387 lines
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387 lines
49 KiB
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<document ID-DOI="10.5281/zenodo.3382576" ID-GBIF-Dataset="48817a66-49e4-4e6f-b6c6-879db64b9ec9" ID-Zenodo-Dep="3382576" approvalRequired="25" approvalRequired_for_bibRefs="5" approvalRequired_for_taxonomicNames="20" checkinTime="1567247981485" checkinUser="plazi" docAuthor="Rauhut, Oliver W. M." docDate="2003" docId="77323C29FFE6B413FF1C9ADFFE1BF6AC" docLanguage="en" docName="Rauhut2003BasalTheropod_ocr.pdf" docOrigin="Special papers in palaeontology 69" docStyle="DocumentStyle{}" docTitle="Abelisauridae Bonaparte and Novas 1985" docType="treatment" docVersion="15" lastPageNumber="34" masterDocId="8B0B4451FFC7B431FFF59E70FFF0FFFF" masterDocTitle="The interrelationships and evolution of basal theropod dinosaurs" masterLastPageNumber="213" masterPageNumber="1" pageNumber="33" updateTime="1663160796219" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The interrelationships and evolution of basal theropod dinosaurs</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rauhut, Oliver W. M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Special papers in palaeontology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2003</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2003-05-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>213</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3382576</mods:identifier>
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<mods:identifier type="GBIF-Dataset">48817a66-49e4-4e6f-b6c6-879db64b9ec9</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3382576</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5123226" ID-GBIF-Taxon="159404301" ID-Zenodo-Dep="5123226" LSID="urn:lsid:plazi:treatment:77323C29FFE6B413FF1C9ADFFE1BF6AC" httpUri="http://treatment.plazi.org/id/77323C29FFE6B413FF1C9ADFFE1BF6AC" lastPageId="34" lastPageNumber="34" pageId="33" pageNumber="33">
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<subSubSection box="[233,947,1199,1235]" pageId="33" pageNumber="33" type="nomenclature">
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<paragraph blockId="33.[231,2001,1140,1456]" box="[233,947,1199,1235]" pageId="33" pageNumber="33">
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<taxonomicName authority="Bonaparte and Novas, 1985" authorityName="Bonaparte and Novas" authorityYear="1985" box="[233,947,1199,1235]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="family">Abelisauridae Bonaparte and Novas, 1985</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="33" type="synonymic_list">
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<paragraph blockId="33.[231,2001,1140,1456]" pageId="33" pageNumber="33">
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<emphasis box="[231,881,1257,1291]" italics="true" pageId="33" pageNumber="33">
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Included taxa.
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<taxonomicName authority="Bonaparte and Novas, 1985" authorityName="Bonaparte and Novas" authorityYear="1985" box="[483,1363,1257,1291]" class="Reptilia" family="Abelisauridae" genus="Abelisaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="comahuensis">Abelisaurus comahuensis Bonaparte and Novas, 1985</taxonomicName>
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</emphasis>
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;
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<taxonomicName authority="Bonaparte, 1985" authorityName="Bonaparte" authorityYear="1985" box="[1395,1992,1257,1291]" class="Reptilia" family="Abelisauridae" genus="Camotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="sastrei">
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<emphasis box="[1395,1705,1257,1291]" italics="true" pageId="33" pageNumber="33">Camotaurus sastrei</emphasis>
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<bibRefCitation author="Bonaparte" box="[1729,1992,1257,1291]" firstAuthor="Bonaparte" journalOrPublisher="National Geographic Research" pageId="33" pageNumber="33" pagination="149 - 151" part="1" refId="ref129212" refString="------ 1985. A homed Cretaceous camosaur from Patagonia. National Geographic Research, 1, 149 - 151." title="A homed Cretaceous camosaur from Patagonia" type="journal article" year="1985">Bonaparte, 1985</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName authority="Accarie, Beaudoin, Dejax, Fries, Michard and Taquet, 1995" authorityName="Accarie, Beaudoin, Dejax, Fries, Michard and Taquet" box="[234,1538,1298,1333]" class="Reptilia" family="Abelisauridae" genus="Genusaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="sisteronis">
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<emphasis box="[234,574,1298,1332]" italics="true" pageId="33" pageNumber="33">Genusaurus sisteronis</emphasis>
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<bibRefCitation author="Accarie, Beaudoin, Dejax, Fries, Michard and Taquet" box="[590,1538,1298,1333]" firstAuthor="Accarie" journalOrPublisher="Comptes Rendus de FAcademie des Sciences de Paris, Serie II" pageId="33" pageNumber="33" pagination="327 - 334" part="320" refId="ref127729" refString="ACCARIE, H " BEAUDOIN, B., DEJAX, J., FRIES, G " MICHARD, J. - G. and TAQUET, P. 1995. Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien. Comptes Rendus de FAcademie des Sciences de Paris, Serie II, 320, 327 - 334." title="Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien" type="journal article" year="1995">Accarie, Beaudoin, Dejax, Friès, Michard and Taquet, 1995</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName authority="Coria and Salgado, 2000" authorityName="Coria and Salgado" authorityYear="2000" class="Reptilia" family="Abelisauridae" genus="Ilokelesia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="aguadagrandensis">
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<emphasis box="[1560,2001,1298,1332]" italics="true" pageId="33" pageNumber="33">Ilokelesia aguadagrandensis</emphasis>
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<bibRefCitation author="Coria and Salgado" box="[232,640,1340,1374]" firstAuthor="Coria" journalOrPublisher="Gaia" pageId="33" pageNumber="33" pagination="89 - 102" part="15" refId="ref131568" refString="------------- 2000. A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina. Gaia, 15, 89 - 102." title="A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina" type="journal article" year="2000">Coria and Salgado, 2000</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName authority="Huene, 1932" authorityName="Huene" authorityYear="1932" box="[667,1222,1340,1374]" class="Reptilia" family="Abelisauridae" genus="Indosuchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="raptorius">
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<emphasis box="[667,997,1340,1374]" italics="true" pageId="33" pageNumber="33">Indosuchus raptorius</emphasis>
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<bibRefCitation author="Huene" box="[1018,1222,1340,1374]" firstAuthor="Huene" journalOrPublisher="Monographien zur Geologie und Palaeontologie, Serie I" pageId="33" pageNumber="33" pagination="1 - 361" part="4" refId="ref135106" refString="------ 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. Monographien zur Geologie und Palaeontologie, Serie I, 4, 1 - 361." title="Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte" type="journal article" year="1932">Huene, 1932</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName authority="Sues and Taquet, 1979" authorityName="Sues and Taquet" authorityYear="1979" box="[1249,1992,1340,1374]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="atopus">
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<emphasis box="[1249,1596,1340,1374]" italics="true" pageId="33" pageNumber="33">Majungatholus atopus</emphasis>
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Sues and Taquet, 1979
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</taxonomicName>
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;
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<taxonomicName authority="Sampson, Carrano and Forster, 2001" authorityName="Sampson, Carrano and Forster" authorityYear="2001" box="[231,1234,1381,1416]" class="Reptilia" family="Noasauridae" genus="Masiakasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="knopfleri">
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<emphasis box="[231,617,1381,1415]" italics="true" pageId="33" pageNumber="33">Masiakasaurus knopfleri</emphasis>
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Sampson, Carrano and Forster, 2001
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</taxonomicName>
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;
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<taxonomicName authority="Bonaparte and Powell, 1980" authorityName="Bonaparte and Powell" authorityYear="1980" box="[1261,1992,1381,1415]" class="Reptilia" family="Noasauridae" genus="Noasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="leali">
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<emphasis box="[1261,1510,1381,1415]" italics="true" pageId="33" pageNumber="33">Noasaurus leali</emphasis>
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Bonaparte and Powell, 1980
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</taxonomicName>
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;
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<taxonomicName authority="Martinez, Gimenez, Rodriguez and Bochatey, 1986" authorityName="Martinez, Gimenez, Rodriguez and Bochatey" authorityYear="1986" box="[231,1458,1422,1456]" class="Reptilia" family="Abelisauridae" genus="Xenotarsosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="bonapartei">
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<emphasis box="[231,664,1422,1456]" italics="true" pageId="33" pageNumber="33">Xenotarsosaurus bonapartei</emphasis>
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Martinez, Giménez, Rodriguez and Bochatey, 1986
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="33" type="distribution">
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<paragraph blockId="33.[234,839,1513,1547]" box="[234,839,1513,1547]" pageId="33" pageNumber="33">
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<emphasis box="[234,484,1513,1547]" italics="true" pageId="33" pageNumber="33">Temporal range.</emphasis>
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Albian-Maastrichtian.
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</paragraph>
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<paragraph blockId="33.[232,1998,1602,1761]" pageId="33" pageNumber="33">
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<emphasis box="[234,418,1602,1636]" italics="true" pageId="33" pageNumber="33">Occurrence.</emphasis>
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Allen Formation, Rio Negro, Argentina; La Colonia Formation, Chubut, Argentina; unnamed unit (green clays and glauconitic sands of Bevons), Alpes de Haute-Provence, France; Lameta Formation, Madhya Pradesh, India; Maevarano Formation, Majunga, Madagascar; Lecho Formation, Salta, Argentina; Bajo-Barreal Formation, Chubut, Argentina.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="33" type="diagnosis">
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<paragraph blockId="33.[233,1999,1810,2029]" pageId="33" pageNumber="33">
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<emphasis box="[233,411,1810,1846]" italics="true" pageId="33" pageNumber="33">Diagnosis.</emphasis>
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Maxilla in front of the antorbital opening very short and high; posterior border of lacrimal convex over its entire length; anterior process of lacrimal reduced; jugal process of postorbital expanded, strongly anteriorly directed and intrudes into the orbit (convergently present in
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<taxonomicName baseAuthorityName="Maleev" baseAuthorityYear="1955" box="[1591,1928,1902,1938]" class="Reptilia" family="Tyrannosauridae" genus="Tarbosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="bataar">
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<emphasis box="[1591,1928,1902,1938]" italics="true" pageId="33" pageNumber="33">Tarbosaurus bataar</emphasis>
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</taxonomicName>
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and
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<emphasis box="[235,560,1947,1983]" italics="true" pageId="33" pageNumber="33">
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<taxonomicName authorityName="Osborn" authorityYear="1905" box="[235,554,1947,1983]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="rex">Tyrannosaurus rex</taxonomicName>
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,
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</emphasis>
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but not in other tyrannosaurids); epipophyses in the cervical vertebrae hypertrophied and higher than the neural spines.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="34" lastPageNumber="34" pageId="33" pageNumber="33" type="discussion">
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<paragraph blockId="33.[231,1999,2075,2567]" pageId="33" pageNumber="33">
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<emphasis box="[231,387,2075,2111]" italics="true" pageId="33" pageNumber="33">Remarks.</emphasis>
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Abelisaurids have recently been recognized as a distinct family of theropod dinosaurs (Bonaparte and Novas 1985). Their diagnosis and taxonomy is still problematic, mainly due to the fragmentary nature of the remains of all members of the family with the exception of
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<emphasis box="[1789,1999,2167,2203]" italics="true" pageId="33" pageNumber="33">Camotaurus</emphasis>
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(
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<bibRefCitation author="Bonaparte" box="[245,644,2213,2249]" etAl="et al." firstAuthor="Bonaparte" journalOrPublisher="Contributions in Science" pageId="33" pageNumber="33" pagination="1 - 42" part="416" refId="ref129397" refString="------------- and CORIA, r. a. 1990. Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia. Contributions in Science, 416, 1 - 42." title="Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia" type="journal article" year="1990">
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Bonaparte
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<emphasis box="[441,532,2213,2249]" italics="true" pageId="33" pageNumber="33">et al.</emphasis>
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1990
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</bibRefCitation>
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) and recently described new specimens of
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<taxonomicName authority="(Sampson et al. 1998)" authorityName="Sampson" authorityYear="1998" baseAuthorityName="Sampson" baseAuthorityYear="1998" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="genus">
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<emphasis box="[1440,1694,2213,2249]" italics="true" pageId="33" pageNumber="33">Majungatholus</emphasis>
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(Sampson
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<emphasis box="[1905,1995,2213,2249]" italics="true" pageId="33" pageNumber="33">et al.</emphasis>
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1998)
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</taxonomicName>
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. Given the many peculiarities in the skeleton of these genera, it seems very probable that the discovery of more complete material of other taxa will provide more diagnostic characters for this family.
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</paragraph>
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<paragraph blockId="33.[231,1999,2075,2567]" lastBlockId="34.[285,2062,201,2387]" lastPageId="34" lastPageNumber="34" pageId="33" pageNumber="33">
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<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[276,609,2395,2431]" class="Reptilia" family="Abelisauridae" genus="Camotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="sastrei">
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<emphasis box="[276,609,2395,2431]" italics="true" pageId="33" pageNumber="33">Camotaurus sastrei</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[630,830,2395,2431]" captionStart="text" captionStartId="38.[281,355,1923,1950]" captionTargetBox="[279,1963,430,2567]" captionTargetPageId="39" captionText="text-fig. 6. Skull reconstructions of representatives of Cretaceous OTUs in left lateral view, a, abelisaurid Camotaurus sastrei, Late Cretaceous (Campanian), Gorro Frigio Formation, Argentina; based on Bonaparte et al. (1990) and MACN CH 894. b, composite skull of a generalized baryonychid; mainly based on Suchomimus tenerensis (MNN GDF 501, 503-506) and Irritator challenged (SMNS 58022), with some elements reconstructed after Baryonyx walkeri (BMNH R 9951). c, carcharodontosaurid Carcharodontosaurus sahariens, Late Cretaceous (Cenomanian), Kem Kem beds, Morocco; based on Sereno et al. (1996) and SGM-Din 1, unpreserved elements shaded. D, dromaeosaurid Velociraptor mongoliensis, Late Cretaceous (Campanian), Djadokhta Formation, Mongolia; based on Barsbold (1983), Barsbold and Osmólska (1999), and AMNH 6515. e, ornithomimosaur Dromiceiomimus brevitertius, Late Cretaceous (Maastrichtian), Horseshoe Canyon Formation, Alberta, Canada; redrawn from Russell (1972). F, oviraptorosaur Oviraptor philoceratops, Late Cretaceous (Campanian), Djadokhta Formation, Mongolia; redrawn from Barsbold et al. (1990). G, therizinosauroid Erlikosaurus andrewsi, Late Cretaceous (Cenomanian- Turonian), Baynshiren Svita, Mongolia; based on Clark et al. (1994). H, tyrannosaurid Gorgosaurus libratus, Late Cretaceous (Campanian), Judith River Formation, Alberta, Canada, redrawn from Molnar et al. (1990). Abbreviations as in Text-figures 4 and 5, and: acc. op., accessory opening of unknown function in the premaxilla of Oviraptor (Barsbold et al. 1990); amf, anterior maxillary foramen; pn, pneumatic openings. Scale bars represent 50 mm (d-g) and 100 mm (a-c, h)." httpUri="https://zenodo.org/record/3382586/files/figure.png" pageId="33" pageNumber="33" targetBox="[279,1963,430,2567]" targetPageId="39">Text-fig. 6a</figureCitation>
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),
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<emphasis box="[862,1244,2395,2431]" italics="true" pageId="33" pageNumber="33">
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<taxonomicName authorityName="Sues and Taquet" authorityYear="1979" box="[862,1238,2395,2431]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="atopus">Majungatholus atopus</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[1321,1747,2395,2431]" class="Reptilia" family="Abelisauridae" genus="Abelisaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="species" species="comahuensis">
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<emphasis box="[1321,1747,2395,2431]" italics="true" pageId="33" pageNumber="33">Abelisaurus comahuensis</emphasis>
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</taxonomicName>
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share all of the cranial synapomorphies listed above, and so their referral to the
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<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[1294,1524,2440,2476]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="33" pageNumber="33" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
is well supported. However, many other, very incompletely known taxa have been assigned to this family (
|
||
|
|
<bibRefCitation author="Martinez" box="[1626,1992,2485,2521]" etAl="et al." firstAuthor="Martinez" journalOrPublisher="Aetas del IV Congreso Argentino de Paleontologia y Biostratigrafia" pageId="33" pageNumber="33" pagination="23 - 31" part="2" refId="ref137698" refString="MARTINEZ, R., GIMENEZ, o., rodri ' guez, J. and BOCHATEY, G. 1986. Xenotarsosaurus bonapartei nov. gen. et sp. (Camosauria, Abelisauridae), un nuevo Theropoda de la Formacion Bajo Barreal, Chubut, Argentina. Aetas del IV Congreso Argentino de Paleontologia y Biostratigrafia, 2, 23 - 31." title="Xenotarsosaurus bonapartei nov. gen. et sp. (Camosauria, Abelisauridae), un nuevo Theropoda de la Formacion Bajo Barreal, Chubut, Argentina" type="journal article" year="1986">
|
||
|
|
Martinez
|
||
|
|
<emphasis box="[1798,1884,2485,2521]" italics="true" pageId="33" pageNumber="33">et al.</emphasis>
|
||
|
|
1986
|
||
|
|
</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation author="Bonaparte" box="[233,597,2531,2567]" etAl="et al." firstAuthor="Bonaparte" journalOrPublisher="Contributions in Science" pageId="33" pageNumber="33" pagination="1 - 42" part="416" refId="ref129397" refString="------------- and CORIA, r. a. 1990. Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia. Contributions in Science, 416, 1 - 42." title="Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia" type="journal article" year="1990">
|
||
|
|
Bonaparte
|
||
|
|
<emphasis box="[416,495,2531,2567]" italics="true" pageId="33" pageNumber="33">et al.</emphasis>
|
||
|
|
1990
|
||
|
|
</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation author="Molnar" bookContentInfo="733 pp." box="[615,836,2531,2567]" editor="weishampel, d. b & dodson, p. & osmOlska, h." firstAuthor="Molnar" journalOrPublisher="University of California Press, Berkeley" pageId="33" pageNumber="33" pagination="306 - 317" refId="ref137890" refString="------ 1990. Problematic Theropoda: ' Camosaurs'. 306 - 317. In weishampel, d. b " dodson, p. and osmOlska, h. (eds). The Dinosauria. University of California Press, Berkeley, 733 pp." title="Problematic Theropoda: ' Camosaurs'" type="book chapter" volumeTitle="The Dinosauria" year="1990">Molnar 1990</bibRefCitation>
|
||
|
|
; Bonaparte 1991
|
||
|
|
<emphasis box="[1121,1142,2531,2567]" italics="true" pageId="33" pageNumber="33">b</emphasis>
|
||
|
|
; Le Loeuff and Buffetaut 1991), and many of these assignments are problematical. Therefore, a short review of poorly known taxa that are included in the
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[289,522,247,283]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
here is given below.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Bonaparte and Powell" authorityYear="1980" box="[329,599,293,329]" class="Reptilia" family="Noasauridae" genus="Noasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="leali">
|
||
|
|
<emphasis box="[329,599,293,329]" italics="true" pageId="34" pageNumber="34">Noasaurus leali</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
was originally assigned to its own family,
|
||
|
|
<taxonomicName authority="(Bonaparte and Powell 1980)" authorityName="Bonaparte and Powell" authorityYear="1980" baseAuthorityName="Bonaparte and Powell" baseAuthorityYear="1980" box="[1333,2053,293,329]" class="Reptilia" family="Noasauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Noasauridae (Bonaparte and Powell 1980)</taxonomicName>
|
||
|
|
. Later,
|
||
|
|
<bibRefCitation author="Bonaparte" box="[399,675,339,375]" firstAuthor="Bonaparte" journalOrPublisher="Historical Biology" pageId="34" pageNumber="34" pagination="1 - 25" part="5" refId="ref129272" refString="------ 1991 a. The Gondwanian theropod families Abelisauridae and Noasauridae. Historical Biology, 5, 1 - 25." title="The Gondwanian theropod families Abelisauridae and Noasauridae" type="journal article" year="1991">Bonaparte (1991</bibRefCitation>
|
||
|
|
/?) noted the close similarities between
|
||
|
|
<taxonomicName authorityName="Bonaparte & Powell" authorityYear="1980" box="[1333,1515,339,375]" class="Reptilia" family="Noasauridae" genus="Noasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[1333,1515,339,375]" italics="true" pageId="34" pageNumber="34">Noasaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and abelisaurids and created the superfamily Abelisauroidea to include these two families (
|
||
|
|
<bibRefCitation author="Bonaparte" box="[1275,1541,384,420]" firstAuthor="Bonaparte" journalOrPublisher="Historical Biology" pageId="34" pageNumber="34" pagination="1 - 25" part="5" refId="ref129272" refString="------ 1991 a. The Gondwanian theropod families Abelisauridae and Noasauridae. Historical Biology, 5, 1 - 25." title="The Gondwanian theropod families Abelisauridae and Noasauridae" type="journal article" year="1991">Bonaparte 1991</bibRefCitation>
|
||
|
|
/?, p. 22).
|
||
|
|
<taxonomicName authorityName="Bonaparte & Powell" authorityYear="1980" box="[1702,1884,384,420]" class="Reptilia" family="Noasauridae" genus="Noasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[1702,1884,384,420]" italics="true" pageId="34" pageNumber="34">Noasaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
shares the first diagnostic character with
|
||
|
|
<emphasis box="[799,1246,431,467]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[799,1010,431,467]" class="Reptilia" family="Abelisauridae" genus="Abelisaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Abelisaurus,</taxonomicName>
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[1025,1239,431,467]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Carnotaurus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
and
|
||
|
|
<emphasis box="[1330,1592,431,467]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[1330,1587,431,467]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Majungatholus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
and the fifth character with
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[290,502,476,512]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[290,502,476,512]" italics="true" pageId="34" pageNumber="34">Carnotaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<emphasis box="[582,845,476,512]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[582,839,476,512]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Majungatholus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
and may therefore be regarded as a close relative of these taxa. Since the differences between the
|
||
|
|
<taxonomicName baseAuthorityName="Bonaparte and Powell" baseAuthorityYear="1980" box="[702,910,522,558]" class="Reptilia" family="Noasauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Noasauridae</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[994,1229,522,558]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
, as defined by
|
||
|
|
<bibRefCitation author="Bonaparte" box="[1482,1760,522,558]" firstAuthor="Bonaparte" journalOrPublisher="Historical Biology" pageId="34" pageNumber="34" pagination="1 - 25" part="5" refId="ref129272" refString="------ 1991 a. The Gondwanian theropod families Abelisauridae and Noasauridae. Historical Biology, 5, 1 - 25." title="The Gondwanian theropod families Abelisauridae and Noasauridae" type="journal article" year="1991">Bonaparte (1991</bibRefCitation>
|
||
|
|
/?), are only slight and may partly reflect size-dependant and/or ontogenetic differences, the recognition of two families is not accepted here. Although
|
||
|
|
<bibRefCitation author="Coria and Salgado" box="[704,1127,614,650]" firstAuthor="Coria" journalOrPublisher="Gaia" pageId="34" pageNumber="34" pagination="89 - 102" part="15" refId="ref131568" refString="------------- 2000. A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina. Gaia, 15, 89 - 102." title="A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina" type="journal article" year="2000">Coria and Salgado (2000)</bibRefCitation>
|
||
|
|
presented a cladogram of neoceratosaurs supporting the distinction of noasaurids and abelisaurids, their results must be seen as preliminary since only some taxa of the Abelisauria were included, characters were restricted to synapomorphies defining each node, and the majority of abelisaurid taxa considered in this analysis are known from less than 20 per cent of the skeleton. A more inclusive analysis might yield quite different results; therefore, all abelisaurs known are included in a single clade,
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[755,990,842,878]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
, here. However, apart from
|
||
|
|
<emphasis box="[1479,1669,842,878]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Bonaparte & Powell" authorityYear="1980" box="[1479,1663,842,878]" class="Reptilia" family="Noasauridae" genus="Noasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Noasaurus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
the recently described
|
||
|
|
<taxonomicName box="[288,544,888,924]" class="Reptilia" family="Noasauridae" genus="Masiakasaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[288,544,888,924]" italics="true" pageId="34" pageNumber="34">Masiakasaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
also indicates an important diversity of small abelisaurids (
|
||
|
|
<bibRefCitation author="Sampson" box="[1564,1917,888,924]" etAl="et al." firstAuthor="Sampson" pageId="34" pageNumber="34" pagination="504 - 506" refId="ref141850" refString="SAMPSON, s. d., carrano, m.. and forster, c. a. 2001. A bizarre predatory dinosaur from the Late Cretaceous of Madagascar. Nature, 409, 504 - 506." type="journal article" year="2001">
|
||
|
|
Sampson
|
||
|
|
<emphasis box="[1731,1815,888,924]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
2001
|
||
|
|
</bibRefCitation>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
Although based on very fragmentary material,
|
||
|
|
<taxonomicName authorityName="Coria and Salgado" authorityYear="2000" box="[1123,1606,933,969]" class="Reptilia" family="Abelisauridae" genus="Ilokelesia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="aguadagrandensis">
|
||
|
|
<emphasis box="[1123,1606,933,969]" italics="true" pageId="34" pageNumber="34">Ilokelesia aguadagrandensis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
can be demonstrated to be an abelisaur with some certainty, since it shares the pecularities of the postorbital and the hypertrophied cervical epipophyses with other abelisaurid taxa (
|
||
|
|
<bibRefCitation author="Coria and Salgado" box="[1133,1549,1025,1061]" firstAuthor="Coria" journalOrPublisher="Gaia" pageId="34" pageNumber="34" pagination="89 - 102" part="15" refId="ref131568" refString="------------- 2000. A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina. Gaia, 15, 89 - 102." title="A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina" type="journal article" year="2000">Coria and Salgado 2000</bibRefCitation>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Huene" authorityYear="1932" box="[328,688,1071,1107]" class="Reptilia" family="Abelisauridae" genus="Indosuchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="raptorius">
|
||
|
|
<emphasis box="[328,688,1071,1107]" italics="true" pageId="34" pageNumber="34">Indosuchus raptorius</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
was described by
|
||
|
|
<bibRefCitation author="Huene" box="[1009,1239,1071,1107]" firstAuthor="Huene" journalOrPublisher="Monographien zur Geologie und Palaeontologie, Serie I" pageId="34" pageNumber="34" pagination="1 - 361" part="4" refId="ref135106" refString="------ 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. Monographien zur Geologie und Palaeontologie, Serie I, 4, 1 - 361." title="Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte" type="journal article" year="1932">Huene (1932)</bibRefCitation>
|
||
|
|
and Huene and Matley (1933) on the basis of a posterior end of a skull roof from the Maastrichtian Lameta Formation of India, and more material was later referred to this taxon (
|
||
|
|
<bibRefCitation author="Chatterjee" box="[759,1034,1163,1199]" firstAuthor="Chatterjee" journalOrPublisher="Journal of Paleontology" pageId="34" pageNumber="34" pagination="570 - 580" part="52" refId="ref130557" refString="CHATTERJEE, s. 1978. Indosuchus and Indosaurus, Cretaceous camosaurs from India. Journal of Paleontology, 52, 570 - 580." title="Indosuchus and Indosaurus, Cretaceous camosaurs from India" type="journal article" year="1978">Chatterjee 1978</bibRefCitation>
|
||
|
|
; Chatterjee and Rudra 1996). Sometimes considered to be a tyrannosaurid (
|
||
|
|
<bibRefCitation author="Chatterjee" box="[541,813,1209,1245]" firstAuthor="Chatterjee" journalOrPublisher="Journal of Paleontology" pageId="34" pageNumber="34" pagination="570 - 580" part="52" refId="ref130557" refString="CHATTERJEE, s. 1978. Indosuchus and Indosaurus, Cretaceous camosaurs from India. Journal of Paleontology, 52, 570 - 580." title="Indosuchus and Indosaurus, Cretaceous camosaurs from India" type="journal article" year="1978">Chatterjee 1978</bibRefCitation>
|
||
|
|
; Paul 1988
|
||
|
|
<emphasis box="[1004,1023,1209,1245]" italics="true" pageId="34" pageNumber="34">«</emphasis>
|
||
|
|
),
|
||
|
|
<taxonomicName box="[1053,1243,1209,1245]" class="Reptilia" family="Abelisauridae" genus="Indosuchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[1053,1243,1209,1245]" italics="true" pageId="34" pageNumber="34">Indosuchus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
has recently been placed in the
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[1774,2006,1209,1245]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
by several authors (Bonaparte and Novas 1985;
|
||
|
|
<bibRefCitation author="Buffetaut" box="[1082,1452,1254,1290]" etAl="et al." firstAuthor="Buffetaut" journalOrPublisher="Comptes Rendus de PAcademie des Sciences de Paris, Serie II" pageId="34" pageNumber="34" pagination="153 - 158" part="306" refId="ref129939" refString="------ MECHIN, p. and MECHIN-SALESSY, A. 1988. Un dinosaure theropode d'affinites gondwanniennes dans le Cretace superieur de Provence. Comptes Rendus de PAcademie des Sciences de Paris, Serie II, 306, 153 - 158." title="Un dinosaure theropode d'affinites gondwanniennes dans le Cretace superieur de Provence" type="journal article" year="1988">
|
||
|
|
Buffetaut
|
||
|
|
<emphasis box="[1258,1343,1254,1290]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1988
|
||
|
|
</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation author="Bonaparte" box="[1477,1860,1254,1290]" etAl="et al." firstAuthor="Bonaparte" journalOrPublisher="Contributions in Science" pageId="34" pageNumber="34" pagination="1 - 42" part="416" refId="ref129397" refString="------------- and CORIA, r. a. 1990. Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia. Contributions in Science, 416, 1 - 42." title="Camotaurus sastrei Bonaparte, the homed, lightly built camosaur from the Middle Cretaceous of Patagonia" type="journal article" year="1990">
|
||
|
|
Bonaparte
|
||
|
|
<emphasis box="[1667,1752,1254,1290]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1990
|
||
|
|
</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation author="Bonaparte" firstAuthor="Bonaparte" journalOrPublisher="Historical Biology" pageId="34" pageNumber="34" pagination="1 - 25" part="5" refId="ref129272" refString="------ 1991 a. The Gondwanian theropod families Abelisauridae and Noasauridae. Historical Biology, 5, 1 - 25." title="The Gondwanian theropod families Abelisauridae and Noasauridae" type="journal article" year="1991">Bonaparte 1991</bibRefCitation>
|
||
|
|
/?). Since there seem to be several large theropods in the Lameta Formation, the material referred to
|
||
|
|
<taxonomicName box="[287,479,1346,1382]" class="Reptilia" family="Abelisauridae" genus="Indosuchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[287,479,1346,1382]" italics="true" pageId="34" pageNumber="34">Indosuchus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
by
|
||
|
|
<bibRefCitation author="Chatterjee" box="[548,844,1346,1382]" firstAuthor="Chatterjee" journalOrPublisher="Journal of Paleontology" pageId="34" pageNumber="34" pagination="570 - 580" part="52" refId="ref130557" refString="CHATTERJEE, s. 1978. Indosuchus and Indosaurus, Cretaceous camosaurs from India. Journal of Paleontology, 52, 570 - 580." title="Indosuchus and Indosaurus, Cretaceous camosaurs from India" type="journal article" year="1978">Chatterjee (1978)</bibRefCitation>
|
||
|
|
cannot be shown to represent this taxon with certainty (
|
||
|
|
<bibRefCitation author="Molnar" bookContentInfo="733 pp." box="[1812,2038,1346,1382]" editor="weishampel, d. b & dodson, p. & osmOlska, h." firstAuthor="Molnar" journalOrPublisher="University of California Press, Berkeley" pageId="34" pageNumber="34" pagination="306 - 317" refId="ref137890" refString="------ 1990. Problematic Theropoda: ' Camosaurs'. 306 - 317. In weishampel, d. b " dodson, p. and osmOlska, h. (eds). The Dinosauria. University of California Press, Berkeley, 733 pp." title="Problematic Theropoda: ' Camosaurs'" type="book chapter" volumeTitle="The Dinosauria" year="1990">Molnar 1990</bibRefCitation>
|
||
|
|
). However, the type specimen shows the same arrangement of the supratemporal fenestrae as found in
|
||
|
|
<emphasis box="[285,774,1437,1473]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[285,490,1437,1473]" class="Reptilia" family="Abelisauridae" genus="Abelisaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Abelisaurus</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName box="[511,768,1437,1473]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Majungatholus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
and
|
||
|
|
<emphasis box="[864,1084,1437,1473]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[864,1074,1437,1473]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Carnotaurus</taxonomicName>
|
||
|
|
;
|
||
|
|
</emphasis>
|
||
|
|
thus, this species is referred to the
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[1693,1929,1437,1473]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[331,702,1483,1519]" class="Reptilia" family="Abelisauridae" genus="Genusaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="sisteronis">
|
||
|
|
<emphasis box="[331,702,1483,1519]" italics="true" pageId="34" pageNumber="34">Genusaurus sisteronis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
was described as a ceratosaurian theropod by
|
||
|
|
<bibRefCitation author="Accarie" box="[1467,1808,1483,1519]" etAl="et al." firstAuthor="Accarie" journalOrPublisher="Comptes Rendus de FAcademie des Sciences de Paris, Serie II" pageId="34" pageNumber="34" pagination="327 - 334" part="320" refId="ref127729" refString="ACCARIE, H " BEAUDOIN, B., DEJAX, J., FRIES, G " MICHARD, J. - G. and TAQUET, P. 1995. Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien. Comptes Rendus de FAcademie des Sciences de Paris, Serie II, 320, 327 - 334." title="Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien" type="journal article" year="1995">
|
||
|
|
Accarie
|
||
|
|
<emphasis box="[1607,1685,1483,1519]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
(1995)
|
||
|
|
</bibRefCitation>
|
||
|
|
on the basis of several vertebrae and an incomplete pelvis and hindlimb.
|
||
|
|
<taxonomicName box="[1293,1492,1529,1565]" class="Reptilia" family="Abelisauridae" genus="Genusaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[1293,1492,1529,1565]" italics="true" pageId="34" pageNumber="34">Genusaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
clearly represents a valid taxon, based on the peculiar morphology of the anterior end of the ilium (
|
||
|
|
<bibRefCitation author="Accarie" box="[1398,1718,1573,1610]" etAl="et al." firstAuthor="Accarie" journalOrPublisher="Comptes Rendus de FAcademie des Sciences de Paris, Serie II" pageId="34" pageNumber="34" pagination="327 - 334" part="320" refId="ref127729" refString="ACCARIE, H " BEAUDOIN, B., DEJAX, J., FRIES, G " MICHARD, J. - G. and TAQUET, P. 1995. Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien. Comptes Rendus de FAcademie des Sciences de Paris, Serie II, 320, 327 - 334." title="Decouverte d'un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l'Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurien" type="journal article" year="1995">
|
||
|
|
Accarie
|
||
|
|
<emphasis box="[1540,1618,1573,1610]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1995
|
||
|
|
</bibRefCitation>
|
||
|
|
, fig. 4a; the anterior end of this element seems to be lacking only minor parts, and its overall morphology is real, not a result of preservation; Carrano, pers. comm. 1999). Although none of the diagnostic characters of the abelisaurids is evident from the published illustrations,
|
||
|
|
<taxonomicName box="[998,1197,1712,1748]" class="Reptilia" family="Abelisauridae" genus="Genusaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[998,1197,1712,1748]" italics="true" pageId="34" pageNumber="34">Genusaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
is referred here to this clade, since it shares two apomorphic characters with
|
||
|
|
<emphasis box="[773,992,1758,1794]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[773,984,1758,1794]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Carnotaurus</taxonomicName>
|
||
|
|
:
|
||
|
|
</emphasis>
|
||
|
|
the almost completely straight dorsal margin of the ilium, and the very long and vertically oriented ischial peduncle of the ilium. These characters are not present in an ilium of
|
||
|
|
<taxonomicName box="[444,699,1848,1884]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[444,699,1848,1884]" italics="true" pageId="34" pageNumber="34">Majungatholus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
(UA 8678; Sampson
|
||
|
|
<emphasis box="[1087,1170,1848,1884]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1998, fig. 2f), indicating that, within abelisaurids,
|
||
|
|
<taxonomicName box="[289,490,1894,1930]" class="Reptilia" family="Abelisauridae" genus="Genusaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[289,490,1894,1930]" italics="true" pageId="34" pageNumber="34">Genusaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
may be more closely related to
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[1045,1255,1894,1930]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[1045,1255,1894,1930]" italics="true" pageId="34" pageNumber="34">Carnotaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
than to
|
||
|
|
<emphasis box="[1397,1660,1894,1930]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[1397,1655,1894,1930]" class="Reptilia" family="Pachycephalosauridae" genus="Majungatholus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Majungatholus</taxonomicName>
|
||
|
|
.
|
||
|
|
</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName box="[328,613,1940,1976]" class="Reptilia" family="Abelisauridae" genus="Xenotarsosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis box="[328,613,1940,1976]" italics="true" pageId="34" pageNumber="34">Xenotarsosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
is based on a dorsal vertebra and hindlimb elements, including femur, tibia, fibula, and astragalocalcaneum, from the Upper Cretaceous of Argentina (
|
||
|
|
<bibRefCitation author="Martinez" box="[1333,1675,1986,2022]" etAl="et al." firstAuthor="Martinez" journalOrPublisher="Aetas del IV Congreso Argentino de Paleontologia y Biostratigrafia" pageId="34" pageNumber="34" pagination="23 - 31" part="2" refId="ref137698" refString="MARTINEZ, R., GIMENEZ, o., rodri ' guez, J. and BOCHATEY, G. 1986. Xenotarsosaurus bonapartei nov. gen. et sp. (Camosauria, Abelisauridae), un nuevo Theropoda de la Formacion Bajo Barreal, Chubut, Argentina. Aetas del IV Congreso Argentino de Paleontologia y Biostratigrafia, 2, 23 - 31." title="Xenotarsosaurus bonapartei nov. gen. et sp. (Camosauria, Abelisauridae), un nuevo Theropoda de la Formacion Bajo Barreal, Chubut, Argentina" type="journal article" year="1986">
|
||
|
|
Martinez
|
||
|
|
<emphasis box="[1495,1573,1986,2022]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1986
|
||
|
|
</bibRefCitation>
|
||
|
|
). Although none of the diagnostic characters of the family as diagnosed here is found in the holotype, the taxon is tentatively referred to the
|
||
|
|
<taxonomicName authorityName="Bonaparte and Novas" authorityYear="1985" box="[561,796,2077,2113]" class="Reptilia" family="Abelisauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="family">Abelisauridae</taxonomicName>
|
||
|
|
, based on the great overall similarity of the preserved elements to the comparable elements of
|
||
|
|
<emphasis box="[708,927,2122,2158]" italics="true" pageId="34" pageNumber="34">
|
||
|
|
<taxonomicName authorityName="Bonaparte" authorityYear="1985" box="[708,922,2122,2158]" class="Reptilia" family="Abelisauridae" genus="Carnotaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="genus">Carnotaurus</taxonomicName>
|
||
|
|
.
|
||
|
|
</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
<paragraph blockId="34.[285,2062,201,2387]" pageId="34" pageNumber="34">
|
||
|
|
The recently described supposed abelisaurid
|
||
|
|
<taxonomicName authority="Le Loeuff and Buffetaut, 1991" authorityName="Le Loeuff and Buffetaut" authorityYear="1991" box="[1091,2047,2168,2204]" class="Reptilia" family="Abelisauridae" genus="Tarascosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="34" pageNumber="34" phylum="Chordata" rank="species" species="salluvicus">
|
||
|
|
<emphasis box="[1091,1516,2168,2204]" italics="true" pageId="34" pageNumber="34">Tarascosaurus salluvicus</emphasis>
|
||
|
|
Le Loeuff and Buffetaut, 1991
|
||
|
|
</taxonomicName>
|
||
|
|
, from the Upper Cretaceous of southern France, is based on extremely fragmentary material that does not show any diagnostic characters and is, therefore, regarded as a
|
||
|
|
<emphasis box="[1355,1611,2259,2295]" italics="true" pageId="34" pageNumber="34">nomen dubium.</emphasis>
|
||
|
|
However, the presence of abelisaurids in the Upper Cretaceous of southern Europe is also supported by other material (
|
||
|
|
<bibRefCitation author="Buffetaut" etAl="et al." firstAuthor="Buffetaut" journalOrPublisher="Comptes Rendus de PAcademie des Sciences de Paris, Serie II" pageId="34" pageNumber="34" pagination="153 - 158" part="306" refId="ref129939" refString="------ MECHIN, p. and MECHIN-SALESSY, A. 1988. Un dinosaure theropode d'affinites gondwanniennes dans le Cretace superieur de Provence. Comptes Rendus de PAcademie des Sciences de Paris, Serie II, 306, 153 - 158." title="Un dinosaure theropode d'affinites gondwanniennes dans le Cretace superieur de Provence" type="journal article" year="1988">
|
||
|
|
Buffetaut
|
||
|
|
<emphasis box="[287,369,2351,2387]" italics="true" pageId="34" pageNumber="34">et al.</emphasis>
|
||
|
|
1988
|
||
|
|
</bibRefCitation>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|