5" year="1917">
<bibRefCitation id="56764B34FFEBBB107199FC58CA7FFBB0" author="Lessert R. de" box="[402,601,998,1025]" pageId="4" pageNumber="5" pagination="1 - 80" refId="ref13208" refString="Lessert R. de 1917. Opilions. Catalogue des invertebres de la Suisse 9: 1 - 80, Museum d'Histoire Naturelle de Geneve, Geneve." type="journal article" year="1917">Lessert 1917: 16</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FBB6CA82FBDA" blockId="4.[189,1399,926,1306]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FBB6CA82FBDA" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FBB6C9BDFB93" box="[189,411,1033,1059]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB1070B6FBB6C9BDFB93" box="[189,411,1033,1059]" italics="true" pageId="4" pageNumber="5">Liobunum rupestre</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB1071AAFBB6C996FB93" bold="true" box="[417,432,1033,1059]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071BEFBB6CAA6FB93" author="Simon E." box="[437,640,1033,1059]" page="181" pageId="4" pageNumber="5" year="1879">
<bibRefCitation id="56764B34FFEBBB1071BEFBB6CAA6FB93" author="Simon E." box="[437,640,1033,1059]" pageId="4" pageNumber="5" refId="ref14212" refString="Simon E. 1879. Les Arachnides de France. Tome 7. Contenant les ordres des Chernetes, Scorpiones et Opiliones. Librairie Encyclopedique de Roret, Paris." type="book" year="1879">Simon 1879: 181</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107281FBB6CA81FB93" bold="true" box="[650,679,1033,1059]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1072A5FBB6CBAFFB93" author="Cantoni E." box="[686,905,1033,1059]" page="194" pageId="4" pageNumber="5" year="1882">
<bibRefCitation id="56764B34FFEBBB1072A5FBB6CBAFFB93" author="Cantoni E." box="[686,905,1033,1059]" pageId="4" pageNumber="5" pagination="191 - 203" refId="ref11277" refString="Cantoni E. 1882. Escursione in Calabria (1877). Chernetidi ed Opilionidi. Bollettino della Societa Entomologica Italiana 14: 191 - 203." type="journal article" year="1882">Cantoni 1882: 194</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107398FBB6CB96FB93" bold="true" box="[915,944,1033,1059]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1073BDFBB5CCABFB94" author="Hansen H. J." box="[950,1165,1033,1060]" page="500" pageId="4" pageNumber="5" year="1884">
<bibRefCitation id="56764B34FFEBBB1073BDFBB5CCABFB94" author="Hansen H. J." box="[950,1165,1033,1060]" pageId="4" pageNumber="5" pagination="491 - 554" refId="ref11846" refString="Hansen H. J. 1884. Arthrogastra Danica: en monographisk fremstilling af de i Danmark levende Meiere og Mosskorpioner med bidrag til sidstnaevnte underordens systematic. Naturhistorisk Tidsskrift (3) 14: 491 - 554. [in Danish]" type="journal article" year="1884">Hansen 1884: 500</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB10749CFBB6CC92FB93" bold="true" box="[1175,1204,1033,1059]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1074B1FBB6C8C8FBF7" author="Kraepelin K." page="222" pageId="4" pageNumber="5" year="1896">
<bibRefCitation id="56764B34FFEBBB1074B1FBB6C8C8FBF7" author="Kraepelin K." pageId="4" pageNumber="5" pagination="219 - 234" refId="ref12976" refString="Kraepelin K. 1896. Phalangiden aus der Umgebung Hamburgs. Mitteilungen aus dem naturhistorischen Museum in Hamburg 13: 219 - 234. Available from http: // biodiversitylibrary. org / page / 29521651 [accessed 23 Jun. 2016]" type="journal article" year="1896">Kraepelin 1896: 222</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB1070F2FB92C931FBF7" bold="true" box="[249,279,1069,1095]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107115FB92C9D8FBF7" author="Roewer C. F." box="[286,510,1069,1095]" page="197" pageId="4" pageNumber="5" year="1910">
<bibRefCitation id="56764B34FFEBBB107115FB92C9D8FBF7" author="Roewer C. F." box="[286,510,1069,1095]" pageId="4" pageNumber="5" pagination="1 - 294" refId="ref13921" refString="Roewer C. F. 1910. Revision der Opiliones Plagiostethi (= Opiliones Palpatores). I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Liobunini, Leptobunini.) Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg 19 (4): 1 - 294." type="journal article" year="1910">Roewer 1910: 197</bibRefCitation>
</treatmentCitation>
, 203.
<emphasis id="0093EAD7FFEBBB10724DFB92CA42FBF7" bold="true" box="[582,612,1069,1095]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107267FB92CB11FBF7" author="Lessert R. de" box="[620,823,1068,1095]" page="15" pageId="4" pageNumber="5" year="1917">
<bibRefCitation id="56764B34FFEBBB107267FB92CB11FBF7" author="Lessert R. de" box="[620,823,1068,1095]" pageId="4" pageNumber="5" pagination="1 - 80" refId="ref13208" refString="Lessert R. de 1917. Opilions. Catalogue des invertebres de la Suisse 9: 1 - 80, Museum d'Histoire Naturelle de Geneve, Geneve." type="journal article" year="1917">Lessert 1917: 15</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107348FB92CB47FBF7" bold="true" box="[835,865,1069,1095]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107363FB92CC63FBF7" author="Roewer C. F." box="[872,1093,1069,1095]" page="890" pageId="4" pageNumber="5" year="1923">
<bibRefCitation id="56764B34FFEBBB107363FB92CC63FBF7" author="Roewer C. F." box="[872,1093,1069,1095]" pageId="4" pageNumber="5" refId="ref13978" refString="Roewer C. F. 1923. Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. G. Fischer Jena." type="book" year="1923">Roewer 1923: 890</bibRefCitation>
</treatmentCitation>
(partim).
<emphasis id="0093EAD7FFEBBB1074B5FB92CCFAFBF7" bold="true" box="[1214,1244,1069,1095]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1074E8FB93C8CBFBDA" author="Hadzi J." page="145" pageId="4" pageNumber="5" year="1931">
<bibRefCitation id="56764B34FFEBBB1074E8FB93C8CBFBDA" author="Hadzi J." pageId="4" pageNumber="5" pagination="107 - 154" refId="ref11762" refString="Hadzi J. 1931. Opilioni Triglavskoga masiva [Die Opilionen des Triglavmassivs]. Prirodoslovne Razprave (Muzejsko Drustvo za Slovenijo) Ljubljana 1: 107 - 154. [in Slovenian, German abstract]" type="journal article" year="1931">Hadži 1931: 145</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB1070F2FBEFC931FBDA" bold="true" box="[249,279,1104,1130]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107115FBF1C9DEFBDA" author="Silhavy V." box="[286,504,1102,1130]" page="174" pageId="4" pageNumber="5" year="1956">
<bibRefCitation id="56764B34FFEBBB107115FBF1C9DEFBDA" author="Silhavy V." box="[286,504,1102,1130]" pageId="4" pageNumber="5" refId="ref14153" refString="Silhavy V. 1956. Sekaci - Opilionidea. Fauna CSR 7. Akademia Nauk, Praha. [in Czech]" type="book" year="1956">Šilhavý 1956: 174</bibRefCitation>
</treatmentCitation>
, figs 437443.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FBCCCB3BFB3D" blockId="4.[189,1399,926,1306]" box="[189,797,1139,1166]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FBCCCB3BFB3D" box="[189,797,1139,1166]" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FBCCC9BCFB3D" box="[189,410,1139,1165]" class="Arachnida" family="Sclerosomatidae" genus="Liobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestris">
<emphasis id="0093EAD7FFEBBB1070B6FBCCC9BCFB3D" box="[189,410,1139,1165]" italics="true" pageId="4" pageNumber="5">Liobunus rupestris</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB1071AAFBCCC996FB3D" bold="true" box="[417,432,1139,1165]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071BCFBCCCB3CFB3D" author="Muller F. &amp; Schenkel E." box="[439,794,1139,1166]" page="819" pageId="4" pageNumber="5" year="1895">
<bibRefCitation id="56764B34FFEBBB1071BCFBCCCB3CFB3D" author="Muller F. &amp; Schenkel E." box="[439,794,1139,1166]" pageId="4" pageNumber="5" pagination="691 - 824" refId="ref13625" refString="Muller F. &amp; Schenkel E. 1895. Verzeichnis der Spinnen von Basel und Umgegend. Verhandlungen der naturforschenden Gesellschaft zu Basel 10: 691 - 824. Available from http: // biodiversitylibrary. org / page / 34087160 [accessed 23 Jun. 2016]" type="journal article" year="1895">Müller &amp; Schenkel 1895: 819</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FB29CAABFB00" blockId="4.[189,1399,926,1306]" box="[189,653,1174,1200]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FB29CAABFB00" box="[189,653,1174,1200]" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FB29C9B8FB00" authorityName="L. Koch" authorityYear="1869" box="[189,414,1174,1200]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFEBBB1070B6FB29C9B8FB00" box="[189,414,1174,1200]" italics="true" pageId="4" pageNumber="5">Liobunum glabrum</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB1071ADFB29C993FB00" bold="true" box="[422,437,1174,1200]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071B7FB29CAAFFB00" author="Simon E." box="[444,649,1174,1200]" page="181" pageId="4" pageNumber="5" year="1879">
<bibRefCitation id="56764B34FFEBBB1071B7FB29CAAFFB00" author="Simon E." box="[444,649,1174,1200]" pageId="4" pageNumber="5" refId="ref14212" refString="Simon E. 1879. Les Arachnides de France. Tome 7. Contenant les ordres des Chernetes, Scorpiones et Opiliones. Librairie Encyclopedique de Roret, Paris." type="book" year="1879">Simon 1879: 181</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FB06CB7AFB63" blockId="4.[189,1399,926,1306]" box="[189,860,1209,1236]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FB06CB7AFB63" box="[189,860,1209,1236]" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FB06C941FB63" box="[189,359,1209,1235]" class="Arachnida" family="Gagrellidae" genus="Nelima" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0093EAD7FFEBBB1070B6FB06C941FB63" box="[189,359,1209,1235]" italics="true" pageId="4" pageNumber="5">Nelima glabra</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB107164FB06C958FB63" bold="true" box="[367,382,1209,1235]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB10718EFB05CA47FB63" author="Roewer C. F." box="[389,609,1209,1236]" page="239" pageId="4" pageNumber="5" year="1910">
<bibRefCitation id="56764B34FFEBBB10718EFB05CA47FB63" author="Roewer C. F." box="[389,609,1209,1236]" pageId="4" pageNumber="5" pagination="1 - 294" refId="ref13921" refString="Roewer C. F. 1910. Revision der Opiliones Plagiostethi (= Opiliones Palpatores). I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Liobunini, Leptobunini.) Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg 19 (4): 1 - 294." type="journal article" year="1910">Roewer 1910: 239</bibRefCitation>
</treatmentCitation>
.
<emphasis id="0093EAD7FFEBBB107267FB06CAAFFB63" bold="true" box="[620,649,1209,1235]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB10729AFB06CB71FB63" author="Hadzi J." box="[657,855,1209,1235]" page="148" pageId="4" pageNumber="5" year="1931">
<bibRefCitation id="56764B34FFEBBB10729AFB06CB71FB63" author="Hadzi J." box="[657,855,1209,1235]" pageId="4" pageNumber="5" pagination="107 - 154" refId="ref11762" refString="Hadzi J. 1931. Opilioni Triglavskoga masiva [Die Opilionen des Triglavmassivs]. Prirodoslovne Razprave (Muzejsko Drustvo za Slovenijo) Ljubljana 1: 107 - 154. [in Slovenian, German abstract]" type="journal article" year="1931">Hadži 1931: 148</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FB63CB1AFAAA" blockId="4.[189,1399,926,1306]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="12F711EBFFEBBB1070B6FB63CB1AFAAA" pageId="4" pageNumber="5">
<taxonomicName id="F5E74D46FFEBBB1070B6FB63C98CFB46" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[189,426,1244,1270]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB1070B6FB63C98CFB46" box="[189,426,1244,1270]" italics="true" pageId="4" pageNumber="5">Leiobunum rupestre</emphasis>
</taxonomicName>
<emphasis id="0093EAD7FFEBBB1071BAFB62C9E6FB47" bold="true" box="[433,448,1245,1271]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1071C3FB64CAA2FB47" author="Silhavy V." box="[456,644,1243,1271]" page="9" pageId="4" pageNumber="5" year="1948">
<bibRefCitation id="56764B34FFEBBB1071C3FB64CAA2FB47" author="Silhavy V." box="[456,644,1243,1271]" pageId="4" pageNumber="5" pagination="1 - 108" refId="ref14107" refString="Silhavy V. 1948. Les Opilions de la steppe de Serpentine pres de la ville de Mohelno. Mohelno. Archiv Svazu na vyzkum a ochranu prirody i krajiny v zemi Moravskoslezske 8: 1 - 108. [in Czech, French summary]" type="journal article" year="1948">Šilhavý 1948: 9</bibRefCitation>
</treatmentCitation>
, 24, table 6, figs 46.
<emphasis id="0093EAD7FFEBBB107382FB62CB80FB47" bold="true" box="[905,934,1245,1271]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB1073A5FB62CCEDFB47" author="Martens J." box="[942,1227,1245,1271]" page="408" pageId="4" pageNumber="5" year="1978">
<bibRefCitation id="56764B34FFEBBB1073A5FB62CCEDFB47" author="Martens J." box="[942,1227,1245,1271]" pageId="4" pageNumber="5" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978: 408412</bibRefCitation>
</treatmentCitation>
, figs 778780, 788.
<emphasis id="0093EAD7FFEBBB1070F3FABFC933FAAA" bold="true" box="[248,277,1280,1306]" pageId="4" pageNumber="5"></emphasis>
<treatmentCitation id="B34610D4FFEBBB107117FABFCA1FFAA9" author="Komposch C." box="[284,569,1279,1306]" page="26" pageId="4" pageNumber="5" year="1998">
<bibRefCitation id="56764B34FFEBBB107117FABFCA1FFAA9" author="Komposch C." box="[284,569,1279,1306]" pageId="4" pageNumber="5" pagination="19 - 40" refId="ref12699" refString="Komposch C. 1998. Leiobunum subalpinum n. sp. - ein neuer Weberknecht aus den Ostalpen (Opiliones: Phalangiidae). Wissenschaftliche Mitteilungen aus dem Nationalpark Hohe Tauern 4: 19 - 40." type="journal article" year="1998">Komposch 1998: 2627</bibRefCitation>
</treatmentCitation>
, 34, figs 5, 10, 14, 21.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325836C5FFEBBB1070B6FAFEC980FAEB" blockId="4.[189,422,1345,1371]" box="[189,422,1345,1371]" pageId="4" pageNumber="5">
<heading id="691081A9FFEBBB1070B6FAFEC980FAEB" bold="true" box="[189,422,1345,1371]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="0093EAD7FFEBBB1070B6FAFEC980FAEB" bold="true" box="[189,422,1345,1371]" pageId="4" pageNumber="5">Taxonomic history</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7AFD654EFFEBBB1170B6FACFC9C3FE3F" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="325836C5FFEBBB1070B6FACFC8DCF881" blockId="4.[189,1399,1392,1841]" pageId="4" pageNumber="5">
At first glance rather complex, but many distributional records from East central, Northwest and North Europe in reality refer to
<taxonomicName id="F5E74D46FFEBBB1071E7FA2CCA56FA1D" authorityName="C. L. Koch" authorityYear="1839" box="[492,624,1427,1453]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="0093EAD7FFEBBB1071E7FA2CCA56FA1D" box="[492,624,1427,1453]" italics="true" pageId="4" pageNumber="5">Leiobunum</emphasis>
</taxonomicName>
<taxonomicName id="F5E74D46FFEBBB107289FA2BCAE9FA1D" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[642,719,1428,1453]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFEBBB107289FA2BCAE9FA1D" box="[642,719,1428,1453]" italics="true" pageId="4" pageNumber="5">tisciae</emphasis>
</taxonomicName>
(see below,
<taxonomicName id="F5E74D46FFEBBB107367FA2BCBC6FA1D" authorityName="Thorell" authorityYear="1876" box="[876,992,1427,1453]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFEBBB107367FA2BCBC6FA1D" box="[876,992,1427,1453]" italics="true" pageId="4" pageNumber="5">L. gracile</emphasis>
</taxonomicName>
).
<taxonomicName id="F5E74D46FFEBBB1073F2FA2BCC5CFA1D" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[1017,1146,1428,1453]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB1073F2FA2BCC5CFA1D" box="[1017,1146,1428,1453]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
is a montane species and well known from low to mid-altitudes of the central and eastern Alps and mountainous areas in the
<collectingCountry id="4AF07655FFEBBB1070B6FA66C950FA43" box="[189,374,1497,1523]" name="Czech Republic" pageId="4" pageNumber="5">Czech Republic</collectingCountry>
and north of the Alps in
<collectingCountry id="4AF07655FFEBBB107298FA66CB24FA43" box="[659,770,1497,1523]" name="Germany" pageId="4" pageNumber="5">Germany</collectingCountry>
. Its genital morphology is quite characteristic and was first reliably depicted by
<bibRefCitation id="56764B34FFEBBB1071D6FA44CAA4F9A6" author="Silhavy V." box="[477,642,1531,1559]" pageId="4" pageNumber="5" pagination="1 - 108" refId="ref14107" refString="Silhavy V. 1948. Les Opilions de la steppe de Serpentine pres de la ville de Mohelno. Mohelno. Archiv Svazu na vyzkum a ochranu prirody i krajiny v zemi Moravskoslezske 8: 1 - 108. [in Czech, French summary]" type="journal article" year="1948">Šilhavý (1948</bibRefCitation>
, figs 46; 1956, figs 437439), fully in accordance with drawings in
<bibRefCitation id="56764B34FFEBBB1070D1F99FC9A1F98A" author="Martens J." box="[218,391,1568,1594]" pageId="4" pageNumber="5" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens (1978</bibRefCitation>
, fig. 778) and
<bibRefCitation id="56764B34FFEBBB107221F9A0CAD0F98A" author="Komposch C." box="[554,758,1567,1594]" pageId="4" pageNumber="5" pagination="19 - 40" refId="ref12699" refString="Komposch C. 1998. Leiobunum subalpinum n. sp. - ein neuer Weberknecht aus den Ostalpen (Opiliones: Phalangiidae). Wissenschaftliche Mitteilungen aus dem Nationalpark Hohe Tauern 4: 19 - 40." type="journal article" year="1998">Komposch (1998</bibRefCitation>
, figs 910).
<taxonomicName id="F5E74D46FFEBBB107388F99FCC21F98A" authorityName="C. L. Koch" authorityYear="1839" box="[899,1031,1568,1594]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="0093EAD7FFEBBB107388F99FCC21F98A" box="[899,1031,1568,1594]" italics="true" pageId="4" pageNumber="5">Leiobunum</emphasis>
</taxonomicName>
populations in the Carpathians,
<collectingCountry id="4AF07655FFEBBB1070B6F9FCC928F9ED" box="[189,270,1603,1629]" name="Poland" pageId="4" pageNumber="5">Poland</collectingCountry>
(
<bibRefCitation id="56764B34FFEBBB107114F9FCC999F9EE" author="Starega W." box="[287,447,1603,1630]" pageId="4" pageNumber="5" pagination="1 - 197" refId="ref14305" refString="Starega W. 1976. Opiliones Kosarze (Arachnoidea). Fauna Polski 5: 1 - 197, Polska Akademia Nauk, Warsaw." type="journal article" year="1976">Staręga 1976</bibRefCitation>
), southern
<collectingCountry id="4AF07655FFEBBB107248F9FCCABBF9ED" box="[579,669,1603,1629]" name="Finland" pageId="4" pageNumber="5">Finland</collectingCountry>
(
<bibRefCitation id="56764B34FFEBBB1072A6F9FCCB4FF9ED" author="Heinajoki M." box="[685,873,1603,1629]" pageId="4" pageNumber="5" pagination="1 - 26" refId="ref11956" refString="Heinajoki M. 1944. Die Opilionidenfauna Finnlands. Acta zoologica Fennica 42: 1 - 26." type="journal article" year="1944">Heinäjoki 1944</bibRefCitation>
), along the Baltic coast hinterland and
<collectingCountry id="4AF07655FFEBBB107537F9FCC8DFF930" name="Denmark" pageId="4" pageNumber="5">Denmark</collectingCountry>
were also largely affiliated to
<taxonomicName id="F5E74D46FFEBBB10725DF9D8CAF2F930" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[598,724,1639,1664]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB10725DF9D8CAF2F930" box="[598,724,1639,1664]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
(
<bibRefCitation id="56764B34FFEBBB1072E9F9D9CBA1F930" author="Martens J." box="[738,903,1638,1664]" pageId="4" pageNumber="5" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978</bibRefCitation>
, fig. 787), though erroneously. These older determinations mostly neglected the genital morphology of these crucial northern populations, which was reconsidered after
<bibRefCitation id="56764B34FFEBBB1071C7F912CA54F977" author="Avram S." box="[460,626,1708,1735]" pageId="4" pageNumber="5" pagination="111 - 125" refId="ref10967" refString="Avram S. 1968. Contribution a l'etude des Opilionides de la zone inondable de la Tisa entre les kilometres fluviaux 698 - 744. Avec la description d'une nouvelle espece: Leiobunum tisciae. Tiscia, Szeged 4: 111 - 125." type="journal article" year="1968">Avram (1968)</bibRefCitation>
published a new species,
<taxonomicName id="F5E74D46FFEBBB1073AFF912CC37F976" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[932,1041,1709,1734]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFEBBB1073AFF912CC37F976" box="[932,1041,1709,1734]" italics="true" pageId="4" pageNumber="5">L. tisciae</emphasis>
</taxonomicName>
, clearly distinct from
<taxonomicName id="F5E74D46FFEBBB10751DF912CD51F976" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[1302,1399,1709,1734]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB10751DF912CD51F976" box="[1302,1399,1709,1734]" italics="true" pageId="4" pageNumber="5">rupestre</emphasis>
</taxonomicName>
but similar in external morphology. Due to its wide distribution, the majority of available names for the eastern and northern populations can be assigned to the synonym of its oldest name
<taxonomicName id="F5E74D46FFEBBB107494F94BCD34F8BD" authorityName="Thorell" authorityYear="1876" box="[1183,1298,1779,1805]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFEBBB107494F94BCD34F8BD" box="[1183,1298,1779,1805]" italics="true" pageId="4" pageNumber="5">L. gracile</emphasis>
</taxonomicName>
(see below).
</paragraph>
<paragraph id="325836C5FFEBBB1170B6F8E8C9C3FE3F" blockId="4.[189,1399,1879,2012]" lastBlockId="5.[189,1399,267,399]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Excluding the complex synonymy of
<taxonomicName id="F5E74D46FFEBBB107280F8E7CADDF8C1" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[651,763,1880,1905]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFEBBB107280F8E7CADDF8C1" box="[651,763,1880,1905]" italics="true" pageId="4" pageNumber="5">L. tisciae</emphasis>
</taxonomicName>
(
<taxonomicName id="F5E74D46FFEBBB10731BF8E8CB45F8C1" authorityName="Thorell" authorityYear="1876" box="[784,867,1879,1905]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFEBBB10731BF8E8CB45F8C1" box="[784,867,1879,1905]" italics="true" pageId="4" pageNumber="5">gracile</emphasis>
</taxonomicName>
), only one uncertain affiliation remains for
<taxonomicName id="F5E74D46FFEBBB1070B6F8C4C967F824" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[189,321,1915,1940]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB1070B6F8C4C967F824" box="[189,321,1915,1940]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
: the name
<taxonomicName id="F5E74D46FFEBBB1071C1F8C4CB49F824" authority="L. Koch, 1869" authorityName="L. Koch" authorityYear="1869" box="[458,879,1914,1941]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFEBBB1071C1F8C4CA97F825" box="[458,689,1915,1941]" italics="true" pageId="4" pageNumber="5">Liobunum glabrum</emphasis>
L.
<bibRefCitation id="56764B34FFEBBB1072D5F8C4CB49F824" author="Koch L." box="[734,879,1914,1941]" pageId="4" pageNumber="5" pagination="149 - 206" refId="ref12226" refString="Koch L. 1869. Beitrag zur Kenntniss der Arachnidenfauna Tirols. Zeitschrift des Ferdinandeums fur Tirol und Vorarlberg, Innsbruck, series 3 (14): 149 - 206." type="journal article" year="1869">Koch, 1869</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="56764B34FFEBBB107370F8C6CC0CF825" author="Silhavy V." box="[891,1066,1913,1941]" pageId="4" pageNumber="5" refId="ref14153" refString="Silhavy V. 1956. Sekaci - Opilionidea. Fauna CSR 7. Akademia Nauk, Praha. [in Czech]" type="book" year="1956">Šilhavý (1956</bibRefCitation>
: fig. 443) originally treated
<emphasis id="0093EAD7FFEBBB1070CCF821C907F808" box="[199,289,1950,1976]" italics="true" pageId="4" pageNumber="5">glabra</emphasis>
as the juvenile
<taxonomicName id="F5E74D46FFEBBB1071EEF820CA4EF808" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[485,616,1951,1976]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB1071EEF820CA4EF808" box="[485,616,1951,1976]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
, but later used this name to distinguish allopatric populations of western
<taxonomicName id="F5E74D46FFEBBB10712BF87DC987F86B" baseAuthorityName="Herbst" baseAuthorityYear="1799" box="[288,417,1986,2011]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEBBB10712BF87DC987F86B" box="[288,417,1986,2011]" italics="true" pageId="4" pageNumber="5">L. rupestre</emphasis>
</taxonomicName>
and eastern
<emphasis id="0093EAD7FFEBBB10723CF87DCA69F86B" box="[567,591,1986,2011]" italics="true" pageId="4" pageNumber="5">L.</emphasis>
<taxonomicName id="F5E74D46FFEBBB10726AF87ECA89F86B" box="[609,687,1985,2011]" class="Arachnida" family="Gagrellidae" genus="Nelima" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0093EAD7FFEBBB10726AF87ECA89F86B" box="[609,687,1985,2011]" italics="true" pageId="4" pageNumber="5">glabra</emphasis>
</taxonomicName>
(later named
<taxonomicName id="F5E74D46FFEBBB107350F87DCBEEF86B" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[859,968,1986,2011]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFEBBB107350F87DCBEEF86B" box="[859,968,1986,2011]" italics="true" pageId="4" pageNumber="5">L. tisciae</emphasis>
</taxonomicName>
) in former
<collectingCountry id="4AF07655FFEBBB10745AF87ECD2DF86B" box="[1105,1291,1985,2011]" name="Czech Republic" pageId="4" pageNumber="5">Czechoslovakia</collectingCountry>
(
<bibRefCitation id="56764B34FFEBBB117510F800C8DBFE95" author="Silhavy V." lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" pagination="204 - 208" refId="ref14175" refString="Silhavy V. 1981. Occurrence of Leiobunum glabrum in Czechoslovakia (Arach., Opilionidea). Vestnik ceskoslovenske Spolecnosti zoologicke [Acta societatis zoologicae bohemoslovacae] 45 (3): 204 - 208." type="journal article" year="1981">Šilhavý 1981</bibRefCitation>
). This was contradicted by
<bibRefCitation id="56764B34FFEABB117257FEB4CB33FE95" author="Starega W." box="[604,789,267,293]" pageId="5" pageNumber="6" pagination="78 - 88" refId="ref14380" refString="Starega W. 2004. Interessante Weberknechtfunde aus Polen (Arachnida: Opiliones). Arachnologische Mitteilungen 27 / 28: 78 - 88." type="journal article" year="2004">Staręga (2004)</bibRefCitation>
, because the
<typeStatus id="ED5C8867FFEABB1173CAFEB3CBD3FE96" box="[961,1013,268,294]" pageId="5" pageNumber="6">type</typeStatus>
locality of
<taxonomicName id="F5E74D46FFEABB117480FEB3CD51FE96" authority=", Meran" authorityName="Meran" box="[1163,1399,267,294]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFEABB117480FEB3CD32FE95" box="[1163,1300,267,293]" italics="true" pageId="5" pageNumber="6">L. glabrum</emphasis>
, Meran
</taxonomicName>
(northern
<collectingCountry id="4AF07655FFEABB11713EFE91C94BFEF8" box="[309,365,302,328]" name="Italy" pageId="5" pageNumber="6">Italy</collectingCountry>
, southern Alps), is situated deeply within the territory of
<taxonomicName id="F5E74D46FFEABB117423FE90CD55FEF9" authority="(Martens 1978)" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[1064,1395,303,329]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEABB117423FE90CC8AFEF8" box="[1064,1196,303,328]" italics="true" pageId="5" pageNumber="6">L. rupestre</emphasis>
(
<bibRefCitation id="56764B34FFEABB1174B7FE90CD4FFEF9" author="Martens J." box="[1212,1385,303,329]" pageId="5" pageNumber="6" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978</bibRefCitation>
)
</taxonomicName>
. We suggest following
<bibRefCitation id="56764B34FFEABB1171C2FEEDCA40FEDC" author="Hadzi J." box="[457,614,338,364]" pageId="5" pageNumber="6" pagination="107 - 154" refId="ref11762" refString="Hadzi J. 1931. Opilioni Triglavskoga masiva [Die Opilionen des Triglavmassivs]. Prirodoslovne Razprave (Muzejsko Drustvo za Slovenijo) Ljubljana 1: 107 - 154. [in Slovenian, German abstract]" type="journal article" year="1931">Hadži (1931)</bibRefCitation>
and
<bibRefCitation id="56764B34FFEABB1172A9FEEDCB7CFEDC" author="Martens J." box="[674,858,338,364]" pageId="5" pageNumber="6" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens (1978)</bibRefCitation>
to correctly place
<taxonomicName id="F5E74D46FFEABB117431FEEDCD0CFEDC" authorityName="in Silhavy" authorityYear="1981" box="[1082,1322,338,364]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFEABB117431FEEDCD0CFEDC" box="[1082,1322,338,364]" italics="true" pageId="5" pageNumber="6">Leiobunum glabrum</emphasis>
</taxonomicName>
in the synonymy of
<taxonomicName id="F5E74D46FFEABB117155FEC9C9F8FE3F" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[350,478,374,399]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEABB117155FEC9C9F8FE3F" box="[350,478,374,399]" italics="true" pageId="5" pageNumber="6">L. rupestre</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7AFD654EFFEABB1C70B6FE09CC2CFE02" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="325836C5FFEABB1170B6FE09C910FE60" blockId="5.[189,310,438,464]" box="[189,310,438,464]" pageId="5" pageNumber="6">
<heading id="691081A9FFEABB1170B6FE09C910FE60" bold="true" box="[189,310,438,464]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="0093EAD7FFEABB1170B6FE09C910FE60" bold="true" box="[189,310,438,464]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="325836C5FFEABB1170B6FE5AC96EFD62" blockId="5.[189,1399,485,722]" pageId="5" pageNumber="6">
A medium-sized
<taxonomicName id="F5E74D46FFEABB117188FE5ACA21FE4F" authorityName="C. L. Koch" authorityYear="1839" box="[387,519,485,511]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="0093EAD7FFEABB117188FE5ACA21FE4F" box="[387,519,485,511]" italics="true" pageId="5" pageNumber="6">Leiobunum</emphasis>
</taxonomicName>
species with blackish upper side, except for broad, white markings on Ceph disto-laterally in male (
<figureCitation id="AADC2A40FFEABB1171C0FDB7CA0AFD92" box="[459,556,520,546]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[189,1398,767,1853]" captionTargetId="figure@5.[189,1398,767,1853]" captionTargetPageId="5" captionText="Fig. 1. Habit of Leiobunum rupestre species group. AB. Leiobunum rupestre Herbst, 1799, Slovenia, Pohorje Mountains, resting at rock faces. A. ♁. B. ♀. CD. Leiobunum apenninicum (Martens, 1969), Italy, Monesi di Triora, at night. C. ♁. D. ♀. EF. Leiobunum gracile Thorell, 1876, Denmark. E. ♁. F. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field." figureDoi="http://doi.org/10.5281/zenodo.3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="5" pageNumber="6">Figs 1A</figureCitation>
,
<figureCitation id="AADC2A40FFEABB11723CFDB7CA78FD92" box="[567,606,520,546]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[408,1184,1137,1848]" captionTargetId="figure@7.[391,1196,1119,1857]" captionTargetPageId="7" captionText="Fig. 3. Body of Leiobunum C.L. Koch, 1839, dorsal view. AB. Leiobunum rupestre Herbst, 1799, Germany, Mt. Arber, CJM136. A. ♁. B. ♀. — CD. Leiobunum gracile Thorell, 1876, Denmark. C. ♁, 2 km N of Skaerbaek, CJM3530. D. ♀, Asp, CJM3531. — EF. Leiobunum apenninicum (Martens, 1969), France, Alpes-Maritimes. E. ♁, CJM1508. F. ♀, CJM2747. Drawings by K. Rehbinder." figureDoi="http://doi.org/10.5281/zenodo.3840155" httpUri="https://zenodo.org/record/3840155/files/figure.png" pageId="5" pageNumber="6">3A</figureCitation>
), broad, blackish irregular saddle-like marking all over the length of the body in female with extended white markings laterally from Ceph to abd Area II and a cross stripe all over area V (
<figureCitation id="AADC2A40FFEABB117171FDF1C9FFFDD9" box="[378,473,590,617]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[189,1398,767,1853]" captionTargetId="figure@5.[189,1398,767,1853]" captionTargetPageId="5" captionText="Fig. 1. Habit of Leiobunum rupestre species group. AB. Leiobunum rupestre Herbst, 1799, Slovenia, Pohorje Mountains, resting at rock faces. A. ♁. B. ♀. CD. Leiobunum apenninicum (Martens, 1969), Italy, Monesi di Triora, at night. C. ♁. D. ♀. EF. Leiobunum gracile Thorell, 1876, Denmark. E. ♁. F. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field." figureDoi="http://doi.org/10.5281/zenodo.3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="5" pageNumber="6">Figs 1B</figureCitation>
,
<figureCitation id="AADC2A40FFEABB1171ECFDF0CA2AFDD9" box="[487,524,591,617]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[408,1184,1137,1848]" captionTargetId="figure@7.[391,1196,1119,1857]" captionTargetPageId="7" captionText="Fig. 3. Body of Leiobunum C.L. Koch, 1839, dorsal view. AB. Leiobunum rupestre Herbst, 1799, Germany, Mt. Arber, CJM136. A. ♁. B. ♀. — CD. Leiobunum gracile Thorell, 1876, Denmark. C. ♁, 2 km N of Skaerbaek, CJM3530. D. ♀, Asp, CJM3531. — EF. Leiobunum apenninicum (Martens, 1969), France, Alpes-Maritimes. E. ♁, CJM1508. F. ♀, CJM2747. Drawings by K. Rehbinder." figureDoi="http://doi.org/10.5281/zenodo.3840155" httpUri="https://zenodo.org/record/3840155/files/figure.png" pageId="5" pageNumber="6">3B</figureCitation>
). In both sexes series of para-median lines of small white spots on the abd areae IV are absent. Contrasting yellowish underside including coxae of all appendages. Wings of truncus penis broad, broadest in lower third part of truncus (
<figureCitation id="AADC2A40FFEABB117389FD2ACBEEFD00" box="[898,968,661,688]" captionStart="Fig" captionStartId="12.[189,232,1859,1885]" captionTargetBox="[197,1377,919,1811]" captionTargetId="figure@12.[189,1398,913,1815]" captionTargetPageId="12" captionText="Fig. 5. Leiobunum, male genitalia. AC. L. rupestre Herbst, 1799, Germany, Baden-Württemberg, CJM1954. A. Ventral view. B. Lateral view. C. Cross-section. — DF. L. gracile Thorell, 1876, Denmark, 2 km N of Skaerbaek, CJM3530. D. Ventral view. E. Lateral view. F. Cross-section. — GJ. L. apenninicum (Martens, 1969), France, Alpes-Maritimes, CJM1508. G. Ventral view. H. Lateral view. J. Cross-section. Arrows indicate the area of the respective cross-sections." figureDoi="http://doi.org/10.5281/zenodo.3840161" httpUri="https://zenodo.org/record/3840161/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
AB). Coxa IV with retrolateral row of granules.
</paragraph>
<caption id="6698664DFFEABB1170B6F8D9CC59F859" ID-DOI="http://doi.org/10.5281/zenodo.3840151" ID-Zenodo-Dep="3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1894,1920]" targetBox="[189,1398,767,1853]" targetPageId="5">
<paragraph id="325836C5FFEABB1170B6F8D9CC59F859" blockId="5.[189,1399,1894,2026]" pageId="5" pageNumber="6">
<emphasis id="0093EAD7FFEABB1170B6F8D9C92AF830" bold="true" box="[189,268,1894,1920]" pageId="5" pageNumber="6">Fig. 1.</emphasis>
Habit of
<taxonomicName id="F5E74D46FFEABB117175F8D9CA4DF830" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[382,619,1894,1920]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEABB117175F8D9CA4DF830" box="[382,619,1894,1920]" italics="true" pageId="5" pageNumber="6">Leiobunum rupestre</emphasis>
</taxonomicName>
species group.
<emphasis id="0093EAD7FFEABB11732FF8D9CB7DF830" bold="true" box="[804,859,1894,1920]" pageId="5" pageNumber="6">AB</emphasis>
.
<taxonomicName id="F5E74D46FFEABB117360F8D9CCDDF830" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" box="[875,1275,1894,1920]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFEABB117360F8D9CC7EF830" box="[875,1112,1894,1920]" italics="true" pageId="5" pageNumber="6">Leiobunum rupestre</emphasis>
<bibRefCitation id="56764B34FFEABB11746BF8D9CCDDF830" author="Herbst J. F. W." box="[1120,1275,1894,1920]" pageId="5" pageNumber="6" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst, 1799</bibRefCitation>
</taxonomicName>
, Slovenia, Pohorje Mountains, resting at rock faces.
<emphasis id="0093EAD7FFEABB1172A0F836CAE6F813" bold="true" box="[683,704,1929,1955]" pageId="5" pageNumber="6">A</emphasis>
. ♁.
<emphasis id="0093EAD7FFEABB1172FEF836CB2FF813" bold="true" box="[757,777,1929,1955]" pageId="5" pageNumber="6">B</emphasis>
. ♀.
<emphasis id="0093EAD7FFEABB117336F836CB50F813" bold="true" box="[829,886,1929,1955]" pageId="5" pageNumber="6">CD</emphasis>
.
<taxonomicName id="F5E74D46FFEABB11738DF836CD54F814" authority="(Martens, 1969)" baseAuthorityName="Martens" baseAuthorityYear="1969" box="[902,1394,1929,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="apenninicum">
<emphasis id="0093EAD7FFEABB11738DF836CC8CF813" box="[902,1194,1929,1955]" italics="true" pageId="5" pageNumber="6">Leiobunum apenninicum</emphasis>
(
<bibRefCitation id="56764B34FFEABB1174B2F836CD4FF814" author="Martens J." box="[1209,1385,1929,1956]" pageId="5" pageNumber="6" pagination="395 - 415" refId="ref13383" refString="Martens J. 1969. Mittel- und sudeuropaische Arten der Gattung Nelima (Arachnida: Opiliones: Leiobunidae). Senckenbergiana biologica 50 (5 / 6): 395 - 415." type="journal article" year="1969">Martens, 1969</bibRefCitation>
)
</taxonomicName>
, Italy, Monesi di Triora, at night.
<emphasis id="0093EAD7FFEABB117248F813CA7EF876" bold="true" box="[579,600,1964,1990]" pageId="5" pageNumber="6">C</emphasis>
. ♁.
<emphasis id="0093EAD7FFEABB117286F812CA84F877" bold="true" box="[653,674,1965,1991]" pageId="5" pageNumber="6">D</emphasis>
. ♀.
<emphasis id="0093EAD7FFEABB1172DCF812CB2AF877" bold="true" box="[727,780,1965,1991]" pageId="5" pageNumber="6">EF</emphasis>
.
<taxonomicName id="F5E74D46FFEABB117310F813CC8AF876" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[795,1196,1964,1990]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFEABB117310F813CBD8F876" box="[795,1022,1964,1990]" italics="true" pageId="5" pageNumber="6">Leiobunum gracile</emphasis>
<bibRefCitation id="56764B34FFEABB11740EF813CC8AF876" author="Thorell T." box="[1029,1196,1964,1990]" pageId="5" pageNumber="6" pagination="452 - 508" refId="ref14658" refString="Thorell T. 1876. Sopra alcuni Opilioni (Phalangidea) d'Europa e dell'Asia occidentale, con un quadro dei generi europei di quest'Ordine. Annali del Museo civico di storia naturale di Genova 8: 452 - 508. Available form http: // biodiversitylibrary. org / page / 7916212 [accessed 23 Jun. 2016]" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
, Denmark.
<emphasis id="0093EAD7FFEABB11753DF812CD6CF877" bold="true" box="[1334,1354,1965,1991]" pageId="5" pageNumber="6">E</emphasis>
. ♁.
<emphasis id="0093EAD7FFEABB1170B6F86FC8E9F85A" bold="true" box="[189,207,2000,2026]" pageId="5" pageNumber="6">F</emphasis>
. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field.
</paragraph>
</caption>
<paragraph id="325836C5FFE9BB1270B6FEB3CA87FEDC" blockId="6.[189,1398,267,364]" pageId="6" pageNumber="7">
BODY (
<figureCitation id="AADC2A40FFE9BB127100FEB4C968FE95" box="[267,334,267,293]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[189,1398,767,1853]" captionTargetId="figure@5.[189,1398,767,1853]" captionTargetPageId="5" captionText="Fig. 1. Habit of Leiobunum rupestre species group. AB. Leiobunum rupestre Herbst, 1799, Slovenia, Pohorje Mountains, resting at rock faces. A. ♁. B. ♀. CD. Leiobunum apenninicum (Martens, 1969), Italy, Monesi di Triora, at night. C. ♁. D. ♀. EF. Leiobunum gracile Thorell, 1876, Denmark. E. ♁. F. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field." figureDoi="http://doi.org/10.5281/zenodo.3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="6" pageNumber="7">Figs 1</figureCitation>
AB, 3AB). In both sexes dorsum smooth, minute granulation not even to be distinguished with magnitude 250 (compare
<bibRefCitation id="56764B34FFE9BB12722EFE92CAE3FEF9" author="Silhavy V." box="[549,709,301,329]" pageId="6" pageNumber="7" pagination="204 - 208" refId="ref14175" refString="Silhavy V. 1981. Occurrence of Leiobunum glabrum in Czechoslovakia (Arach., Opilionidea). Vestnik ceskoslovenske Spolecnosti zoologicke [Acta societatis zoologicae bohemoslovacae] 45 (3): 204 - 208." type="journal article" year="1981">Šilhavý 1981</bibRefCitation>
: fig. 4), few scattered pointed granules present; eye mound in both sexes without spines or tubercles.
</paragraph>
<paragraph id="325836C5FFE9BB1270B6FE2BCD76FE1E" blockId="6.[189,1360,403,430]" box="[189,1360,403,430]" pageId="6" pageNumber="7">
DORSAL PATTERN. Male and female (
<figureCitation id="AADC2A40FFE9BB127253FE2CCAB8FE1D" box="[600,670,403,429]" captionStart="Fig" captionStartId="5.[189,232,1894,1920]" captionTargetBox="[189,1398,767,1853]" captionTargetId="figure@5.[189,1398,767,1853]" captionTargetPageId="5" captionText="Fig. 1. Habit of Leiobunum rupestre species group. AB. Leiobunum rupestre Herbst, 1799, Slovenia, Pohorje Mountains, resting at rock faces. A. ♁. B. ♀. CD. Leiobunum apenninicum (Martens, 1969), Italy, Monesi di Triora, at night. C. ♁. D. ♀. EF. Leiobunum gracile Thorell, 1876, Denmark. E. ♁. F. ♀. Photographs: AD by A.L.Schönhofer; EF by S. Toft, all taken in the field." figureDoi="http://doi.org/10.5281/zenodo.3840151" httpUri="https://zenodo.org/record/3840151/files/figure.png" pageId="6" pageNumber="7">Figs 1</figureCitation>
AB, 3AB): contrasting black and white (see Diagnosis).
</paragraph>
<paragraph id="325836C5FFE9BB1270B6FE6ACC95FD2F" blockId="6.[189,1399,469,671]" pageId="6" pageNumber="7">
PEDIPALPS. Male (
<figureCitation id="AADC2A40FFE9BB127186FE6AC9F0FE5F" box="[397,470,469,495]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[199,1377,266,1667]" captionTargetId="figure@10.[199,1388,259,1671]" captionTargetPageId="10" captionText="Fig. 4. Pedipalps of the Leiobunum rupestregroup, wholepedipalps inlateral view, single femora inmedial view. — AF. ♁♁. AB. Leiobunum rupestre Herbst, 1799, Germany, Baden-Württemberg, CJM1954. A. Pedipalpus lateral. B. Femur medial. CD. L. gracile Thorell, 1876, Denmark, Asp, CJM3531. C. Pedipalpus lateral. D. Femur medial. — EF. Leiobunum apenninicum (Martens, 1969), France, Alpes- Maritimes, CJM2747. E. Pedipalpus lateral. F. Femur medial. — GM. ♀♀. GH. Leiobunum rupestre Herbst, 1799, Germany, Mt. Arber, CJM136. G. Pedipalpus lateral. H. Femur medial. JK. L. gracile Thorell, 1876, Denmark, Asp, CJM3531. J. Pedipalpus lateral. K. Femur medial. LM. L. apenninicum (Martens, 1969), France, Alpes-Maritimes, CJM2747. L. Pedipalpus lateral. M. Femur medial. Arrows indicate characters mentioned in the descriptions." figureDoi="http://doi.org/10.5281/zenodo.3840157" httpUri="https://zenodo.org/record/3840157/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
AB):
<emphasis id="0093EAD7FFE9BB127221FE6ACA6FFE5F" bold="true" box="[554,585,469,495]" pageId="6" pageNumber="7">Fe</emphasis>
with irregular rows of marked triangular spines ventrally; dorsally nearly unarmed, ventral spines more slender than in
<taxonomicName id="F5E74D46FFE9BB12732AFE46CBB5FDA2" authorityName="Thorell" authorityYear="1876" box="[801,915,504,530]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE9BB12732AFE46CBB5FDA2" box="[801,915,504,530]" italics="true" pageId="6" pageNumber="7">L. gracile</emphasis>
</taxonomicName>
;
<emphasis id="0093EAD7FFE9BB1273A9FE47CB98FDA2" bold="true" box="[930,958,504,530]" pageId="6" pageNumber="7">Pt</emphasis>
slightly armed with pointed denticles laterally;
<emphasis id="0093EAD7FFE9BB127139FDA4C96BFD85" bold="true" box="[306,333,539,565]" pageId="6" pageNumber="7">Ti</emphasis>
markedly convex in basal part and concave at ventro-distal end, therefore forming a well-marked S-bend, field of fine dark coloured granules all over the ventral side, well to be seen;
<emphasis id="0093EAD7FFE9BB12755CFD81CD52FDE8" bold="true" box="[1367,1396,574,600]" pageId="6" pageNumber="7">Ta</emphasis>
continuously and slightly bent to ventral. Female (
<figureCitation id="AADC2A40FFE9BB12730CFDDECB6AFDCC" box="[775,844,609,636]" captionStart="Fig" captionStartId="10.[189,232,1718,1744]" captionTargetBox="[199,1377,266,1667]" captionTargetId="figure@10.[199,1388,259,1671]" captionTargetPageId="10" captionText="Fig. 4. Pedipalps of the Leiobunum rupestregroup, wholepedipalps inlateral view, single femora inmedial view. — AF. ♁♁. AB. Leiobunum rupestre Herbst, 1799, Germany, Baden-Württemberg, CJM1954. A. Pedipalpus lateral. B. Femur medial. CD. L. gracile Thorell, 1876, Denmark, Asp, CJM3531. C. Pedipalpus lateral. D. Femur medial. — EF. Leiobunum apenninicum (Martens, 1969), France, Alpes- Maritimes, CJM2747. E. Pedipalpus lateral. F. Femur medial. — GM. ♀♀. GH. Leiobunum rupestre Herbst, 1799, Germany, Mt. Arber, CJM136. G. Pedipalpus lateral. H. Femur medial. JK. L. gracile Thorell, 1876, Denmark, Asp, CJM3531. J. Pedipalpus lateral. K. Femur medial. LM. L. apenninicum (Martens, 1969), France, Alpes-Maritimes, CJM2747. L. Pedipalpus lateral. M. Femur medial. Arrows indicate characters mentioned in the descriptions." figureDoi="http://doi.org/10.5281/zenodo.3840157" httpUri="https://zenodo.org/record/3840157/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
GH): similar to male, less spines on Fe and Pt, few spines on Ti proximally and no ventral granulation, Ta strait, slightly bent distally.
</paragraph>
<paragraph id="325836C5FFE9BB1270B6FD79CD51FC66" blockId="6.[189,1399,710,983]" pageId="6" pageNumber="7">
GENITAL MORPHOLOGY (
<figureCitation id="AADC2A40FFE9BB1271C3FD79CA29FD51" box="[456,527,710,737]" captionStart="Fig" captionStartId="12.[189,232,1859,1885]" captionTargetBox="[197,1377,919,1811]" captionTargetId="figure@12.[189,1398,913,1815]" captionTargetPageId="12" captionText="Fig. 5. Leiobunum, male genitalia. AC. L. rupestre Herbst, 1799, Germany, Baden-Württemberg, CJM1954. A. Ventral view. B. Lateral view. C. Cross-section. — DF. L. gracile Thorell, 1876, Denmark, 2 km N of Skaerbaek, CJM3530. D. Ventral view. E. Lateral view. F. Cross-section. — GJ. L. apenninicum (Martens, 1969), France, Alpes-Maritimes, CJM1508. G. Ventral view. H. Lateral view. J. Cross-section. Arrows indicate the area of the respective cross-sections." figureDoi="http://doi.org/10.5281/zenodo.3840161" httpUri="https://zenodo.org/record/3840161/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
AC). Truncus penis stout, in ventral/dorsal view from basal opening to insertion of glans slightly and continuously tapering; distal wings covering less than half of truncus. Wings consist of two independent parts: a ventral shell-like plate, sharply cut off horizontally at upper third of wing structure, from its upper margin continuously extending to lateral side of truncus, thus partly embracing the lateral wings which extend from dorsal side of truncus via lateral side to ventral side. Their lower two thirds are covered by the ventral plate. Surface of the upper margin of the lateral wings and membranes adjacent to the truncus is invaginated forming a long double-walled internal sack. At the very distal end of truncus a small membranous oval sack with a distal opening is attached ventrally.
</paragraph>
<paragraph id="325836C5FFE9BB1270B6FC41CD28FB8B" blockId="6.[189,1399,1022,1084]" pageId="6" pageNumber="7">From lateral view truncus slightly curved (concave on ventral side), slightly tapering towards glans. Wings massively enlarged on ventral side. Glans stout, in lateral view tapering towards stylus.</paragraph>
<caption id="6698664DFFE9BB1270B6F89FCC3DF85B" ID-DOI="http://doi.org/10.5281/zenodo.3840153" ID-Zenodo-Dep="3840153" httpUri="https://zenodo.org/record/3840153/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1824,1850]" targetBox="[186,1399,1156,1766]" targetPageId="6">
<paragraph id="325836C5FFE9BB1270B6F89FCC3DF85B" blockId="6.[189,1399,1824,2027]" pageId="6" pageNumber="7">
<emphasis id="0093EAD7FFE9BB1270B6F89FC92DF88A" bold="true" box="[189,267,1824,1850]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
Distribution of the species of the
<taxonomicName id="F5E74D46FFE9BB1272ABF89ECBABF88A" authorityName="Herbst" authorityYear="1799" box="[672,909,1825,1851]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE9BB1272ABF89ECBABF88A" box="[672,909,1825,1851]" italics="true" pageId="6" pageNumber="7">Leiobunum rupestre</emphasis>
</taxonomicName>
species group. Red squares =
<taxonomicName id="F5E74D46FFE9BB1274FDF89EC971F8EE" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE9BB1274FDF89ECD51F88A" box="[1270,1399,1825,1850]" italics="true" pageId="6" pageNumber="7">L. rupestre</emphasis>
<bibRefCitation id="56764B34FFE9BB1270B6F8FBC971F8EE" author="Herbst J. F. W." box="[189,343,1860,1886]" pageId="6" pageNumber="7" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst, 1799</bibRefCitation>
</taxonomicName>
; blue circles =
<taxonomicName id="F5E74D46FFE9BB12720DF8FACB06F8EE" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[518,800,1860,1886]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE9BB12720DF8FACA51F8EE" box="[518,631,1860,1886]" italics="true" pageId="6" pageNumber="7">L. gracile</emphasis>
<bibRefCitation id="56764B34FFE9BB127276F8FBCB06F8EE" author="Thorell T." box="[637,800,1860,1886]" pageId="6" pageNumber="7" pagination="452 - 508" refId="ref14658" refString="Thorell T. 1876. Sopra alcuni Opilioni (Phalangidea) d'Europa e dell'Asia occidentale, con un quadro dei generi europei di quest'Ordine. Annali del Museo civico di storia naturale di Genova 8: 452 - 508. Available form http: // biodiversitylibrary. org / page / 7916212 [accessed 23 Jun. 2016]" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
; green triangles =
<taxonomicName id="F5E74D46FFE9BB1273FDF8FACD54F8EF" authority="(Martens, 1969)" baseAuthorityName="Martens" baseAuthorityYear="1969" box="[1014,1394,1860,1887]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="apenninicum">
<emphasis id="0093EAD7FFE9BB1273FDF8FACC8AF8EE" box="[1014,1196,1861,1886]" italics="true" pageId="6" pageNumber="7">L. apenninicum</emphasis>
(
<bibRefCitation id="56764B34FFE9BB1274B1F8FBCD4FF8EF" author="Martens J." box="[1210,1385,1860,1887]" pageId="6" pageNumber="7" pagination="395 - 415" refId="ref13383" refString="Martens J. 1969. Mittel- und sudeuropaische Arten der Gattung Nelima (Arachnida: Opiliones: Leiobunidae). Senckenbergiana biologica 50 (5 / 6): 395 - 415." type="journal article" year="1969">Martens, 1969</bibRefCitation>
)
</taxonomicName>
; stippled line circumscribes area of
<taxonomicName id="F5E74D46FFE9BB127263F8D7CBCBF831" authority="Komposch, 1998" authorityName="Komposch" authorityYear="1998" box="[616,1005,1895,1921]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="subalpinum">
<emphasis id="0093EAD7FFE9BB127263F8D7CB35F831" box="[616,787,1895,1921]" italics="true" pageId="6" pageNumber="7">L. subalpinum</emphasis>
<bibRefCitation id="56764B34FFE9BB127316F8D8CBCBF831" author="Komposch C." box="[797,1005,1895,1921]" pageId="6" pageNumber="7" pagination="19 - 40" refId="ref12699" refString="Komposch C. 1998. Leiobunum subalpinum n. sp. - ein neuer Weberknecht aus den Ostalpen (Opiliones: Phalangiidae). Wissenschaftliche Mitteilungen aus dem Nationalpark Hohe Tauern 4: 19 - 40." type="journal article" year="1998">Komposch, 1998</bibRefCitation>
</taxonomicName>
. Type localities are indicated by abbreviated taxon names (ape =
<taxonomicName id="F5E74D46FFE9BB127227F834CAE2F814" baseAuthorityName="Martens" baseAuthorityYear="1969" box="[556,708,1931,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="apenninicum">
<emphasis id="0093EAD7FFE9BB127227F834CAE2F814" box="[556,708,1931,1956]" italics="true" pageId="6" pageNumber="7">apenninicum</emphasis>
</taxonomicName>
; gla =
<taxonomicName id="F5E74D46FFE9BB127305F835CB57F814" authorityName="in Silhavy" authorityYear="1981" box="[782,881,1930,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="glabrum">
<emphasis id="0093EAD7FFE9BB127305F835CB57F814" box="[782,881,1930,1956]" italics="true" pageId="6" pageNumber="7">glabrum</emphasis>
</taxonomicName>
; gra =
<taxonomicName id="F5E74D46FFE9BB1273B6F835CC36F814" authorityName="Thorell" authorityYear="1876" box="[957,1040,1930,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE9BB1273B6F835CC36F814" box="[957,1040,1930,1956]" italics="true" pageId="6" pageNumber="7">gracile</emphasis>
</taxonomicName>
; lae =
<taxonomicName id="F5E74D46FFE9BB127453F835CCB0F814" authorityName="Thorell" authorityYear="1876" box="[1112,1174,1930,1956]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laeve">
<emphasis id="0093EAD7FFE9BB127453F835CCB0F814" box="[1112,1174,1930,1956]" italics="true" pageId="6" pageNumber="7">laeve</emphasis>
</taxonomicName>
; nor =
<taxonomicName id="F5E74D46FFE9BB1274EFF834CD48F814" authorityName="Strand" authorityYear="1900" box="[1252,1390,1931,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="norvegicum">
<emphasis id="0093EAD7FFE9BB1274EFF834CD48F814" box="[1252,1390,1931,1956]" italics="true" pageId="6" pageNumber="7">norvegicum</emphasis>
</taxonomicName>
; rup =
<taxonomicName id="F5E74D46FFE9BB127108F811C942F877" authorityName="Herbst" authorityYear="1799" box="[259,356,1966,1991]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE9BB127108F811C942F877" box="[259,356,1966,1991]" italics="true" pageId="6" pageNumber="7">rupestre</emphasis>
</taxonomicName>
; sub
<taxonomicName id="F5E74D46FFE9BB1271AFF812CA0BF877" authorityName="Komposch" authorityYear="1998" box="[420,557,1965,1991]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="subalpinum">
<emphasis id="0093EAD7FFE9BB1271AFF812CA0BF877" box="[420,557,1965,1991]" italics="true" pageId="6" pageNumber="7">subalpinum</emphasis>
</taxonomicName>
; tis =
<taxonomicName id="F5E74D46FFE9BB12727CF811CAE2F877" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[631,708,1966,1991]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFE9BB12727CF811CAE2F877" box="[631,708,1966,1991]" italics="true" pageId="6" pageNumber="7">tisciae</emphasis>
</taxonomicName>
). Arrows indicate a specified locality, if unprecisely given, the general area is indicated (
<taxonomicName id="F5E74D46FFE9BB12721DF86ECA51F85A" authorityName="Herbst" authorityYear="1799" box="[534,631,2001,2026]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE9BB12721DF86ECA51F85A" box="[534,631,2001,2026]" italics="true" pageId="6" pageNumber="7">rupestre</emphasis>
</taxonomicName>
= Saxony;
<taxonomicName id="F5E74D46FFE9BB1272F6F86ECB76F85B" authorityName="Thorell" authorityYear="1876" box="[765,848,2001,2027]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE9BB1272F6F86ECB76F85B" box="[765,848,2001,2027]" italics="true" pageId="6" pageNumber="7">gracile</emphasis>
</taxonomicName>
,
<taxonomicName id="F5E74D46FFE9BB127354F86ECBBBF85B" authorityName="Thorell" authorityYear="1876" box="[863,925,2001,2027]" class="Arachnida" family="Phalangiidae" genus="Liobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laeve">
<emphasis id="0093EAD7FFE9BB127354F86ECBBBF85B" box="[863,925,2001,2027]" italics="true" pageId="6" pageNumber="7">laeve</emphasis>
</taxonomicName>
= Scania).
</paragraph>
</caption>
<paragraph id="325836C5FFE8BB1370B6FEB4C990FE95" blockId="7.[189,438,267,293]" box="[189,438,267,293]" pageId="7" pageNumber="8">
<emphasis id="0093EAD7FFE8BB1370B6FEB4C970FE95" bold="true" box="[189,342,267,293]" pageId="7" pageNumber="8">Distribution</emphasis>
(
<figureCitation id="AADC2A40FFE8BB13716DFEB4C98BFE95" box="[358,429,267,293]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[186,1399,1156,1766]" captionTargetId="figure@6.[185,1400,1154,1767]" captionTargetPageId="6" captionText="Fig. 2. Distribution of the species of the Leiobunum rupestre species group. Red squares = L. rupestre Herbst, 1799; blue circles = L. gracile Thorell, 1876; green triangles = L. apenninicum (Martens, 1969); stippled line circumscribes area of L. subalpinum Komposch, 1998. Type localities are indicated by abbreviated taxon names (ape = apenninicum; gla = glabrum; gra = gracile; lae = laeve; nor = norvegicum; rup = rupestre; sub subalpinum; tis = tisciae). Arrows indicate a specified locality, if unprecisely given, the general area is indicated (rupestre = Saxony; gracile, laeve = Scania)." figureDoi="http://doi.org/10.5281/zenodo.3840153" httpUri="https://zenodo.org/record/3840153/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
)
</paragraph>
<paragraph id="325836C5FFE8BB1370B6FE85CA55FDB4" blockId="7.[189,1399,314,516]" pageId="7" pageNumber="8">
Main area are the eastern Alps from the eastern part of
<collectingCountry id="4AF07655FFE8BB13735FFE85CBC5FEE4" box="[852,995,314,340]" name="Switzerland" pageId="7" pageNumber="8">Switzerland</collectingCountry>
, all over
<collectingCountry id="4AF07655FFE8BB137459FE85CC8DFEE4" box="[1106,1195,314,340]" name="Austria" pageId="7" pageNumber="8">Austria</collectingCountry>
, the German and Slovenian alpine parts and extending to the north-western Balkan Peninsula. Records extend southwards to
<collectingCountry id="4AF07655FFE8BB1370D0FE3FC902FE2A" box="[219,292,384,410]" name="Serbia" pageId="7" pageNumber="8">Serbia</collectingCountry>
(CJM6363, CIK1336, CIK1337),
<collectingCountry id="4AF07655FFE8BB1372B8FE3FCB21FE2A" box="[691,775,384,410]" name="Croatia" pageId="7" pageNumber="8">Croatia</collectingCountry>
(
<bibRefCitation id="56764B34FFE8BB13731DFE3FCB86FE2A" author="Babic K." box="[790,928,384,410]" pageId="7" pageNumber="8" pagination="169 - 179" refId="ref11083" refString="Babic K. 1916. Opilionidi hrvatskog zemaljskog zooloskog muzeja u Zagrebu [Die Opilioniden des kroatischen zoologischen Landesmuseums zu Zagreb]. Glasnik hrvatskoga prirodoslovnoga drustva 28: 169 - 179. [in Croatian] Available from http: // biodiversitylibrary. org / page / 11087817 [accessed 23 Jun. 2016]" type="journal article" year="1916">Babić 1916</bibRefCitation>
;
<bibRefCitation id="56764B34FFE8BB1373A7FE3FCC6FFE2B" author="Novak T." box="[940,1097,384,411]" pageId="7" pageNumber="8" pagination="231 - 296" refId="ref13748" refString="Novak T. 2004 b. An overview of harvestmen (Arachnida: Opiliones) in Croatia [Pregled faune kosaca (Aachnida: Opiliones) u Hrvatskoj]. Natura Croatica 13 (3): 231 - 296." type="journal article" year="2004">Novak 2004b</bibRefCitation>
) and Bosnia-Hercegovina (
<bibRefCitation id="56764B34FFE8BB1370CEFE1BC94CFE0E" author="Martens J." box="[197,362,420,446]" pageId="7" pageNumber="8" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978</bibRefCitation>
;
<bibRefCitation id="56764B34FFE8BB13717EFE1CCA2EFE0E" author="Novak T." box="[373,520,419,446]" pageId="7" pageNumber="8" pagination="301 - 350" refId="ref13789" refString="Novak T. 2005. An overview of harvestmen (Arachnida: Opiliones) in Bosnia and Herzegovina. Natura Croatica 14 (4): 301 - 350." type="journal article" year="2005">Novak 2005</bibRefCitation>
). To the Northeast, the area extends into the
<collectingCountry id="4AF07655FFE8BB13740BFE1CCC98FE0D" box="[1024,1214,419,445]" name="Czech Republic" pageId="7" pageNumber="8">Czech Republic</collectingCountry>
: many localities in
<collectingCountry id="4AF07655FFE8BB1370D1FE78C962FE51" box="[218,324,455,481]" name="Czech Republic" pageId="7" pageNumber="8">Bohemia</collectingCountry>
and
<collectingRegion id="F023F827FFE8BB137171FE78C9C6FE51" box="[378,480,455,481]" country="Czech Republic" name="Jihomoravsky" pageId="7" pageNumber="8">Moravia</collectingRegion>
, mainly in the northern and southern mountains marking the borders to
<collectingCountry id="4AF07655FFE8BB137512FE78CD54FE51" box="[1305,1394,455,481]" name="Austria" pageId="7" pageNumber="8">Austria</collectingCountry>
,
<collectingCountry id="4AF07655FFE8BB1370B6FE55C90CFDB4" box="[189,298,490,516]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
and
<collectingCountry id="4AF07655FFE8BB137168FE55C992FDB4" box="[355,436,490,516]" name="Poland" pageId="7" pageNumber="8">Poland</collectingCountry>
(
<bibRefCitation id="56764B34FFE8BB1371CFFE57CA40FDB4" author="Silhavy V." box="[452,614,488,516]" pageId="7" pageNumber="8" pagination="204 - 208" refId="ref14175" refString="Silhavy V. 1981. Occurrence of Leiobunum glabrum in Czechoslovakia (Arach., Opilionidea). Vestnik ceskoslovenske Spolecnosti zoologicke [Acta societatis zoologicae bohemoslovacae] 45 (3): 204 - 208." type="journal article" year="1981">Šilhavý 1981</bibRefCitation>
).
</paragraph>
<paragraph id="325836C5FFE8BB1370B6FD94CBDBFD46" blockId="7.[189,1399,555,758]" pageId="7" pageNumber="8">
North of the Alps this main Alpine/Balkan area extends in isolated patches towards the northern border of low mountain ranges in
<collectingCountry id="4AF07655FFE8BB1371F1FDF0CA41FDD9" box="[506,615,591,617]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
(Black Forest, Schwäbische Alb, Frankenalb, Bavarian Forest and Bohemian Forest; all
<bibRefCitation id="56764B34FFE8BB1371B4FDCDCA40FD3C" author="Martens J." box="[447,614,626,652]" pageId="7" pageNumber="8" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978</bibRefCitation>
), Fichtelgebirge (
<bibRefCitation id="56764B34FFE8BB13733DFDCDCBEDFD3D" author="Staudt A." box="[822,971,626,653]" pageId="7" pageNumber="8" refId="ref14405" refString="Staudt A. 2016. Nachweiskarten der Spinnentiere Deutschlands (Arachnida: Araneae, Opiliones, Pseudoscorpiones): Weberknechte, Sclerosomatidae, Leiobunum rupestre. Available online from: http: // www. spiderling. de / arages / Verbreitungskarten / species _ eng. php? name = Leiobunum rupestre [accessed 3 Jan. 2016]" type="url" year="2016">Staudt 2016</bibRefCitation>
), Thüringer Wald (CJM 5392), Erzgebirge (
<bibRefCitation id="56764B34FFE8BB13712EFD2AC9EEFD1F" author="Buttner K." box="[293,456,661,687]" pageId="7" pageNumber="8" pagination="22 - 25" refId="ref11250" refString="Buttner K. 1930. Beitrag zur Kenntnis der der Kankerfauna Sudwestsachsens. Jahrbuch des Vereins fur Naturkunde Zwickau 1928 - 1930: 22 - 25." type="journal article" year="1930">Büttner 1930</bibRefCitation>
), Lausitzer Gebirge (
<bibRefCitation id="56764B34FFE8BB1372C8FD2ACB4CFD1F" author="Hiebsch H." box="[707,874,661,687]" pageId="7" pageNumber="8" pagination="1 - 32" refId="ref11974" refString="Hiebsch H. 1972. Beitrage zur Spinnen- und Weberknechtfauna des Neissetales bei Ostritz. Abhandlungen und Berichte des Naturkundemuseums Gorlitz 47: 1 - 32." type="journal article" year="1972">Hiebsch 1972</bibRefCitation>
) and northerly up to the Saxonian Elbsandsteingebirge (CJM 4752, 4753). “
<collectingRegion id="F023F827FFE8BB137246FD07CA93FD62" box="[589,693,696,722]" country="Germany" name="Sachsen" pageId="7" pageNumber="8">Sachsen</collectingRegion>
”, i.e.,
<collectingRegion id="F023F827FFE8BB13730AFD07CB7CFD62" box="[769,858,696,722]" country="Germany" name="Sachsen" pageId="7" pageNumber="8">Saxony</collectingRegion>
, is the
<typeStatus id="ED5C8867FFE8BB1373B9FD06CBC3FD63" box="[946,997,697,723]" pageId="7" pageNumber="8">type</typeStatus>
locality of
<taxonomicName id="F5E74D46FFE8BB137479FD06CCF5FD62" authorityName="Herbst" authorityYear="1799" box="[1138,1235,697,722]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB137479FD06CCF5FD62" box="[1138,1235,697,722]" italics="true" pageId="7" pageNumber="8">rupestre</emphasis>
</taxonomicName>
, and the
<typeStatus id="ED5C8867FFE8BB13754FFD06CD51FD63" box="[1348,1399,697,723]" pageId="7" pageNumber="8">type</typeStatus>
material of
<bibRefCitation id="56764B34FFE8BB13714FFD64C9C1FD45" author="Herbst J. F. W." box="[324,487,731,757]" pageId="7" pageNumber="8" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst (1799)</bibRefCitation>
likely originated from present South
<collectingRegion id="F023F827FFE8BB1373ABFD64CBD0FD45" box="[928,1014,731,757]" country="Germany" name="Sachsen" pageId="7" pageNumber="8">Saxony</collectingRegion>
.
</paragraph>
<paragraph id="325836C5FFE8BB1370B6FCA2CBEEFBBA" blockId="7.[189,1399,797,1035]" pageId="7" pageNumber="8">
For
<collectingCountry id="4AF07655FFE8BB1370FBFCA2C962FC87" box="[240,324,797,823]" name="Poland" pageId="7" pageNumber="8">Poland</collectingCountry>
,
<bibRefCitation id="56764B34FFE8BB137158FCA2CA25FC88" author="Starega W." box="[339,515,797,824]" pageId="7" pageNumber="8" pagination="1 - 197" refId="ref14305" refString="Starega W. 1976. Opiliones Kosarze (Arachnoidea). Fauna Polski 5: 1 - 197, Polska Akademia Nauk, Warsaw." type="journal article" year="1976">Staręga (1976)</bibRefCitation>
indicated a number of “
<taxonomicName id="F5E74D46FFE8BB137324FCA1CBB6FC87" authorityName="Herbst" authorityYear="1799" box="[815,912,798,823]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB137324FCA1CBB6FC87" box="[815,912,798,823]" italics="true" pageId="7" pageNumber="8">rupestre</emphasis>
</taxonomicName>
” records in the southern mountains, the Karkonosze (in German “Riesengebirge”) close to the Czech border. From the opposite Czech side, in the same mountain stock, the Krkonoše,
<bibRefCitation id="56764B34FFE8BB13729CFCDECB63FCCD" author="Silhavy V." box="[663,837,865,893]" pageId="7" pageNumber="8" pagination="204 - 208" refId="ref14175" refString="Silhavy V. 1981. Occurrence of Leiobunum glabrum in Czechoslovakia (Arach., Opilionidea). Vestnik ceskoslovenske Spolecnosti zoologicke [Acta societatis zoologicae bohemoslovacae] 45 (3): 204 - 208." type="journal article" year="1981">Šilhavý (1981)</bibRefCitation>
pinpointed records for (true)
<taxonomicName id="F5E74D46FFE8BB137495FCDBCCD9FCCD" authorityName="Herbst" authorityYear="1799" box="[1182,1279,868,893]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB137495FCDBCCD9FCCD" box="[1182,1279,868,893]" italics="true" pageId="7" pageNumber="8">rupestre</emphasis>
</taxonomicName>
, which he compared to “
<taxonomicName id="F5E74D46FFE8BB137168FC39C997FC10" box="[355,433,902,928]" class="Arachnida" family="Gagrellidae" genus="Nelima" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0093EAD7FFE8BB137168FC39C997FC10" box="[355,433,902,928]" italics="true" pageId="7" pageNumber="8">glabra</emphasis>
</taxonomicName>
” from his country. As these close-to-border Polish localities are situated in mountainous areas, too, they probably belong to
<taxonomicName id="F5E74D46FFE8BB1372CAFC15CB04FC73" authorityName="Herbst" authorityYear="1799" box="[705,802,938,963]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB1372CAFC15CB04FC73" box="[705,802,938,963]" italics="true" pageId="7" pageNumber="8">rupestre</emphasis>
</taxonomicName>
as well. Besides, the Polish
<taxonomicName id="F5E74D46FFE8BB13748AFC15CCE8FC73" baseAuthorityName="Martens" baseAuthorityYear="1978" box="[1153,1230,938,963]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tisciae">
<emphasis id="0093EAD7FFE8BB13748AFC15CCE8FC73" box="[1153,1230,938,963]" italics="true" pageId="7" pageNumber="8">tisciae</emphasis>
</taxonomicName>
(i.e.,
<taxonomicName id="F5E74D46FFE8BB137517FC15CD49FC74" authorityName="Thorell" authorityYear="1876" box="[1308,1391,938,964]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE8BB137517FC15CD49FC74" box="[1308,1391,938,964]" italics="true" pageId="7" pageNumber="8">gracile</emphasis>
</taxonomicName>
, below) records are all situated in lower stretches of central and eastern parts of the country (
<bibRefCitation id="56764B34FFE8BB137516FC72C8DCFBBA" author="Starega W." pageId="7" pageNumber="8" pagination="1 - 197" refId="ref14305" refString="Starega W. 1976. Opiliones Kosarze (Arachnoidea). Fauna Polski 5: 1 - 197, Polska Akademia Nauk, Warsaw." type="journal article" year="1976">Staręga 1976</bibRefCitation>
,
<bibRefCitation id="56764B34FFE8BB13710DFC4FC963FBBA" author="Starega W." box="[262,325,1008,1034]" pageId="7" pageNumber="8" pagination="78 - 88" refId="ref14380" refString="Starega W. 2004. Interessante Weberknechtfunde aus Polen (Arachnida: Opiliones). Arachnologische Mitteilungen 27 / 28: 78 - 88." type="journal article" year="2004">2004</bibRefCitation>
), far from any (possible) mountainous
<taxonomicName id="F5E74D46FFE8BB137306FC4ECB48FBBA" authorityName="Herbst" authorityYear="1799" box="[781,878,1009,1034]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB137306FC4ECB48FBBA" box="[781,878,1009,1034]" italics="true" pageId="7" pageNumber="8">rupestre</emphasis>
</taxonomicName>
record.
</paragraph>
<caption id="6698664DFFE8BB1370B6F8D9CD30F85A" ID-DOI="http://doi.org/10.5281/zenodo.3840155" ID-Zenodo-Dep="3840155" httpUri="https://zenodo.org/record/3840155/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1894,1920]" targetBox="[408,1184,1137,1848]" targetPageId="7">
<paragraph id="325836C5FFE8BB1370B6F8D9CD30F85A" blockId="7.[189,1399,1894,2027]" pageId="7" pageNumber="8">
<emphasis id="0093EAD7FFE8BB1370B6F8D9C928F830" bold="true" box="[189,270,1894,1920]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
Body of
<taxonomicName id="F5E74D46FFE8BB13718CF8D9CAC2F831" authority="C. L. Koch, 1839" authorityName="C. L. Koch" authorityYear="1839" box="[391,740,1894,1921]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="0093EAD7FFE8BB13718CF8D9CA2DF830" box="[391,523,1894,1920]" italics="true" pageId="7" pageNumber="8">Leiobunum</emphasis>
C.L. Koch, 1839
</taxonomicName>
, dorsal view. AB.
<taxonomicName id="F5E74D46FFE8BB1373D2F8D9CD57F831" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" box="[985,1393,1894,1921]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE8BB1373D2F8D9CCEFF830" box="[985,1225,1894,1920]" italics="true" pageId="7" pageNumber="8">Leiobunum rupestre</emphasis>
<bibRefCitation id="56764B34FFE8BB1374DFF8D9CD57F831" author="Herbst J. F. W." box="[1236,1393,1894,1921]" pageId="7" pageNumber="8" refId="ref12001" refString="Herbst J. F. W. 1799. Natursystem der ungeflugelten Insekten 3. Fortsetzung der Naturgeschichte der Insectengattung Opilio. Vol 3. Lange, Berlin." type="book" year="1799">Herbst, 1799</bibRefCitation>
</taxonomicName>
, Germany, Mt. Arber, CJM136.
<emphasis id="0093EAD7FFE8BB137227F836CA67F813" bold="true" box="[556,577,1929,1955]" pageId="7" pageNumber="8">A</emphasis>
. ♁.
<emphasis id="0093EAD7FFE8BB137278F835CAA1F814" bold="true" box="[627,647,1930,1956]" pageId="7" pageNumber="8">B</emphasis>
. ♀. —
<emphasis id="0093EAD7FFE8BB1372D6F836CB30F813" bold="true" box="[733,790,1929,1955]" pageId="7" pageNumber="8">CD</emphasis>
.
<taxonomicName id="F5E74D46FFE8BB13732EF835CC88F814" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[805,1198,1930,1956]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="0093EAD7FFE8BB13732EF835CC22F814" box="[805,1028,1930,1956]" italics="true" pageId="7" pageNumber="8">Leiobunum gracile</emphasis>
<bibRefCitation id="56764B34FFE8BB137400F835CC88F814" author="Thorell T." box="[1035,1198,1930,1956]" pageId="7" pageNumber="8" pagination="452 - 508" refId="ref14658" refString="Thorell T. 1876. Sopra alcuni Opilioni (Phalangidea) d'Europa e dell'Asia occidentale, con un quadro dei generi europei di quest'Ordine. Annali del Museo civico di storia naturale di Genova 8: 452 - 508. Available form http: // biodiversitylibrary. org / page / 7916212 [accessed 23 Jun. 2016]" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
, Denmark.
<emphasis id="0093EAD7FFE8BB13753DF836CD6DF813" bold="true" box="[1334,1355,1929,1955]" pageId="7" pageNumber="8">C</emphasis>
. ♁, 2 km N of Skaerbaek, CJM3530.
<emphasis id="0093EAD7FFE8BB13725BF812CA43F877" bold="true" box="[592,613,1965,1991]" pageId="7" pageNumber="8">D</emphasis>
. ♀, Asp, CJM3531. —
<emphasis id="0093EAD7FFE8BB137385F812CBE4F877" bold="true" box="[910,962,1965,1991]" pageId="7" pageNumber="8">EF</emphasis>
.
<taxonomicName id="F5E74D46FFE8BB1373DFF812C922F85B" authority="(Martens, 1969)" baseAuthorityName="Martens" baseAuthorityYear="1969" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="apenninicum">
<emphasis id="0093EAD7FFE8BB1373DFF812CCDDF877" box="[980,1275,1965,1991]" italics="true" pageId="7" pageNumber="8">Leiobunum apenninicum</emphasis>
(
<bibRefCitation id="56764B34FFE8BB137506F812C8DDF85B" author="Martens J." pageId="7" pageNumber="8" pagination="395 - 415" refId="ref13383" refString="Martens J. 1969. Mittel- und sudeuropaische Arten der Gattung Nelima (Arachnida: Opiliones: Leiobunidae). Senckenbergiana biologica 50 (5 / 6): 395 - 415." type="journal article" year="1969">Martens, 1969</bibRefCitation>
)
</taxonomicName>
, France, Alpes-Maritimes.
<emphasis id="0093EAD7FFE8BB137249F86FCA70F85A" bold="true" box="[578,598,2000,2026]" pageId="7" pageNumber="8">E</emphasis>
. ♁, CJM1508.
<emphasis id="0093EAD7FFE8BB137307F86FCB38F85A" bold="true" box="[780,798,2000,2026]" pageId="7" pageNumber="8">F</emphasis>
. ♀, CJM2747. Drawings by K. Rehbinder.
</paragraph>
</caption>
<paragraph id="325836C5FFE7BB1C70B6FEB4CC2CFE02" blockId="8.[189,1399,267,435]" pageId="8" pageNumber="9">
The
<taxonomicName id="F5E74D46FFE7BB1C70FDFEB3C95DFE95" authorityName="Herbst" authorityYear="1799" box="[246,379,268,293]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE7BB1C70FDFEB3C95DFE95" box="[246,379,268,293]" italics="true" pageId="8" pageNumber="9">L. rupestre</emphasis>
</taxonomicName>
area as depicted in
<bibRefCitation id="56764B34FFE7BB1C7272FEB3CB0BFE95" author="Martens J." box="[633,813,267,294]" pageId="8" pageNumber="9" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens (1978</bibRefCitation>
: fig. 787) needs to be reduced, substracting the distribution area of.
<emphasis id="0093EAD7FFE7BB1C71ADFE90CA06FEF9" box="[422,544,303,329]" italics="true" pageId="8" pageNumber="9">L. tisciae</emphasis>
(i.e.,
<emphasis id="0093EAD7FFE7BB1C726AFE90CA9AFEF9" box="[609,700,303,329]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="F5E74D46FFE7BB1C726AFE90CA91FEF9" authorityName="Thorell" authorityYear="1876" box="[609,695,303,329]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gracile">gracile</taxonomicName>
,
</emphasis>
below), stretching from the Carpathians in a broad northerly area to the Baltic coast up to South
<collectingCountry id="4AF07655FFE7BB1C7250FEEDCA93FEDC" box="[603,693,338,364]" name="Finland" pageId="8" pageNumber="9">Finland</collectingCountry>
and in the West to
<collectingCountry id="4AF07655FFE7BB1C739FFEEDCC27FEDC" box="[916,1025,338,364]" name="Denmark" pageId="8" pageNumber="9">Denmark</collectingCountry>
and northern
<collectingCountry id="4AF07655FFE7BB1C74A8FEEDCD36FEDC" box="[1187,1296,338,364]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
(
<figureCitation id="AADC2A40FFE7BB1C752BFEEDCD4EFEDC" box="[1312,1384,338,364]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[186,1399,1156,1766]" captionTargetId="figure@6.[185,1400,1154,1767]" captionTargetPageId="6" captionText="Fig. 2. Distribution of the species of the Leiobunum rupestre species group. Red squares = L. rupestre Herbst, 1799; blue circles = L. gracile Thorell, 1876; green triangles = L. apenninicum (Martens, 1969); stippled line circumscribes area of L. subalpinum Komposch, 1998. Type localities are indicated by abbreviated taxon names (ape = apenninicum; gla = glabrum; gra = gracile; lae = laeve; nor = norvegicum; rup = rupestre; sub subalpinum; tis = tisciae). Arrows indicate a specified locality, if unprecisely given, the general area is indicated (rupestre = Saxony; gracile, laeve = Scania)." figureDoi="http://doi.org/10.5281/zenodo.3840153" httpUri="https://zenodo.org/record/3840153/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). However, the remaining distribution area of
<taxonomicName id="F5E74D46FFE7BB1C72B4FEC9CB18FE3F" authorityName="Herbst" authorityYear="1799" box="[703,830,374,399]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE7BB1C72B4FEC9CB18FE3F" box="[703,830,374,399]" italics="true" pageId="8" pageNumber="9">L. rupestre</emphasis>
</taxonomicName>
slightly changed with respect to the Balkan area, then largely unknown, and new records in central
<collectingCountry id="4AF07655FFE7BB1C7303FE27CB53FE02" box="[776,885,408,434]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
were added.
</paragraph>
</subSubSection>
<subSubSection id="7AFD654EFFE7BB1C70B6FE65C927FC2C" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="325836C5FFE7BB1C70B6FE65C906FE44" blockId="8.[189,288,474,500]" box="[189,288,474,500]" pageId="8" pageNumber="9">
<heading id="691081A9FFE7BB1C70B6FE65C906FE44" bold="true" box="[189,288,474,500]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="0093EAD7FFE7BB1C70B6FE65C906FE44" bold="true" box="[189,288,474,500]" pageId="8" pageNumber="9">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="325836C5FFE7BB1C70B6FDB7C9FEFD45" blockId="8.[189,1399,520,758]" pageId="8" pageNumber="9">
This is a forest species with a high demand of air moisture. Therefore, it is confined to moist places close to streams and creeks, on tree trunks, under rotten wood and on moist rock faces. It hardly occurs in open areas, in agricultural land, urban gardens or on house walls in villages and towns, if not situated in moist forests (
<bibRefCitation id="56764B34FFE7BB1C7117FDCDC9E3FD3C" author="Martens J." box="[284,453,626,652]" pageId="8" pageNumber="9" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens 1978</bibRefCitation>
), or at higher altitudes.
<bibRefCitation id="56764B34FFE7BB1C72E8FDCDCBBCFD3C" author="Martens J." box="[739,922,626,652]" pageId="8" pageNumber="9" refId="ref13416" refString="Martens J. 1978. Spinnentiere, Arachnida - Weberknechte, Opiliones. Die Tierwelt Deutschlands 64, VEB Gustav Fischer, Jena." type="journal volume" year="1978">Martens (1978)</bibRefCitation>
indicated height records from
<quantity id="F51F9B20FFE7BB1C7500FDCDCD71FD3C" box="[1291,1367,626,652]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="9" unit="m" value="260.0">260 m</quantity>
to
<quantity id="F51F9B20FFE7BB1C70B6FD2AC93EFD1F" box="[189,280,661,687]" metricMagnitude="3" metricUnit="m" metricValue="2.16" pageId="8" pageNumber="9" unit="m" value="2160.0">2160 m</quantity>
a.s.l. in the Austrian Alps, but likely included localities for the closely related
<taxonomicName id="F5E74D46FFE7BB1C74CDFD29CD49FD1F" authorityName="Komposch" authorityYear="1998" box="[1222,1391,661,687]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subalpinum">
<emphasis id="0093EAD7FFE7BB1C74CDFD29CD49FD1F" box="[1222,1391,661,687]" italics="true" pageId="8" pageNumber="9">L. subalpinum</emphasis>
</taxonomicName>
.
<bibRefCitation id="56764B34FFE7BB1C70B6FD07CA32FD62" author="Komposch C. &amp; Gruber J." box="[189,532,696,722]" pageId="8" pageNumber="9" pagination="485 - 534" refId="ref12950" refString="Komposch C. &amp; Gruber J. 2004. Die Weberknechte Osterreichs (Arachnida, Opiliones). Denisia NS 12: 485 - 534." type="journal article" year="2004">Komposch &amp; Gruber (2004)</bibRefCitation>
set the Austrian uppermost
<taxonomicName id="F5E74D46FFE7BB1C7366FD06CBE8FD62" authorityName="Herbst" authorityYear="1799" box="[877,974,697,722]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE7BB1C7366FD06CBE8FD62" box="[877,974,697,722]" italics="true" pageId="8" pageNumber="9">rupestre</emphasis>
</taxonomicName>
limit at
<quantity id="F51F9B20FFE7BB1C7431FD07CCB0FD63" box="[1082,1174,696,723]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="9" unit="m" value="1680.0">1680 m</quantity>
a.s.l. and for
<taxonomicName id="F5E74D46FFE7BB1C754FFD07C93BFD45" authorityName="Komposch" authorityYear="1998" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subalpinum">
<emphasis id="0093EAD7FFE7BB1C754FFD07C93BFD45" italics="true" pageId="8" pageNumber="9">subalpinum</emphasis>
</taxonomicName>
at
<quantity id="F51F9B20FFE7BB1C714AFD64C9BCFD46" box="[321,410,731,758]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="9" unit="m" value="2100.0">2100 m</quantity>
a.s.l.
</paragraph>
<paragraph id="325836C5FFE7BB1C70B6FCA5C927FC2C" blockId="8.[189,1399,793,924]" pageId="8" pageNumber="9">
In all cases populations of
<taxonomicName id="F5E74D46FFE7BB1C71FDFCA5CA71FC83" authorityName="Herbst" authorityYear="1799" box="[502,599,794,819]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE7BB1C71FDFCA5CA71FC83" box="[502,599,794,819]" italics="true" pageId="8" pageNumber="9">rupestre</emphasis>
</taxonomicName>
are confined to mountainous areas of different altitudes. Only in the Saxonian Elbsandsteingebirge does
<taxonomicName id="F5E74D46FFE7BB1C7266FC82CAE8FCE6" authorityName="Herbst" authorityYear="1799" box="[621,718,829,854]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rupestre">
<emphasis id="0093EAD7FFE7BB1C7266FC82CAE8FCE6" box="[621,718,829,854]" italics="true" pageId="8" pageNumber="9">rupestre</emphasis>
</taxonomicName>
live at the bottom of deeply recessed valleys which are permanently moist and where sun radiation is largely excluded from the narrow valleys by the steep rock faces.
</paragraph>
</subSubSection>
</treatment>
</document>