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<document id="59553C9C2A72C9F5273F71F5BB6E7CA2" ID-CLB-Dataset="55477" ID-DOI="10.11646/zootaxa.5138.3.4" ID-GBIF-Dataset="456f68b2-d5a3-406f-b4d4-f3610cfc11f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="6559737" ID-ZooBank="E516B443-1E3B-4A6E-B8F3-E24FD80F2610" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652866623385" checkinUser="plazi" docAuthor="Cartwright, David I. &amp; Dostine, Peter" docDate="2022" docId="03F7903AFFF0682F45CA30AABE30FE04" docLanguage="en" docName="zootaxa.5138.3.4.pdf" docOrigin="Zootaxa 5138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neboissomina topendica Cartwright &amp; Dostine 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="290" masterDocId="FFCEE842FFF56828455D350BBA7BFF8D" masterDocTitle="Five new species and new records of caddisflies (Insecta: Trichoptera) from Australia’s ‘ Top End’" masterLastPageNumber="304" masterPageNumber="283" pageNumber="288" updateTime="1699337787172" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="235A9FF8FB9CA76E4504B1C9F8F6FBC1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FA51222908B85BBE347111979BE4BC65">
<mods:title id="11753360C680597627DD5102E2D8D03C">Five new species and new records of caddisflies (Insecta: Trichoptera) from Australia’s ‘ Top End’</mods:title>
</mods:titleInfo>
<mods:name id="1751778AB88C891935C86BDC2E6E2501" type="personal">
<mods:role id="37BF6D983C52F3E912D9078F77865F19">
<mods:roleTerm id="0221A16C6AA1222514EB7DD06B438DFC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82B1A3CDCEFD7E7BBA150FABFFED3338">Cartwright, David I.</mods:namePart>
<mods:affiliation id="C52C2E62C9F0A35CCBF3AEB256E5E16D">13 Brolga Crescent, Wandana Heights, Vic 3216, Australia.</mods:affiliation>
</mods:name>
<mods:name id="BD55C32943A0A21221F6073A541B57AA" type="personal">
<mods:role id="E012E7F0EC39A6C7F7741BE92FC9451E">
<mods:roleTerm id="8E32D7B3A804D7F1C4C5D00C6B46CFFF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D00524FAED36F2B78CA8336FFE469211">Dostine, Peter</mods:namePart>
<mods:affiliation id="C7605188C9CEF15299B03FAF45DDFC44">Department of Environment, Parks and Water Security, PO Box 496, Palmerston, NT 0831, Australia.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="ABC7D1040013FB7EB987FC493B05FA39">text</mods:typeOfResource>
<mods:relatedItem id="1AB64565287EFC751E32ABD4BA9A3DB9" type="host">
<mods:titleInfo id="04570547102F7D172FBABA2334F5D168">
<mods:title id="832525849B2DCC25C2D6A051EF9F7138">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7B36509A4A1711C45B78D8EB1FC9AE73">
<mods:date id="F437AFD3230BA6AC5C6A4BAC88891453">2022</mods:date>
<mods:detail id="BF570B2E25D0EAA247474D93F3F15989" type="pubDate">
<mods:number id="20A90E78A3ABE1427A8D6F40D2972A23">2022-05-18</mods:number>
</mods:detail>
<mods:detail id="E1B1531FCB580F272D98D8DA891E3CBA" type="volume">
<mods:number id="693367A97A0B6CEAC4A6DFF577CB34CD">5138</mods:number>
</mods:detail>
<mods:detail id="32728ED9EA7DC8A3F957D93094474BB9" type="issue">
<mods:number id="447059E32B6599C7BE3AE48C820FD8B8">3</mods:number>
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<mods:start id="B0E92165A4BB95E293ADFE7FB764FE7F">283</mods:start>
<mods:end id="8FD1C4143961897232834412E78404F1">304</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="82446409C4386C64D05FEA7398128889">journal article</mods:classification>
<mods:identifier id="4CCDDF2207923F31514FF7F73F4ED53E" type="CLB-Dataset">55477</mods:identifier>
<mods:identifier id="9A9DE070253F8A481F34E54487D8F166" type="DOI">10.11646/zootaxa.5138.3.4</mods:identifier>
<mods:identifier id="9FD92EA872BAC15FD19090F07127FF69" type="GBIF-Dataset">456f68b2-d5a3-406f-b4d4-f3610cfc11f9</mods:identifier>
<mods:identifier id="60F67D8268530D7F767C829CEB2667D1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9FDC60888EDE8EF4879B2BD537C4858E" type="Zenodo-Dep">6559737</mods:identifier>
<mods:identifier id="05DB773D6FB563835041A22541593D6F" type="ZooBank">E516B443-1E3B-4A6E-B8F3-E24FD80F2610</mods:identifier>
</mods:mods>
<treatment id="03F7903AFFF0682F45CA30AABE30FE04" ID-DOI="http://doi.org/10.5281/zenodo.6559633" ID-GBIF-Taxon="195535824" ID-Zenodo-Dep="6559633" LSID="urn:lsid:plazi:treatment:03F7903AFFF0682F45CA30AABE30FE04" httpUri="http://treatment.plazi.org/id/03F7903AFFF0682F45CA30AABE30FE04" lastPageId="7" lastPageNumber="290" pageId="5" pageNumber="288">
<subSubSection id="C34472A7FFF0682D45CA30AAB85FFA36" box="[151,548,1441,1468]" pageId="5" pageNumber="288" type="nomenclature">
<paragraph id="8BE1212CFFF0682D45CA30AAB85FFA36" blockId="5.[151,548,1441,1503]" box="[151,548,1441,1468]" pageId="5" pageNumber="288">
<heading id="D0A99640FFF0682D45CA30AAB85FFA36" bold="true" box="[151,548,1441,1468]" fontSize="11" level="1" pageId="5" pageNumber="288" reason="1">
<emphasis id="B92AFD3EFFF0682D45CA30AAB85FFA36" bold="true" box="[151,548,1441,1468]" pageId="5" pageNumber="288">
<taxonomicName id="4C5E5AAFFFF0682D45CA30AABBC4FA31" authority="Cartwright &amp; Dostine, 2022" authorityName="Cartwright &amp; Dostine" authorityYear="2022" box="[151,447,1441,1468]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="topendica" status="sp. nov.">
<emphasis id="B92AFD3EFFF0682D45CA30AABBC4FA31" bold="true" box="[151,447,1441,1468]" italics="true" pageId="5" pageNumber="288">Neboissomina topendica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194045FFF0682D449B30AAB85FFA36" box="[454,548,1441,1467]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF0682D45CA30CEBB67FA52" box="[151,284,1477,1503]" pageId="5" pageNumber="288" type="description">
<paragraph id="8BE1212CFFF0682D45CA30CEBB67FA52" blockId="5.[151,548,1441,1503]" box="[151,284,1477,1503]" pageId="5" pageNumber="288">
<figureCitation id="13653DA9FFF0682D45CA30CEBB67FA52" box="[151,284,1477,1503]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="5" pageNumber="288">Figures 6–9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF0682D45CA3306B81BF8A9" pageId="5" pageNumber="288" type="materials_examined">
<paragraph id="8BE1212CFFF0682D45CA3306BB7DF9C1" blockId="5.[151,1437,1549,2008]" pageId="5" pageNumber="288">
<emphasis id="B92AFD3EFFF0682D45CA3306BB3EF9AA" bold="true" box="[151,325,1549,1575]" pageId="5" pageNumber="288">Type material.</emphasis>
<materialsCitation id="3B362B71FFF0682D44163306BB7DF9C1" ID-GBIF-Occurrence="3775806325" collectingDate="2015-05-21" collectionCode="MVM" collectorName="P. Dostine" location="Umbrawarra Gorge" pageId="5" pageNumber="288" specimenCode="T-22578" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="54E59F8EFFF0682D44163306BBC8F9AA" box="[331,435,1549,1575]" pageId="5" pageNumber="288" type="holotype">Holotype</typeStatus>
male (CT-807),
<collectingRegion id="499AEFCEFFF0682D47363306B941F9AA" box="[619,826,1549,1575]" country="Australia" name="Northern Territory" pageId="5" pageNumber="288">Northern Territory</collectingRegion>
,
<location id="8E8177F7FFF0682D46183306BE5AF9AA" LSID="urn:lsid:plazi:treatment:03F7903AFFF0682F45CA30AABE30FE04:8E8177F7FFF0682D46183306BE5AF9AA" box="[837,1057,1549,1575]" name="Umbrawarra Gorge" pageId="5" pageNumber="288" stateProvince="Northern Territory">Umbrawarra Gorge</location>
,
<date id="FFE007ECFFF0682D41713306BEBFF9A5" box="[1068,1220,1549,1576]" pageId="5" pageNumber="288" value="2015-05-21">
<collectingDate id="EFA4FE04FFF0682D41713306BEBFF9A5" box="[1068,1220,1549,1576]" pageId="5" pageNumber="288" value="2015-05-21">21 May 2015</collectingDate>
</date>
,
<collectorName id="26AB44FAFFF0682D41933305BF44F9AA" box="[1230,1343,1549,1576]" pageId="5" pageNumber="288">P. Dostine</collectorName>
(
<collectionCode id="ED4FB9E9FFF0682D40133306BFE3F9AA" box="[1358,1432,1549,1575]" pageId="5" pageNumber="288">MVM</collectionCode>
,
<specimenCode id="DBF88957FFF0682D45CA333ABA80F9C1" box="[151,251,1585,1612]" collectionCode="T-" pageId="5" pageNumber="288">T-22578</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE1212CFFF0682D459A335DB87FF91E" blockId="5.[151,1437,1549,2008]" pageId="5" pageNumber="288">
<materialsCitation id="3B362B71FFF0682D459A335DBF78F9E2" ID-GBIF-Occurrence="3775806491" box="[199,1283,1621,1648]" collectingDate="2015-05-21" collectionCode="MVM" collectorName="P. Dostine" location="Umbrawarra Gorge" pageId="5" pageNumber="288" specimenCount="1" specimenCount-male="1" stateProvince="Northern Territory" typeStatus="paratype">
<typeStatus id="54E59F8EFFF0682D459A335DBB43F9FD" box="[199,312,1622,1648]" pageId="5" pageNumber="288" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="499AEFCEFFF0682D441C335EB86AF9E2" box="[321,529,1621,1647]" country="Australia" name="Northern Territory" pageId="5" pageNumber="288">Northern Territory</collectingRegion>
.
<specimenCount id="9D58EAA5FFF0682D4746335EB81DF9E2" box="[539,614,1621,1647]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
(CT-667/DC-312 figured), collected with holotype (
<collectionCode id="ED4FB9E9FFF0682D41F2335EBE80F9E2" box="[1199,1275,1621,1647]" pageId="5" pageNumber="288">MVM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B362B71FFF0682D4050335DB87BF91E" ID-GBIF-Occurrence="3775806535" collectingDate="2015-05-21" collectionCode="MAGNT" collectorName="P. Dostine" location="Umbrawarra Gorge" pageId="5" pageNumber="288" specimenCount="7" specimenCount-male="7" stateProvince="Northern Territory" typeStatus="paratype">
<specimenCount id="9D58EAA5FFF0682D4050335DBF1DF9E2" box="[1293,1382,1621,1648]" pageId="5" pageNumber="288" type="male">7 males</specimenCount>
, collected with holotype (
<collectionCode id="ED4FB9E9FFF0682D44D63372BB83F91E" box="[395,504,1657,1683]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34986" lsid="urn:lsid:biocol.org:col:34986" name="Museum and Art Gallery of the Northern Territory" pageId="5" pageNumber="288" type="Museum">MAGNT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE1212CFFF0682D459A3396B81BF8A9" blockId="5.[151,1437,1549,2008]" pageId="5" pageNumber="288">
Other material examined.
<materialsCitation id="3B362B71FFF0682D44B23396BE15F935" ID-GBIF-Occurrence="3775806479" box="[495,1134,1693,1720]" collectingDate="2015-05-14" collectionCode="R" collectorName="Blackmore" location="Northern Territory" pageId="5" pageNumber="288" specimenCount="1" specimenCount-male="1" stateProvince="Northern Territory">
<collectingRegion id="499AEFCEFFF0682D44B23396B8BBF93A" box="[495,704,1693,1719]" country="Australia" name="Northern Territory" pageId="5" pageNumber="288">Northern Territory</collectingRegion>
.
<specimenCount id="9D58EAA5FFF0682D47913396B960F93A" box="[716,795,1693,1719]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
,
<collectorName id="26AB44FAFFF0682D467A3396B9D9F93A" box="[807,930,1693,1719]" pageId="5" pageNumber="288">Blackmore</collectorName>
<collectionCode id="ED4FB9E9FFF0682D46F73395B9B9F935" box="[938,962,1694,1720]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="288">R</collectionCode>
.,
<date id="FFE007ECFFF0682D468E3396BE15F935" box="[979,1134,1693,1720]" pageId="5" pageNumber="288" value="2015-05-14">
<collectingDate id="EFA4FE04FFF0682D468E3396BE15F935" box="[979,1134,1693,1720]" pageId="5" pageNumber="288" value="2015-05-14">14 May 2015</collectingDate>
</date>
</materialsCitation>
, P. Dostine;
<specimenCount id="9D58EAA5FFF0682D41A13396BF2DF93A" box="[1276,1366,1693,1719]" pageId="5" pageNumber="288" type="male">2 males</specimenCount>
, same loc. and coll.,
<date id="FFE007ECFFF0682D446733CABBB3F951" box="[314,456,1729,1756]" pageId="5" pageNumber="288" value="2015-06-17">17 Jun 2015</date>
;
<specimenCount id="9D58EAA5FFF0682D448833CAB85FF956" box="[469,548,1729,1755]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
, same loc. and coll.,
<date id="FFE007ECFFF0682D464833CAB9D7F951" box="[789,940,1729,1756]" pageId="5" pageNumber="288" value="2015-08-13">13 Aug 2015</date>
;
<specimenCount id="9D58EAA5FFF0682D46E433CABE73F956" box="[953,1032,1729,1755]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
, same loc. and coll.,
<date id="FFE007ECFFF0682D41A433CABFF9F956" box="[1273,1410,1729,1756]" pageId="5" pageNumber="288" value="2016-08-03">3 Aug 2016</date>
;
<specimenCount id="9D58EAA5FFF0682D40D333CABAABF972" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
, same loc. and coll.,
<date id="FFE007ECFFF0682D44EA33EEB831F972" box="[439,586,1765,1792]" pageId="5" pageNumber="288" value="2016-08-31">31 Aug 2016</date>
;
<specimenCount id="9D58EAA5FFF0682D470933EEB8DAF972" box="[596,673,1765,1791]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
, same loc. and coll.,
<date id="FFE007ECFFF0682D46DA33EEBE67F972" box="[903,1052,1765,1792]" pageId="5" pageNumber="288" value="2016-11-28">28 Nov 2016</date>
;
<specimenCount id="9D58EAA5FFF0682D417B33EEBE08F972" box="[1062,1139,1765,1791]" pageId="5" pageNumber="288" type="male">1 male</specimenCount>
, Sandy Billabong, Kakadu N.P.,
<date id="FFE007ECFFF0682D45893202BB17F8AE" box="[212,364,1801,1828]" pageId="5" pageNumber="288" value="2017-05-26">26 May 2017</date>
,
<materialsCitation id="3B362B71FFF0682D442A3201B827F8A9" ID-GBIF-Occurrence="3775806441" box="[375,604,1801,1828]" collectionCode="FFD" collectorName="Dostine" pageId="5" pageNumber="288" specimenCount="1" stateProvince="Northern Territory">
P.
<collectorName id="26AB44FAFFF0682D44CF3202BB92F8AE" box="[402,489,1801,1827]" pageId="5" pageNumber="288">Dostine</collectorName>
(all
<collectionCode id="ED4FB9E9FFF0682D47413201B82FF8A9" box="[540,596,1802,1828]" pageId="5" pageNumber="288">FFD</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF0682D459A3226B936F83E" pageId="5" pageNumber="288" type="diagnosis">
<paragraph id="8BE1212CFFF0682D459A3226B936F83E" blockId="5.[151,1437,1549,2008]" pageId="5" pageNumber="288">
<emphasis id="B92AFD3EFFF0682D459A3226BB39F8CA" bold="true" box="[199,322,1837,1863]" pageId="5" pageNumber="288">Diagnosis.</emphasis>
The male of
<taxonomicName id="4C5E5AAFFFF0682D44803226B888F8CA" authority="Cartwright &amp; Dostine, 2022" authorityName="Cartwright &amp; Dostine" authorityYear="2022" box="[477,755,1837,1863]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="topendica" status="sp. nov.">
<emphasis id="B92AFD3EFFF0682D44803226B888F8CA" box="[477,755,1837,1863]" italics="true" pageId="5" pageNumber="288">Neboissomina topendica</emphasis>
</taxonomicName>
<emphasis id="B92AFD3EFFF0682D47A13226B92CF8CA" bold="true" box="[764,855,1837,1863]" pageId="5" pageNumber="288">
<taxonomicNameLabel id="A2194045FFF0682D47A13226B92CF8CA" box="[764,855,1837,1863]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to those of
<taxonomicName id="4C5E5AAFFFF0682D41C93225BF74F8CA" authorityName="Cartwright" authorityYear="2011" box="[1172,1295,1837,1863]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="jardinei">
<emphasis id="B92AFD3EFFF0682D41C93225BF74F8CA" box="[1172,1295,1837,1863]" italics="true" pageId="5" pageNumber="288">N. jardinei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5E5AAFFFF0682D40433225BB2AF8E6" authority="Cartwright 2011" authorityName="Cartwright" authorityYear="2011" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="kuranya">
<emphasis id="B92AFD3EFFF0682D40433225BFE7F8CA" box="[1310,1436,1837,1863]" italics="true" pageId="5" pageNumber="288">N. kuranya</emphasis>
<bibRefCitation id="EFCF5CDDFFF0682D45CA325ABB2AF8E6" author="Cartwright, D. I." box="[151,337,1873,1899]" pageId="5" pageNumber="288" pagination="17 - 30" refId="ref14740" refString="Cartwright, D. I. (2011) Descriptions of Neboissomina, new genus and 6 new species of Ecnomidae from Australia (Trichoptera). Zootaxa, 2736 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 2736.1.2" type="journal article" year="2011">Cartwright 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5E5AAFFFF0682D44013259BBBBF8E6" authorityName="Cartwright" authorityYear="2011" box="[348,448,1873,1899]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="riyala">
<emphasis id="B92AFD3EFFF0682D44013259BBBBF8E6" box="[348,448,1873,1899]" italics="true" pageId="5" pageNumber="288">N. riyala</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5E5AAFFFF0682D44A03259B93CF8E6" authority="Cartwright 2011" authorityName="Cartwright" authorityYear="2011" box="[509,839,1873,1899]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="philsuteri">
<emphasis id="B92AFD3EFFF0682D44A03259B8F2F8E6" box="[509,649,1873,1899]" italics="true" pageId="5" pageNumber="288">N. philsuteri</emphasis>
<bibRefCitation id="EFCF5CDDFFF0682D47CD325AB93CF8E6" author="Cartwright, D. I." box="[656,839,1873,1899]" pageId="5" pageNumber="288" pagination="17 - 30" refId="ref14740" refString="Cartwright, D. I. (2011) Descriptions of Neboissomina, new genus and 6 new species of Ecnomidae from Australia (Trichoptera). Zootaxa, 2736 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 2736.1.2" type="journal article" year="2011">
Cartwright
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</bibRefCitation>
</taxonomicName>
in possessing inferior appendages which are fused to form a single large sub-rectangular plate, but it is distinguished by the three, minute, acute projections apicolaterally and apicomesally on each inferior appendage, in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF0682F459A32B6BBE4FE90" lastPageId="7" lastPageNumber="290" pageId="5" pageNumber="288" type="description">
<paragraph id="8BE1212CFFF0682E459A32B6BF00FECC" blockId="5.[151,1437,1549,2008]" lastBlockId="6.[151,1436,151,321]" lastPageId="6" lastPageNumber="289" pageId="5" pageNumber="288">
<emphasis id="B92AFD3EFFF0682D459A32B6BBE0F85A" bold="true" box="[199,411,1981,2007]" pageId="5" pageNumber="288">Description, male</emphasis>
. Spur formula: 3:4:4. Head, body, and wings light brown, abdomen paler ventrally; wings (
<figureCitation id="13653DA9FFF3682E45FD359CBA98FF3F" box="[160,227,151,178]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Fig. 6</figureCitation>
) similar to those of
<taxonomicName id="4C5E5AAFFFF3682E4499359CB846FF3C" authorityName="Cartwright" authorityYear="2011" box="[452,573,151,177]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="jardinei">
<emphasis id="B92AFD3EFFF3682E4499359CB846FF3C" box="[452,573,151,177]" italics="true" pageId="6" pageNumber="289">N. jardinei</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCF5CDDFFF3682E4716359CB97CFF3C" author="Cartwright, D. I." box="[587,775,151,177]" pageId="6" pageNumber="289" pagination="17 - 30" refId="ref14740" refString="Cartwright, D. I. (2011) Descriptions of Neboissomina, new genus and 6 new species of Ecnomidae from Australia (Trichoptera). Zootaxa, 2736 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 2736.1.2" type="journal article" year="2011">Cartwright 2011</bibRefCitation>
, fig. 2), length of each forewing: male
<quantity id="4CA68CC9FFF3682E41E6359CBF3FFF3C" box="[1211,1348,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="6" pageNumber="289" unit="mm" value="2.7" valueMax="3.1" valueMin="2.3">2.3–3.1 mm</quantity>
, female
<quantity id="4CA68CC9FFF3682E45CA35B0BB58FF58" box="[151,291,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="6" pageNumber="289" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
, length about 3.0 times width; length of each hind wing 3.2–3.3 times width. Wing venation: Each forewing with forks 2, 3, 4, and 5 present; fork 2 sessile, length about 1.5 times length of fork 3; fork 3 with length of fork about twice length of footstalk, fork 4 about 0.8 times length of fork 3, fork 5 about 1.6–1.7 times length of fork 4. Hind wings each with forks 2, 3, and 5 present; fork 2 sessile, its length about 1.7 times length of fork 3.
</paragraph>
<caption id="DF2171A4FFF3682E45CA333ABBC0F973" ID-DOI="http://doi.org/10.5281/zenodo.6559743" ID-Zenodo-Dep="6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289" startId="6.[151,264,1585,1609]" targetBox="[362,1223,356,1555]" targetPageId="6">
<paragraph id="8BE1212CFFF3682E45CA333ABBC0F973" blockId="6.[151,1437,1585,1790]" pageId="6" pageNumber="289">
<emphasis id="B92AFD3EFFF3682E45CA333ABB32F9C7" bold="true" box="[151,329,1585,1610]" pageId="6" pageNumber="289">FIGURES 6–12.</emphasis>
<taxonomicName id="4C5E5AAFFFF3682E440D3339BBBEF9C7" authorityName="Ulmer" authorityYear="1903" box="[336,453,1586,1610]" class="Insecta" family="Ecnomidae" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="family">Ecnomidae</taxonomicName>
. Figures 6–9,
<taxonomicName id="4C5E5AAFFFF3682E47053339B92FF9C7" authority="Cartwright &amp; Dostine, 2022" authorityName="Cartwright &amp; Dostine" authorityYear="2022" box="[600,852,1586,1610]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="topendica" status="sp. nov.">
<emphasis id="B92AFD3EFFF3682E47053339B92FF9C7" box="[600,852,1586,1610]" italics="true" pageId="6" pageNumber="289">Neboissomina topendica</emphasis>
</taxonomicName>
<emphasis id="B92AFD3EFFF3682E46063339B9D6F9C7" bold="true" box="[859,941,1586,1610]" pageId="6" pageNumber="289">
<taxonomicNameLabel id="A2194045FFF3682E46063339B9D6F9C7" box="[859,941,1586,1610]" pageId="6" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12,
<taxonomicName id="4C5E5AAFFFF3682E4625335DBE2BF9E3" authority="Cartwright &amp; Dostine, 2022" authorityName="Cartwright &amp; Dostine" authorityYear="2022" box="[888,1104,1622,1646]" class="Insecta" family="Ecnomidae" genus="Wellsomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="tamoides" status="sp. nov.">
<emphasis id="B92AFD3EFFF3682E4625335DBE2BF9E3" box="[888,1104,1622,1646]" italics="true" pageId="6" pageNumber="289">Wellsomina tamoides</emphasis>
</taxonomicName>
<emphasis id="B92AFD3EFFF3682E4104335DBED0F9E3" bold="true" box="[1113,1195,1622,1646]" pageId="6" pageNumber="289">
<taxonomicNameLabel id="A2194045FFF3682E4104335DBED0F9E3" box="[1113,1195,1622,1646]" pageId="6" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male genitalia: 10, left lateral; 11, ventral; 12, dorsal. Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired).
</paragraph>
</caption>
<paragraph id="8BE1212CFFF3682F459A3227B9A2FF74" blockId="6.[151,1437,1836,2042]" lastBlockId="7.[151,1436,150,393]" lastPageId="7" lastPageNumber="290" pageId="6" pageNumber="289">
<emphasis id="B92AFD3EFFF3682E459A3227BB73F8CB" box="[199,264,1836,1862]" italics="true" pageId="6" pageNumber="289">Male.</emphasis>
Tergum X membranous, with pair of long, slender dorsal sclerotized processes converging near middle and diverging posteriorly (
<figureCitation id="13653DA9FFF3682E449F325BB87CF8E7" box="[450,519,1872,1898]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Fig. 9</figureCitation>
). Superior appendages short and robust (
<figureCitation id="13653DA9FFF3682E468E325BBE43F8E7" box="[979,1080,1872,1898]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Figs 7, 9</figureCitation>
), in lateral view appearing subrectangular, very slightly tapered posteriorly, length about 2.8 times width, apices slightly rounded (
<figureCitation id="13653DA9FFF3682E41B8327FBF53F803" box="[1253,1320,1908,1934]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Fig. 7</figureCitation>
), in dorsal view slightly dilated in posterior third (
<figureCitation id="13653DA9FFF3682E47143293B8F7F83F" box="[585,652,1944,1970]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Fig. 9</figureCitation>
); pair of long, slender curved processes (mesal processes of tergum X) ventral to superior appendages. Phallus simple, tube-like (
<figureCitation id="13653DA9FFF3682E464632B7B906F85B" box="[795,893,1980,2006]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Figs 7, 8</figureCitation>
) with elongate, slender process partly embedded and emergent posteriorly (
<figureCitation id="13653DA9FFF3682E449D32EBB87DF877" box="[448,518,2016,2042]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Fig. 7</figureCitation>
). Inferior appendages slightly longer than superior appendages, fused (
<figureCitation id="13653DA9FFF3682E407532EBBFF4F877" box="[1320,1423,2016,2042]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="6" pageNumber="289">Figs 7–9</figureCitation>
), in ventral view large, subrectangular shield, truncate posteriorly, length about 1.5 times width, with acute apicolateral corners and acute apicomesal projection (
<figureCitation id="13653DA9FFF2682F47FD35B0B89EFF58" box="[672,741,187,213]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="7" pageNumber="290">Fig. 8</figureCitation>
), in lateral view robust, tapered basally, very slightly tapered posteriorly, appearing sub-trapezoidal, length about 3 times width (
<figureCitation id="13653DA9FFF2682F46DA35D4B9B0FF74" box="[903,971,223,249]" captionStart="FIGURES 6–12" captionStartId="6.[151,264,1585,1609]" captionTargetBox="[362,1223,356,1555]" captionTargetId="figure-132@6.[352,1235,345,1562]" captionTargetPageId="6" captionText="FIGURES 6–12. Ecnomidae. Figures 6–9, Neboissomina topendica sp. nov.: 6, apices of right fore- and hind wings, dorsal. 7–9, male genitalia: 7, left lateral; 8, ventral; 9, dorsal. Figures 10–12, Wellsomina tamoides sp. nov. male genitalia: 10, left lateral; 11, ventral; 12, dorsal.Abbreviations: F2 = fork 2; F3 = fork 3; F4 = fork 4; F5 = fork 5; ap. sp. X = apical spine tergum X; inf. app. = inferior appendage (paired); mes. pro. terg. X = mesal process tergum X (paired); pha. = phallus; sup. app. = superior appendage (paired); sup. app. dors. = dorsal branch of superior appendage (paired); sup. app. vent. = ventral branch of superior appendage (paired)." figureDoi="http://doi.org/10.5281/zenodo.6559743" httpUri="https://zenodo.org/record/6559743/files/figure.png" pageId="7" pageNumber="290">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BE1212CFFF2682F459A3408BBE4FE90" blockId="7.[151,1436,150,393]" box="[199,415,259,285]" pageId="7" pageNumber="290">
<emphasis id="B92AFD3EFFF2682F459A3408BB58FE90" box="[199,291,259,285]" italics="true" pageId="7" pageNumber="290">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF2682F459A342CBF55FECC" box="[199,1326,295,321]" pageId="7" pageNumber="290" type="etymology">
<paragraph id="8BE1212CFFF2682F459A342CBF55FECC" blockId="7.[151,1436,150,393]" box="[199,1326,295,321]" pageId="7" pageNumber="290">
<emphasis id="B92AFD3EFFF2682F459A342CBB35FECC" bold="true" box="[199,334,295,321]" pageId="7" pageNumber="290">Etymology.</emphasis>
<emphasis id="B92AFD3EFFF2682F4408342CBBB2FECC" box="[341,457,295,321]" italics="true" pageId="7" pageNumber="290">Topendica</emphasis>
—based on the colloquial regional name for the locality (Top End of the NT).
</paragraph>
</subSubSection>
<subSubSection id="C34472A7FFF2682F459A3440BE30FE04" pageId="7" pageNumber="290" type="discussion">
<paragraph id="8BE1212CFFF2682F459A3440BE30FE04" blockId="7.[151,1436,150,393]" pageId="7" pageNumber="290">
<emphasis id="B92AFD3EFFF2682F459A3440BB40FEE8" bold="true" box="[199,315,331,357]" pageId="7" pageNumber="290">Remarks.</emphasis>
At least thirty males of
<taxonomicName id="4C5E5AAFFFF2682F47123440B8A4FEE8" authorityName="Cartwright &amp; Dostine" authorityYear="2022" box="[591,735,331,357]" class="Insecta" family="Ecnomidae" genus="Neboissomina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="topendica">
<emphasis id="B92AFD3EFFF2682F47123440B8A4FEE8" box="[591,735,331,357]" italics="true" pageId="7" pageNumber="290">N. topendica</emphasis>
</taxonomicName>
have been collected from 3 localities in the Top End (latitude range 12.80–
<geoCoordinate id="EE6A47EBFFF2682F44773465BBF0FE04" box="[298,395,366,393]" degrees="14.59" direction="south" orientation="latitude" pageId="7" pageNumber="290" precision="555" value="-14.59">14.59 °S</geoCoordinate>
), including sites in Kakadu NP and in the greater Darwin area.
</paragraph>
</subSubSection>
</treatment>
</document>
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