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<document id="E718BD7BD95E4B7A7231390CA95FFA1E" ID="10.11646/zootaxa.4169.1.8" ID-DOI="10.11646/zootaxa.4169.1.8" ID-GBIF-Dataset="a78cff31-17bb-4b28-bf38-cd97d4feb6b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="262091" ID-ZooBank="B54A654B-D5FA-4707-9C31-BAFAF98D5F53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474269705738" checkinUser="plazi" docAuthor="Borges-Nojosa, Diva Maria, Caramaschi, Ulisses &amp; Rodrigues, Miguel Trefaut" docDate="2016" docId="03AB93352A24FFF4C3FCF8C24A20599E" docLanguage="en" docName="zootaxa.4169.1.8.pdf" docOrigin="Zootaxa 4169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Placosoma limaverdorum Borges-Nojosa, Caramaschi &amp; Rodrigues, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="164" masterDocId="FF92EB4D2A25FFF0C36BFF974E385C67" masterDocTitle="A new species of lizard Placosoma Tschudi, 1847 (Squamata: Gymnophthalmidae) from the relictual forest mountains of the State of Ceará, Brazil" masterLastPageNumber="170" masterPageNumber="160" pageNumber="161" updateTime="1698386260174" updateUser="plazi">
<mods:mods id="B430CF8D904415872E3443F8DB1E9412" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9CBD1B8084019E12CC47FA7D5231A890">
<mods:title id="D8BFD87E70997F57A8F1FEFACB214F0E">A new species of lizard Placosoma Tschudi, 1847 (Squamata: Gymnophthalmidae) from the relictual forest mountains of the State of Ceará, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="15C4F15677DD95DE971C11A81290BE67" type="personal">
<mods:role id="1F2AB13F6E595010A465B6B8CB4E27FD">
<mods:roleTerm id="2F9A896C6BAAFCB39FF7736FC1518EC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9789DCB7B804C85698EA5B2FCDD5BE08">Borges-Nojosa, Diva Maria</mods:namePart>
</mods:name>
<mods:name id="9007911EA1AAFDC024F5B5457A11F1D2" type="personal">
<mods:role id="FE4BAB03946ADB8BC1A40C87112D25C5">
<mods:roleTerm id="41B3025D4464977F0B4E4A167D5C6899">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3443656DCA1F66DD67E94DF1F470561">Caramaschi, Ulisses</mods:namePart>
</mods:name>
<mods:name id="22D15FF7E9543580719A8F98CA39CDA5" type="personal">
<mods:role id="EB4CA8E924B3E05475EF376732AB51B4">
<mods:roleTerm id="ACF6D76864D495559869156568235ED4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F97214EBA7FCA812DBA4ADB4668EAD2E">Rodrigues, Miguel Trefaut</mods:namePart>
</mods:name>
<mods:typeOfResource id="82E9B128A43607AC5E2A16D26679419D">text</mods:typeOfResource>
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<mods:titleInfo id="71A2B88383115E84CB6F6090D662E2BA">
<mods:title id="DFAA3F6D5E781888840E87E7153D6E57">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D14483F3D0C319896C7C1B0E4423D238">
<mods:date id="95DAB1B2B443CF35F350D2D297F081F3">2016</mods:date>
<mods:detail id="7341C6FAFB28A196752D32A074EAFCE1" type="volume">
<mods:number id="05D642EB8D1F3AE258FD0E1507CCBBB5">4169</mods:number>
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<mods:detail id="8A7F16881387A456EC3556686E0F6418" type="issue">
<mods:number id="EB08A0A1424DAE73A52144C792069A80">1</mods:number>
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<mods:start id="8D6CE24184657E06D221AC0BA3078437">160</mods:start>
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<mods:classification id="ABCE5B2DEDA89954E12AA047AF2AADEF">journal article</mods:classification>
<mods:identifier id="3CD0FD7529390A2DB1CBF72798721321" type="DOI">10.11646/zootaxa.4169.1.8</mods:identifier>
<mods:identifier id="B5A446C5FFF9B9B00B6F989DEE73AE00" type="GBIF-Dataset">a78cff31-17bb-4b28-bf38-cd97d4feb6b1</mods:identifier>
<mods:identifier id="FA2B471B35C5441F2530E4399EC2B227" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="616FD780E39854AE054329A5B036F8FE" type="Zenodo-Dep">262091</mods:identifier>
<mods:identifier id="271512B6060694AEF2E1D4F853F238F0" type="ZooBank">B54A654B-D5FA-4707-9C31-BAFAF98D5F53</mods:identifier>
</mods:mods>
<treatment id="03AB93352A24FFF4C3FCF8C24A20599E" ID-DOI="http://doi.org/10.5281/zenodo.5680789" ID-GBIF-Taxon="124552019" ID-Zenodo-Dep="5680789" LSID="urn:lsid:plazi:treatment:03AB93352A24FFF4C3FCF8C24A20599E" httpUri="http://treatment.plazi.org/id/03AB93352A24FFF4C3FCF8C24A20599E" lastPageId="4" lastPageNumber="164" pageId="1" pageNumber="161">
<subSubSection id="C31871A82A24FFF1C3FCF8C24F165BF7" pageId="1" pageNumber="161" type="nomenclature">
<paragraph id="8BBD22232A24FFF1C3FCF8C24C0A5B08" blockId="1.[151,562,1877,1936]" box="[151,562,1877,1903]" pageId="1" pageNumber="161">
<heading id="D0F5954F2A24FFF1C3FCF8C24C0A5B08" bold="true" box="[151,562,1877,1903]" fontSize="11" level="1" pageId="1" pageNumber="161" reason="1">
<emphasis id="B976FE312A24FFF1C3FCF8C24C0A5B08" bold="true" box="[151,562,1877,1903]" pageId="1" pageNumber="161">
<taxonomicName id="4C0259A02A24FFF1C3FCF8C24FF55B08" box="[151,461,1877,1903]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="161" phylum="Chordata" rank="species" species="limaverdorum" status="sp. nov.">
<emphasis id="B976FE312A24FFF1C3FCF8C24FF55B08" bold="true" box="[151,461,1877,1903]" italics="true" pageId="1" pageNumber="161">Placosoma limaverdorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245434A2A24FFF1C2BFF8C24C0A5B08" box="[468,562,1877,1903]" pageId="1" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD22232A24FFF1C3FCF8E04F165BF7" blockId="1.[151,562,1877,1936]" box="[151,302,1911,1936]" pageId="1" pageNumber="161">
(
<figureCitation id="13393EA62A24FFF1C3F5F8E04F1D5BF7" box="[158,293,1911,1936]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="5.[151,250,1968,1990]" captionStartId-1="6.[151,250,1344,1366]" captionStartId-2="6.[151,250,1849,1871]" captionTargetBox-0="[349,1222,245,1876]" captionTargetBox-1="[208,1367,252,1272]" captionTargetBox-2="[151,1436,1416,1827]" captionTargetId-0="figure@5.[343,1226,195,1950]" captionTargetId-1="figure@6.[151,1435,193,1323]" captionTargetId-2="figure@6.[151,1436,1416,1827]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="FIGURE 1. Placosoma limaverdorum sp. nov., Holotype (CHUFC L 1998). (A) dorsal (B) lateral, (C) ventral views of the head. (Drawing by P. Maçaranduba)." captionText-1="FIGURE 2. Placosoma limaverdorum sp. nov., Holotype (CHUFC L 1998) — Cloacal región. (Drawing by P. Maçaranduba)." captionText-2="FIGURE 3. Color in life of two paratypes of Placosoma limaverdorum sp. nov. (A) CHUFC L 4170, adult male and (B) CHUFC L 4171, adult female. (Photos: D. M. Borges-Nojosa)." httpUri-0="https://zenodo.org/record/262092/files/figure.png" httpUri-1="https://zenodo.org/record/262093/files/figure.png" httpUri-2="https://zenodo.org/record/262094/files/figure.png" pageId="1" pageNumber="161">Figures 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31871A82A24FFF2C3FCF8574F645E0E" lastPageId="2" lastPageNumber="162" pageId="1" pageNumber="161" type="materials_examined">
<paragraph id="8BBD22232A24FFF2C3FCF8574FCC5CB0" blockId="1.[151,1436,1984,2009]" lastBlockId="2.[151,1437,151,2021]" lastPageId="2" lastPageNumber="162" pageId="1" pageNumber="161">
<bibRef id="52A2B6D52A24FFF2C3FCF8574FCC5CB0" lastPageId="2" lastPageNumber="162" pageId="1" pageNumber="161" type="journal article">
<materialsCitation id="3B6A287E2A24FFF2C3FCF8574FCC5CB0" ID-GBIF-Occurrence="1313779560" collectionCode="CHUFC" collectorName="Macico de Baturite &amp; Sitio Olho d'Agua dos Tangaras &amp; Datum &amp; J. A. Carneiro &amp; December &amp; Figures" country="Brazil" lastPageId="2" lastPageNumber="162" latitude="-4.235947" location="Macico de Baturite" longLatPrecision="1" longitude="-38.91699" municipality="Pacoti Municipality" pageId="1" pageNumber="161" specimenCount="1" stateProvince="Ceara" typeStatus="holotype">
<bibRefCitation id="EF935FD22A24FFF1C3FCF8574FFD5BBF" author="Holotype" box="[151,453,1984,2009]" pageId="1" pageNumber="161" refString="Holotype. CHUFC L 1998, an adult male, collected at the Macico de Baturite (Sitio Olho d'Agua dos Tangaras), Pacoti Municipality, state of Ceara, Brazil (04 o 14 ' 9,41 &quot; S / 38 o 55 ' 1,15 &quot; W; Datum WGS 84) by J. A. Carneiro in December, 1996 (Figures 1 - 2)." type="journal article" year="1998">
<author id="53BFCAC62A24FFF1C3FCF8574FB35BBF" box="[151,395,1984,2009]" pageId="1" pageNumber="161">
<emphasis id="B976FE312A24FFF1C3FCF8574F335BBE" bold="true" box="[151,267,1984,2009]" pageId="1" pageNumber="161">
<typeStatus id="54B99C812A24FFF1C3FCF8574F3E5BBE" box="[151,262,1984,2009]" pageId="1" pageNumber="161" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED13BAE62A24FFF1C278F8574F485BBF" box="[275,368,1984,2008]" pageId="1" pageNumber="161">CHUFC</collectionCode>
L
</author>
<year id="FFAA63702A24FFF1C2E7F8574FFD5BBF" box="[396,453,1984,2008]" pageId="1" pageNumber="161">1998</year>
</bibRefCitation>
,
<title id="F906C9152A24FFF2C2B9F8564B345CD4" jopScore="0.87216794" lastPageId="2" lastPageNumber="162" pageId="1" pageNumber="161" type="_" typeScore="80">
an adult male, collected at the
<location id="8EDD74F82A24FFF1C05AF8574A335BBE" LSID="urn:lsid:plazi:treatment:03AB93352A24FFF4C3FCF8C24A20599E:8EDD74F82A24FFF1C05AF8574A335BBE" box="[817,1035,1984,2009]" country="Brazil" latitude="-4.235947" longLatPrecision="1" longitude="-38.91699" municipality="Pacoti Municipality" name="Macico de Baturite" pageId="1" pageNumber="161" stateProvince="Ceara">
<collectorName id="26F747F52A24FFF1C05AF8574A335BBE" box="[817,1035,1984,2009]" pageId="1" pageNumber="161">Maciço de Baturité</collectorName>
</location>
(
<collectorName id="26F747F52A24FFF1C777F8574BA85BBF" box="[1052,1424,1984,2009]" pageId="1" pageNumber="161">Sítio Olho d’Água dos Tangarás</collectorName>
),
<collectingMunicipality id="6BD9B8592A27FFF2C3FCFF0D4F415CD4" box="[151,377,154,179]" pageId="2" pageNumber="162">Pacoti Municipality</collectingMunicipality>
, state of
<collectingRegion id="49C6ECC12A27FFF2C28DFF0C4C125CD4" box="[486,554,155,179]" country="Brazil" name="Ceara" pageId="2" pageNumber="162">Ceará</collectingRegion>
,
<collectingCountry id="F31562B32A27FFF2C153FF0D4C445CD4" box="[568,636,154,179]" name="Brazil" pageId="2" pageNumber="162">Brazil</collectingCountry>
(
<geoCoordinate id="EE3644E42A27FFF2C1E4FF0C4D1E5CD4" box="[655,806,151,179]" direction="south" orientation="latitude" pageId="2" pageNumber="162" precision="1" value="-4.235947">04o14’9,41”S</geoCoordinate>
/
<geoCoordinate id="EE3644E42A27FFF2C029FF0C4DD85CD4" box="[834,992,151,179]" direction="west" orientation="longitude" pageId="2" pageNumber="162" precision="1" value="-38.91699">38o55'1,15&quot;W</geoCoordinate>
;
<collectorName id="26F747F52A27FFF2C085FF0C4A035CD5" box="[1006,1083,155,178]" pageId="2" pageNumber="162">Datum</collectorName>
WGS 84) by
<collectorName id="26F747F52A27FFF2C788FF0C4B345CD4" box="[1251,1292,155,179]" pageId="2" pageNumber="162">J.A</collectorName>
</title>
.
<journalOrPublisher id="51C328EF2A27FFF2C671FF0D4F345CB0" jopScore="0.89074683" pageId="2" pageNumber="162">
Carneiro in
<collectorName id="26F747F52A27FFF2C3FCFF294F345CB0" box="[151,268,190,215]" pageId="2" pageNumber="162">December</collectorName>
</journalOrPublisher>
,
<part id="FFA688BE2A27FFF2C27CFF294F775CB0" box="[279,335,190,215]" pageId="2" pageNumber="162" type="volume">1996</part>
(
<figureCitation id="13393EA62A27FFF2C236FF294FDF5CB0" box="[349,487,190,215]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="5.[151,250,1968,1990]" captionStartId-1="6.[151,250,1344,1366]" captionTargetBox-0="[349,1222,245,1876]" captionTargetBox-1="[208,1367,252,1272]" captionTargetId-0="figure@5.[343,1226,195,1950]" captionTargetId-1="figure@6.[151,1435,193,1323]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 1. Placosoma limaverdorum sp. nov., Holotype (CHUFC L 1998). (A) dorsal (B) lateral, (C) ventral views of the head. (Drawing by P. Maçaranduba)." captionText-1="FIGURE 2. Placosoma limaverdorum sp. nov., Holotype (CHUFC L 1998) — Cloacal región. (Drawing by P. Maçaranduba)." httpUri-0="https://zenodo.org/record/262092/files/figure.png" httpUri-1="https://zenodo.org/record/262093/files/figure.png" pageId="2" pageNumber="162">
<collectorName id="26F747F52A27FFF2C236FF294F8B5CB0" box="[349,435,190,215]" pageId="2" pageNumber="162">Figures</collectorName>
<pagination id="B2CBDCD72A27FFF2C2D1FF284FDF5CB0" box="[442,487,191,215]" pageId="2" pageNumber="162" type="pagination">1–2</pagination>
</figureCitation>
).
</materialsCitation>
</bibRef>
</paragraph>
<paragraph id="8BBD22232A27FFF2C3ACFF754F645E0E" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="162">
<emphasis id="B976FE312A27FFF2C3ACFF754F075C9C" bold="true" box="[199,319,226,251]" pageId="2" pageNumber="162">Paratypes</emphasis>
. CHUFC L1958, juvenile male,
<date id="FFBC04E32A27FFF2C1D3FF754DA65C9C" box="[696,926,226,251]" pageId="2" pageNumber="162" value="1995-11-15">November 15, 1995</date>
, and MZUSP 88071 (CHUFC L2055), adult male,
<date id="FFBC04E32A27FFF2C38BFE914F825D78" box="[224,442,262,287]" pageId="2" pageNumber="162" value="1998-02-19">February 19, 1998</date>
, collected by D.M. Borges-Nojosa; CHUFC L2527, adult male, CHUFC L2525 and CHUFC L2526, adult females,
<date id="FFBC04E32A27FFF2C29CFEBC4C8C5D24" box="[503,692,299,323]" pageId="2" pageNumber="162" value="1998-08-27">August 27, 1998</date>
, collected by D.C. Lima; CHUFC L3116, adult female,
<date id="FFBC04E32A27FFF2C642FEBD4EC75D00" pageId="2" pageNumber="162" value="2003-12-12">December 12, 2003</date>
, collected by Y. Quinet; CHUFC L4170 (
<figureCitation id="13393EA62A27FFF2C18CFED94D745D00" box="[743,844,334,359]" captionStart="FIGURE 3" captionStartId="6.[151,250,1849,1871]" captionTargetBox="[151,1436,1416,1827]" captionTargetId="figure@6.[151,1436,1416,1827]" captionTargetPageId="6" captionText="FIGURE 3. Color in life of two paratypes of Placosoma limaverdorum sp. nov. (A) CHUFC L 4170, adult male and (B) CHUFC L 4171, adult female. (Photos: D. M. Borges-Nojosa)." httpUri="https://zenodo.org/record/262094/files/figure.png" pageId="2" pageNumber="162">Figure 3</figureCitation>
), adult male, and CHUFC L4171 (
<figureCitation id="13393EA62A27FFF2C78DFED94B735D00" box="[1254,1355,334,359]" captionStart="FIGURE 3" captionStartId="6.[151,250,1849,1871]" captionTargetBox="[151,1436,1416,1827]" captionTargetId="figure@6.[151,1436,1416,1827]" captionTargetPageId="6" captionText="FIGURE 3. Color in life of two paratypes of Placosoma limaverdorum sp. nov. (A) CHUFC L 4170, adult male and (B) CHUFC L 4171, adult female. (Photos: D. M. Borges-Nojosa)." httpUri="https://zenodo.org/record/262094/files/figure.png" pageId="2" pageNumber="162">Figure 3</figureCitation>
), adult female,
<date id="FFBC04E32A27FFF2C39EFEE44FA35DEC" box="[245,411,370,395]" pageId="2" pageNumber="162" value="2010-06-26">June 26, 2010</date>
, collected by D.C. Lima and T.A. Sousa; CHUFC L4194, adult male,
<date id="FFBC04E32A27FFF2C7BEFEE44BA05DEC" box="[1237,1432,371,395]" pageId="2" pageNumber="162" value="2010-08-22">August 22, 2010</date>
, collected by D.C. Lima; MNRJ 26049 (CHUFC L5649), adult male,
<date id="FFBC04E32A27FFF2C0C9FE014ABF5DC8" box="[930,1159,406,431]" pageId="2" pageNumber="162" value="2014-09-27">September 27, 2014</date>
,
<materialsCitation id="3B6A287E2A27FFF2C7F8FE004E875E79" ID-GBIF-Occurrence="1313779559" collectingDate="2014-10-18" collectionCode="MPEG, CHUFC" collectorName="D. M. Borges-Nojosa &amp; D. P. Castro &amp; Serra de Maranguape" country="Brazil" county="State" latitude="-4.05" location="Serra de Maranguape" longLatPrecision="1307" longitude="-38.666668" municipality="Maranguape Municipality" pageId="2" pageNumber="162" specimenCode="MPEG 32210" specimenCount="1" stateProvince="Ceara">
<specimenCode id="DBA48A582A27FFF2C7F8FE004B165DC8" box="[1171,1326,407,431]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="162">
<collectionCode id="ED13BAE62A27FFF2C7F8FE004AD95DC8" box="[1171,1249,407,431]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="162">MPEG</collectionCode>
32210
</specimenCode>
(
<collectionCode id="ED13BAE62A27FFF2C656FE004BA45DC8" box="[1341,1436,407,431]" pageId="2" pageNumber="162">CHUFC</collectionCode>
L5783), juvenile male,
<collectingDate id="EFF8FD0B2A27FFF2C2C9FE2D4C545DB4" box="[418,620,442,467]" pageId="2" pageNumber="162" value="2014-10-18">
<date id="FFBC04E32A27FFF2C2C9FE2D4C545DB4" box="[418,620,442,467]" pageId="2" pageNumber="162" value="2014-10-18">October 18, 2014</date>
</collectingDate>
, collected by
<collectorName id="26F747F52A27FFF2C067FE2C4DC25DB4" box="[780,1018,442,467]" pageId="2" pageNumber="162">D.M. Borges-Nojosa</collectorName>
,
<collectorName id="26F747F52A27FFF2C76CFE2C4AB05DB4" box="[1031,1160,443,467]" pageId="2" pageNumber="162">D.P. Castro</collectorName>
et al.: all from
<collectorName id="26F747F52A27FFF2C655FE2C4F1E5D9D" pageId="2" pageNumber="162">
<location id="8EDD74F82A27FFF2C655FE2C4F1E5D9D" LSID="urn:lsid:plazi:treatment:03AB93352A24FFF4C3FCF8C24A20599E:8EDD74F82A27FFF2C655FE2C4F1E5D9D" country="Brazil" county="State" latitude="-4.05" longLatPrecision="1307" longitude="-38.666668" municipality="Maranguape Municipality" name="Serra de Maranguape" pageId="2" pageNumber="162" stateProvince="Ceara">Serra de Maranguape</location>
</collectorName>
,
<collectingMunicipality id="6BD9B8592A27FFF2C258FE754C625D9D" box="[307,602,481,506]" pageId="2" pageNumber="162">Maranguape Municipality</collectingMunicipality>
,
<collectingCounty id="62DC5AAF2A27FFF2C103FE754CA75D9D" box="[616,671,482,506]" pageId="2" pageNumber="162">State</collectingCounty>
of
<collectingRegion id="49C6ECC12A27FFF2C1A2FE754D355D9D" box="[713,781,482,506]" country="Brazil" name="Ceara" pageId="2" pageNumber="162">Ceará</collectingRegion>
,
<collectingCountry id="F31562B32A27FFF2C070FE764D675D9D" box="[795,863,481,506]" name="Brazil" pageId="2" pageNumber="162">Brazil</collectingCountry>
(03o54'–
<geoCoordinate id="EE3644E42A27FFF2C0AFFE764A1B5D9D" box="[964,1059,478,506]" direction="south" orientation="latitude" pageId="2" pageNumber="162" precision="925" value="-4.05">04o03' S</geoCoordinate>
/ 38o32'–
<geoCoordinate id="EE3644E42A27FFF2C7E4FE754AC25D9E" box="[1167,1274,478,506]" direction="west" orientation="longitude" pageId="2" pageNumber="162" precision="925" value="-38.666668">38o40' W</geoCoordinate>
; Datum WGS 84)
</materialsCitation>
.
<materialsCitation id="3B6A287E2A27FFF2C3BEFD914F645E0E" ID-GBIF-Occurrence="1313779561" collectingDate="2005-03" collectingDateMax="2005-07-31" collectingDateMin="2005-03" collectionCode="CHUFC" collectorName="L. B. M. Brito &amp; Macico de Baturite" country="Brazil" county="State" latitude="-4.6666665" location="Macico de Baturite" longLatPrecision="1306" longitude="-39.166668" municipality="Guaramiranga Municipality" pageId="2" pageNumber="162" specimenCount="1" stateProvince="Ceara">
<collectionCode id="ED13BAE62A27FFF2C3BEFD914F0A5E79" box="[213,306,518,542]" pageId="2" pageNumber="162">CHUFC</collectionCode>
L3720, juvenile male,
<collectingDate id="EFF8FD0B2A27FFF2C108FD924D0E5E79" box="[611,822,517,542]" pageId="2" pageNumber="162" value="2005-03" valueMax="2005-07-31" valueMin="2005-03-00">
<date id="FFBC04E32A27FFF2C108FD924D0E5E79" box="[611,822,517,542]" pageId="2" pageNumber="162" value="2005-03" valueMax="2005-07-31" valueMin="2005-03-00">March–July 2005</date>
</collectingDate>
, collected by
<collectorName id="26F747F52A27FFF2C09AFD914AAB5E79" box="[1009,1171,517,542]" pageId="2" pageNumber="162">L.B.M. Brito</collectorName>
:
<collectorName id="26F747F52A27FFF2C7C3FD924BA05E79" box="[1192,1432,517,542]" pageId="2" pageNumber="162">
<location id="8EDD74F82A27FFF2C7C3FD924BA05E79" LSID="urn:lsid:plazi:treatment:03AB93352A24FFF4C3FCF8C24A20599E:8EDD74F82A27FFF2C7C3FD924BA05E79" box="[1192,1432,517,542]" country="Brazil" county="State" latitude="-4.6666665" longLatPrecision="1306" longitude="-39.166668" municipality="Guaramiranga Municipality" name="Macico de Baturite" pageId="2" pageNumber="162" stateProvince="Ceara">Maciço de Baturité</location>
</collectorName>
,
<collectingMunicipality id="6BD9B8592A27FFF2C3FCFDBB4FED5E22" box="[151,469,556,581]" pageId="2" pageNumber="162">Guaramiranga Municipality</collectingMunicipality>
(Sítio Guaramiranga),
<collectingCounty id="62DC5AAF2A27FFF2C181FDBA4D1A5E22" box="[746,802,557,581]" pageId="2" pageNumber="162">State</collectingCounty>
of
<collectingRegion id="49C6ECC12A27FFF2C039FDBA4DAE5E22" box="[850,918,557,581]" country="Brazil" name="Ceara" pageId="2" pageNumber="162">Ceará</collectingRegion>
,
<collectingCountry id="F31562B32A27FFF2C0C3FDBB4DD45E22" box="[936,1004,556,581]" name="Brazil" pageId="2" pageNumber="162">Brazil</collectingCountry>
(04o05–
<geoCoordinate id="EE3644E42A27FFF2C724FDBB4A8C5E22" box="[1103,1204,553,581]" direction="south" orientation="latitude" pageId="2" pageNumber="162" precision="925" value="-4.6666665">04o40’ S</geoCoordinate>
/ 38o30'–
<geoCoordinate id="EE3644E42A27FFF2C643FDBA4BA05E23" box="[1320,1432,553,581]" direction="west" orientation="longitude" pageId="2" pageNumber="162" precision="925" value="-39.166668">39o10' W</geoCoordinate>
; Datum WGS 84).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31871A82A27FFF2C3ACFDE34FFC5F7A" pageId="2" pageNumber="162" type="etymology">
<paragraph id="8BBD22232A27FFF2C3ACFDE34FFC5F7A" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="162">
<emphasis id="B976FE312A27FFF2C3ACFDE34F765EEA" bold="true" box="[199,334,628,653]" pageId="2" pageNumber="162">Etymology.</emphasis>
The species honors Prof. Dr. José Santiago Lima-Verde, an important collaborator of herpetological studies in the State of
<collectingRegion id="49C6ECC12A27FFF2C15CFD0E4C435ED6" box="[567,635,665,689]" country="Brazil" name="Ceara" pageId="2" pageNumber="162">Ceará</collectingRegion>
, who started the Herpetological Collection of the Universidade Federal do
<collectingRegion id="49C6ECC12A27FFF2C3D5FD2A4EC75EB2" box="[190,255,701,725]" country="Brazil" name="Ceara" pageId="2" pageNumber="162">Ceará</collectingRegion>
(CHUFC) and the Núcleo Regional de Ofiologia da UFC (NUROF-UFC). This description is also a tribute to the agronomist Wilson Luiz Lima-Verde, owner of the cottage (Sítio Olho d’Água dos Tangarás) where the
<typeStatus id="54B99C812A27FFF2C3ABFC934F1A5F7A" box="[192,290,772,797]" pageId="2" pageNumber="162" type="holotype">holotype</typeStatus>
was obtained.
</paragraph>
</subSubSection>
<subSubSection id="C31871A82A27FFF3C3ACFCBF4C785E07" lastPageId="3" lastPageNumber="163" pageId="2" pageNumber="162" type="diagnosis">
<paragraph id="8BBD22232A27FFF2C3ACFCBF4FAE5807" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="162">
<emphasis id="B976FE312A27FFF2C3ACFCBF4F795F26" bold="true" box="[199,321,808,833]" pageId="2" pageNumber="162">Diagnosis.</emphasis>
Body long, depressed; head long with a pronounced acute snout; tail longer than body; tongue elongate, arrow headed. Limbs slender, pentadactyl, all clawed. Ear opening and eyelid distinct, with an undivided transparent palpebral disk. Frontonasal single; prefrontals, frontoparietals, parietals, interparietal and two series of occipitals present; four supraoculars. Nostril in the center of a divided nasal. Tympanic recess deep. Collar absent, gular fold present; four enlarged pairs of chinshields, posteriormost reduced. Dorsal scales smooth, subimbricate, juxtaposed, disposed in regular transverse rows. Lateral scales distinct in shape and much smaller than dorsals and ventrals, disposed in irregular transverse rows. Ventrals quadrangular, smooth, imbricate, larger than dorsals, disposed in regular transverse and longitudinal rows. Males with femoral pores, preanal pores absent; females without femoral pores.
</paragraph>
<paragraph id="8BBD22232A27FFF2C3ACFBFB4B015B32" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="162">
The systematics of gymnophthalmid lizards is going through a period of effervescence since the advent of the first comprehensive molecular study showed that morphologic character convergence was widespread, introducing uncertainty in their taxonomy (
<bibRefCitation id="EF935FD22A27FFF2C297FB234CC358AA" author="Pellegrino" box="[508,763,1204,1229]" pageId="2" pageNumber="162" refString="Pellegrino, K. C. M., Rodrigues, M. T., Yonenaga-Yassuda, Y. &amp; Sites, J. W. Jr. (2001) A molecular perspective on the evolution of microteiid lizards (Squamata, Gymnophthalmidae), and a new classification for the family. Biological Journal of the Linnean Society, 74 (3), 315 - 338. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2001. tb 01395. x" type="journal article" year="2001">
Pellegrino
<emphasis id="B976FE312A27FFF2C111FB224C8D58AA" box="[634,693,1204,1229]" italics="true" pageId="2" pageNumber="162">et al.</emphasis>
2001
</bibRefCitation>
). Since then, several genera and suprageneric groups have been defined molecularly but we have no consistent morphological synapomorphies supporting either the present admitted genera and tribes (
<bibRefCitation id="EF935FD22A27FFF2C2A4FB6B4CFE5972" author="Rodrigues" box="[463,710,1276,1301]" pageId="2" pageNumber="162" refString="Rodrigues, M. T., Cassimiro, J., Pavan, D., Curcio, F. F., Verdade, V. K. &amp; Pellegrino, K. C. M. (2009) A new genus of Microteiid lizard from the Caparao Mountains, southeastern Brazil, with a discussion of relationships among Gymnophthalminae (Squamata). American Museum Novitates, 3673, 1 - 27. http: // dx. doi. org / 10.1206 / 622.1" type="journal article" year="2009">
Rodrigues
<emphasis id="B976FE312A27FFF2C120FB6A4CBC5972" box="[587,644,1276,1301]" italics="true" pageId="2" pageNumber="162">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EF935FD22A27FFF2C1BAFB6B4A395972" author="Torres-Carvajal" box="[721,1025,1276,1301]" pageId="2" pageNumber="162" refString="Torres-Carvajal, O., Lobos, S. E., Venegas, P. J., Chavez, G., Aguirre-Penafiel, V., Zurita, D. &amp; Echevarria, L. Y. (2016) Phylogeny and biogeography of the most diverse clade of South American gymnophthalmid lizards (Squamata, Gymnophthalmidae, Cercosaurinae). Molecular Phylogenetics and Evolution, 99, 63 - 75. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.006" type="journal article" year="2016">
Torres-Carvajal
<emphasis id="B976FE312A27FFF2C0ECFB6A4DF85972" box="[903,960,1276,1301]" italics="true" pageId="2" pageNumber="162">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EF935FD22A27FFF2C766FB6B4B2D5972" author="Goicoechea" box="[1037,1301,1276,1301]" pageId="2" pageNumber="162" refString="Goicoechea, N., Frost, D. R., De la Riva, I., Pellegrino, K. C. M., Sites, J., Rodrigues, M. T. &amp; Padial, J. M. (2016) Molecular systematics of teioid lizards (Teioidea / Gymnophthalmoidea: Squamata) based on the analysis of 48 loci under treealignment and similarity-alignment. Cladistics, 2016, 1 - 48. http: // dx. doi. org / 10.1111 / cla. 12150" type="journal article" year="2016">
Goicoechea
<emphasis id="B976FE312A27FFF2C7F2FB6A4AE95972" box="[1177,1233,1276,1301]" italics="true" pageId="2" pageNumber="162">et al.</emphasis>
2016
</bibRefCitation>
). The genus
<taxonomicName id="4C0259A02A27FFF2C3FCFAB74F2C595E" box="[151,276,1312,1337]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C3FCFAB74F2C595E" box="[151,276,1312,1337]" italics="true" pageId="2" pageNumber="162">Placosoma</emphasis>
</taxonomicName>
has been consistently recovered in molecular studies, nested among the Cercosaurini with high support (
<bibRefCitation id="EF935FD22A27FFF2C3F5FAD34F93593A" author="Pellegrino" box="[158,427,1348,1373]" pageId="2" pageNumber="162" refString="Pellegrino, K. C. M., Rodrigues, M. T., Yonenaga-Yassuda, Y. &amp; Sites, J. W. Jr. (2001) A molecular perspective on the evolution of microteiid lizards (Squamata, Gymnophthalmidae), and a new classification for the family. Biological Journal of the Linnean Society, 74 (3), 315 - 338. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2001. tb 01395. x" type="journal article" year="2001">
Pellegrino
<emphasis id="B976FE312A27FFF2C249FAD24F61593A" box="[290,345,1348,1373]" italics="true" pageId="2" pageNumber="162">et al</emphasis>
. 2001
</bibRefCitation>
,
<bibRefCitation id="EF935FD22A27FFF2C2D6FAD34C0F593A" author="Kok" box="[445,567,1348,1373]" pageId="2" pageNumber="162" refString="Kok, P. J. R. (2015) A new species of the Pantepui endemic genus Riolama (Squamata: Gymnophthalmidae) from the summit of Murisipan-tepui, with the erection of a new gymnophthalmid subfamily. Zoological Journal of the Linnean Society, 174 (3), 500 - 518. http: // dx. doi. org / 10.1111 / zoj. 12241" type="journal article" year="2015">Kok 2015</bibRefCitation>
,
<bibRefCitation id="EF935FD22A27FFF2C122FAD34DA8593A" author="Torres-Carvajal" box="[585,912,1348,1373]" pageId="2" pageNumber="162" refString="Torres-Carvajal, O., Lobos, S. E., Venegas, P. J., Chavez, G., Aguirre-Penafiel, V., Zurita, D. &amp; Echevarria, L. Y. (2016) Phylogeny and biogeography of the most diverse clade of South American gymnophthalmid lizards (Squamata, Gymnophthalmidae, Cercosaurinae). Molecular Phylogenetics and Evolution, 99, 63 - 75. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.006" type="journal article" year="2016">
Torres-Carvajal
<emphasis id="B976FE312A27FFF2C06CFAD24D7F593A" box="[775,839,1348,1373]" italics="true" pageId="2" pageNumber="162">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EF935FD22A27FFF2C0C8FAD34AF8593A" author="Goicoechea" box="[931,1216,1348,1373]" pageId="2" pageNumber="162" refString="Goicoechea, N., Frost, D. R., De la Riva, I., Pellegrino, K. C. M., Sites, J., Rodrigues, M. T. &amp; Padial, J. M. (2016) Molecular systematics of teioid lizards (Teioidea / Gymnophthalmoidea: Squamata) based on the analysis of 48 loci under treealignment and similarity-alignment. Cladistics, 2016, 1 - 48. http: // dx. doi. org / 10.1111 / cla. 12150" type="journal article" year="2016">
Goicoechea
<emphasis id="B976FE312A27FFF2C75EFAD24A56593A" box="[1077,1134,1348,1373]" italics="true" pageId="2" pageNumber="162">et al</emphasis>
. 2016
</bibRefCitation>
). Nevertheless, no morphological synapomorphies have been proposed either to the genus or to the tribe Cercosaurini. Considering the absence of molecular data for the species herein described and while waiting for a sound morphological study of the entire tribe to better characterize its genera we use the following character combination to support the inclusion of the new species in the Cercosaurini and in the genus
<taxonomicName id="4C0259A02A27FFF2C0C5FA434A12598A" box="[942,1066,1492,1517]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C0C5FA434A12598A" box="[942,1066,1492,1517]" italics="true" pageId="2" pageNumber="162">Placosoma</emphasis>
</taxonomicName>
: presence of an eyelid and ear opening, a long body and tail, pentadactyl limbs, presence of an elongate and acute snout, two series of occipital scales, homogeneous and slightly imbricate quadrangular dorsal and ventral scales, lateral scales distinct in size and shape from dorsal and ventral ones but never granular. The genus
<taxonomicName id="4C0259A02A27FFF2C0DFF9D74A095A3E" box="[948,1073,1600,1625]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C0DFF9D74A095A3E" box="[948,1073,1600,1625]" italics="true" pageId="2" pageNumber="162">Placosoma</emphasis>
</taxonomicName>
differs from the stream adapted genera
<taxonomicName id="4C0259A02A27FFF2C39BF9F34F475A1A" box="[240,383,1636,1661]" class="Reptilia" family="Gymnophthalmidae" genus="Echinosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C39BF9F34F475A1A" box="[240,383,1636,1661]" italics="true" pageId="2" pageNumber="162">Echinosaura</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0259A02A27FFF2C2FEF9F24C235A1B" box="[405,539,1637,1660]" class="Reptilia" family="Gymnophthalmidae" genus="Neusticurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C2FEF9F24C235A1B" box="[405,539,1637,1660]" italics="true" pageId="2" pageNumber="162">Neusticurus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0259A02A27FFF2C15AF9F24C995A1B" box="[561,673,1637,1660]" class="Reptilia" family="Gymnophthalmidae" genus="Potamites" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C15AF9F24C995A1B" box="[561,673,1637,1660]" italics="true" pageId="2" pageNumber="162">Potamites</emphasis>
</taxonomicName>
, and
<emphasis id="B976FE312A27FFF2C185F9F34DB35A1A" box="[750,907,1636,1661]" italics="true" pageId="2" pageNumber="162">Gelanosaurus</emphasis>
by having a homogeneous dorsal scalation (heterogeneous in size and shape and ornamentation in those genera). From
<taxonomicName id="4C0259A02A27FFF2C775F91E4A995AC7" box="[1054,1185,1673,1696]" class="Reptilia" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C775F91E4A995AC7" box="[1054,1185,1673,1696]" italics="true" pageId="2" pageNumber="162">Cercosaura</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0259A02A27FFF2C78BF91F4BB85AC6" box="[1248,1408,1672,1697]" class="Squamata" family="Gymnophthalmidae" genus="Pantodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C78BF91F4BB85AC6" box="[1248,1408,1672,1697]" italics="true" pageId="2" pageNumber="162">Pantodactylus</emphasis>
</taxonomicName>
it differs by presenting eight longitudinal series of quadrangular scales at midbody and no granular scales in the flanks (six or less enlarged rows of ventrals and granular scales on flanks). From
<taxonomicName id="4C0259A02A27FFF2C7F5F9474AD75A8E" box="[1182,1263,1744,1769]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C7F5F9474AD75A8E" box="[1182,1263,1744,1769]" italics="true" pageId="2" pageNumber="162">Anadia</emphasis>
</taxonomicName>
,
<emphasis id="B976FE312A27FFF2C66DF9474F705B6A" italics="true" pageId="2" pageNumber="162">
<taxonomicName id="4C0259A02A27FFF2C66DF9474BA05A8E" box="[1286,1432,1744,1769]" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">Euspondylus</taxonomicName>
,
<taxonomicName id="4C0259A02A27FFF2C3FCF9634F7C5B6A" box="[151,324,1780,1805]" class="Reptilia" family="Gymnophthalmidae" genus="Macropholidus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">Macropholidus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C0259A02A27FFF2C2EDF9624FF75B6B" box="[390,463,1781,1804]" class="Reptilia" family="Gymnophthalmidae" genus="Riama" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C2EDF9624FF75B6B" box="[390,463,1781,1804]" italics="true" pageId="2" pageNumber="162">Riama</emphasis>
</taxonomicName>
it can be distinguished by having a series of lateral scales smaller than dorsal and ventral scales but never granular (generally identical to dorsals, without distinction). From
<taxonomicName id="4C0259A02A27FFF2C7FFF88E4B275B57" box="[1172,1311,1817,1840]" class="Reptilia" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C7FFF88E4B275B57" box="[1172,1311,1817,1840]" italics="true" pageId="2" pageNumber="162">Proctoporus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0259A02A27FFF2C646F88F4BA45B56" box="[1325,1436,1816,1841]" class="Reptilia" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C646F88F4BA45B56" box="[1325,1436,1816,1841]" italics="true" pageId="2" pageNumber="162">Petracola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0259A02A27FFF2C3ADF8AB4F635B32" box="[198,347,1852,1877]" class="Reptilia" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="genus">
<emphasis id="B976FE312A27FFF2C3ADF8AB4F635B32" box="[198,347,1852,1877]" italics="true" pageId="2" pageNumber="162">Pholidobolus</emphasis>
</taxonomicName>
it differs by the absence of a lateral fold with granular scales on sides of body (present).
</paragraph>
<paragraph id="8BBD22232A27FFF3C3ACF8F74C785E07" blockId="2.[151,1437,151,2021]" lastBlockId="3.[151,1437,151,2018]" lastPageId="3" lastPageNumber="163" pageId="2" pageNumber="162">
<taxonomicName id="4C0259A02A27FFF2C3ACF8F74FD75B1E" box="[199,495,1888,1913]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="162" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A27FFF2C3ACF8F74FD75B1E" box="[199,495,1888,1913]" italics="true" pageId="2" pageNumber="162">Placosoma limaverdorum</emphasis>
</taxonomicName>
differs from all congeners by having an undivided transparent palpebral disc (divided with scales in all other species), an entirely divided nasal with a nostril in the center, a deeply tympanic recess, absence of preanal pores, and by the presence of a light vertebral stripe extending from frontonasal to tip of tail, with about one scale wide, bordered laterally by a dark pigmentation, irregularly interrupted by interspaced ocelli (two paravertebral light stripes in
<emphasis id="B976FE312A26FFF3C10AFF0E4DF75CD7" box="[609,975,151,176]" italics="true" pageId="3" pageNumber="163">
<taxonomicName id="4C0259A02A26FFF3C10AFF0E4CC65CD7" box="[609,766,151,176]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cordylinum">P. cordylinum</taxonomicName>
,
<taxonomicName id="4C0259A02A26FFF3C067FF0E4DF35CD7" box="[780,971,151,176]" class="Squamata" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="champsonotus">P. champsonotus</taxonomicName>
,
</emphasis>
and
<emphasis id="B976FE312A26FFF3C762FF0E4AAA5CD7" box="[1033,1170,151,176]" italics="true" pageId="3" pageNumber="163">P. g l ab e l l um</emphasis>
, and two paravertebral stripes and a lateral one in
<taxonomicName id="4C0259A02A26FFF3C2B6FF2A4C665CB3" box="[477,606,189,212]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cipoense">
<emphasis id="B976FE312A26FFF3C2B6FF2A4C665CB3" box="[477,606,189,212]" italics="true" pageId="3" pageNumber="163">P. cipoense</emphasis>
</taxonomicName>
). It further differs from
<taxonomicName id="4C0259A02A26FFF3C0EFFF2A4A2C5CB2" box="[900,1044,188,213]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="glabellum">
<emphasis id="B976FE312A26FFF3C0EFFF2A4A2C5CB2" box="[900,1044,188,213]" italics="true" pageId="3" pageNumber="163">P. glabellum</emphasis>
</taxonomicName>
by having squared ventral scales (posterior part of scales rounded), and from
<taxonomicName id="4C0259A02A26FFF3C1C3FF764D515C9F" box="[680,873,223,248]" class="Squamata" family="Gymnophthalmidae" genus="Prionodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="champsonotus">
<emphasis id="B976FE312A26FFF3C1C3FF764D515C9F" box="[680,873,223,248]" italics="true" pageId="3" pageNumber="163">P. champsonotus</emphasis>
</taxonomicName>
by having smooth dorsal scales (keeled) and a maximum of six enlarged dorsal scales between fore limbs (eight or more).
<taxonomicName id="4C0259A02A26FFF3C08EFE934B3D5D7A" box="[997,1285,260,285]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A26FFF3C08EFE934B3D5D7A" box="[997,1285,260,285]" italics="true" pageId="3" pageNumber="163">Placosoma limaverdorum</emphasis>
</taxonomicName>
share with
<taxonomicName id="4C0259A02A26FFF3C6ECFE924F3F5D27" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="glabellum">
<emphasis id="B976FE312A26FFF3C6ECFE924F3F5D27" italics="true" pageId="3" pageNumber="163">P. glabellum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0259A02A26FFF3C27EFEBE4F975D27" box="[277,431,295,320]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cordylinum">
<emphasis id="B976FE312A26FFF3C27EFEBE4F975D27" box="[277,431,295,320]" italics="true" pageId="3" pageNumber="163">P. cordylinum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0259A02A26FFF3C28CFEBE4C5D5D27" box="[487,613,297,320]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cipoense">
<emphasis id="B976FE312A26FFF3C28CFEBE4C5D5D27" box="[487,613,297,320]" italics="true" pageId="3" pageNumber="163">P. cipoense</emphasis>
</taxonomicName>
a low number of enlarged dorsal scales between forelimbs (maximum of six) but differ from those species in number of femoral pores 21–26 (
<quantity id="4CFA8FC62A26FFF3C0FCFEDB4DC35D02" box="[919,1019,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="3" pageNumber="163" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
<taxonomicName id="4C0259A02A26FFF3C76AFEDA4AB45D02" box="[1025,1164,332,357]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="glabellum">
<emphasis id="B976FE312A26FFF3C76AFEDA4AB45D02" box="[1025,1164,332,357]" italics="true" pageId="3" pageNumber="163">P. glabellum</emphasis>
</taxonomicName>
,
<quantity id="4CFA8FC62A26FFF3C7F1FEDB4AC45D02" box="[1178,1276,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="8.128" metricValueMin="6.858" pageId="3" pageNumber="163" unit="in" value="29.5" valueMax="32.0" valueMin="27.0">27–32 in</quantity>
<taxonomicName id="4C0259A02A26FFF3C669FEDA4BA35D02" box="[1282,1435,332,357]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cordylinum">
<emphasis id="B976FE312A26FFF3C669FEDA4BA35D02" box="[1282,1435,332,357]" italics="true" pageId="3" pageNumber="163">P. cordylinum</emphasis>
</taxonomicName>
and
<quantity id="4CFA8FC62A26FFF3C3A1FEF84F3E5DEF" box="[202,262,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="3" pageNumber="163" unit="in" value="26.0">26 in</quantity>
<taxonomicName id="4C0259A02A26FFF3C27BFEE64FA95DEF" box="[272,401,369,392]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cipoense">
<emphasis id="B976FE312A26FFF3C27BFEE64FA95DEF" box="[272,401,369,392]" italics="true" pageId="3" pageNumber="163">P. cipoense</emphasis>
</taxonomicName>
).
<taxonomicName id="4C0259A02A26FFF3C2C7FEF84CAD5DEF" box="[428,661,367,392]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="cipoense">
<emphasis id="B976FE312A26FFF3C2C7FEF84CAD5DEF" box="[428,661,367,392]" italics="true" pageId="3" pageNumber="163">Placosoma cipoense</emphasis>
</taxonomicName>
is the only species in the genus to have the superior part of nasal semidivided; the suture in also present in the lower part of nasal in
<taxonomicName id="4C0259A02A26FFF3C080FE024A945DCA" box="[1003,1196,404,429]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A26FFF3C080FE024A945DCA" box="[1003,1196,404,429]" italics="true" pageId="3" pageNumber="163">P. limaverdorum</emphasis>
</taxonomicName>
. Additionally, in
<taxonomicName id="4C0259A02A26FFF3C6ECFE024F0D5DB7" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A26FFF3C6ECFE024F0D5DB7" italics="true" pageId="3" pageNumber="163">P. limaverdorum</emphasis>
</taxonomicName>
, the frontal scale has anterior and posteriorly the same approximate width, is smaller than parietal and internasal and is either totally separated from the first supraocular or in punctual contact with it. In all other species the frontal scale is longer than parietal and internasal and is in broad contact with the first supraocular. Another difference is that the suture between frontoparietals is relatively longer in
<taxonomicName id="4C0259A02A26FFF3C7FBFDB24B685E5A" box="[1168,1360,548,573]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="163" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A26FFF3C7FBFDB24B685E5A" box="[1168,1360,548,573]" italics="true" pageId="3" pageNumber="163">P. limaverdorum</emphasis>
</taxonomicName>
when compared with those of its congeners.
</paragraph>
</subSubSection>
<subSubSection id="C31871A82A26FFF4C3ACFDFB4BA55F3A" lastPageId="4" lastPageNumber="164" pageId="3" pageNumber="163" type="description">
<paragraph id="8BBD22232A26FFF4C3ACFDFB4BA55D02" blockId="3.[151,1437,151,2018]" lastBlockId="4.[151,1437,151,1334]" lastPageId="4" lastPageNumber="164" pageId="3" pageNumber="163">
<emphasis id="B976FE312A26FFF3C3ACFDFB4C1B5EE2" bold="true" box="[199,547,620,645]" pageId="3" pageNumber="163">
Description of the
<typeStatus id="54B99C812A26FFF3C2C4FDFB4C265EE2" box="[431,542,620,645]" pageId="3" pageNumber="163" type="holotype">Holotype</typeStatus>
.
</emphasis>
Measurements (mm): snout-vent length 52,8; tail length 60,2 (regenerated); head length 13,4, head width 8,2 and head height 7,0; right forelimb and hindlimb,13,8 and 20,5, respectively. Snout long and pointed, neck distinct, with a conspicuous gular fold, collar fold absent. Rostral broad, visible dorsally, wider than high, in contact with frontonasal, nasal and first supralabial. Frontonasal pentagonal, long, in lateral contact with the nasal, indenting posteriorly a pair of prefrontals. Frontal hexagonal, longer than wide, as long as frontonasal and frontoparietal suture, contacting laterally first (punctualy), second (largely) and third (moderately) supraoculars and posteriorly a pair of longer than wide pentagonal frontoparietals. Interparietal as wide as and longer than frontal, posteriorly rounded, indenting anteriorly the frontoparietals. Parietals wide, irregularly hexagonal, rounded laterally, contacting anteriorly fourth supraocular and frontoparietals, laterally temporals and posteriorly the occipitals. A row of three occipitals following parietal and interparietal the central one smaller the lateral ones in contact with the temporal and a posterior pair of nucals. Supraoculars four, first the smallest, the other three irregular in shape but with the same approximate size. Nasal large, divided above and below the median nostril, contacting inferiorly first and second supralabials. Loreal single, higher than wide, contacting posterior part of nasal, prefrontal, first superciliary, frenorbital, preocular and second supralabial. Frenorbital triangular, smaller than posterior part of nasal, in broad contact with first superciliar and preocular. Superciliaries six/five (R/L), first largest. Lower eyelid smooth, without scale division, with a transparent palpebral disc. Suboculars six/five (R/L), posterior three the largest. Postoculars one/two (R/L). Supralabials eight, 4th under the eye, 4th and 6th the largest. A series of temporals irregular in size and shape, those close to dorsal ones much enlarged. Ear opening semicircular bordered by series of small and flat elongated scales. Tympanum distinct, and close to head surface; external auditory meatus reduced. Mental small, semicircular, in straight contact with a pentagonal posmental which is as long as wide, wider posteriorly and in contact with first infralabial. Three enlarged pairs of genials, the first two in broad contact, the third pair separated by irregular scales in the middle, all contacting infralabials. A smaller pair of chinshields broadely separated at midline and by the infralabials by an elongated scale. Infralabials elongated, eight/seven (R/L), the latest very small, the fifth the longest. Dorsal region of the neck with a transverse row of two enlarged and wider than long nuchals at midline following occipitals. A series of six-seven pairs of midline dorsals, smooth, slightly smaller than the nucals following them and gradually becoming identical to other dorsals. Other dorsal scales quadrangular, with straight edges or slightly rounded, imbricate, smooth, and organized in 38 regular transverse rows (between nucals and posterior level of hindlimbs). Six enlarged dorsals between forelimbs. Lateral region with smaller scales, smooth, imbricate, elongated, not well defined in regular transverse rows. Gular region with 12 transverse rows of irregular scales between the last pair of chinshields and the interbrachial region (included). Interbrachial region with a distinct gular fold, with nine scales, all quadrangular, medial ones larger. A distinctive granular area near forelimb and hindlimb insertion. Ventral scales squared, smooth, slightly imbricate, organized in longitudinal rows and 23 regular transverse rows from the gular fold to the row before the upper preanals. Scales around midbody 33: 10 ventrals, 4+7 laterals, 12 dorsals. Preanal region with two series of smooth and almost juxtaposed scales. Upper row with four scales, central ones largest and wider than long, six scales in the lower row, all longer than large. Twenty one total femoral
</paragraph>
<paragraph id="8BBD22232A21FFF4C3ACFEF84CB05ECF" blockId="4.[151,1437,151,1334]" pageId="4" pageNumber="164">
<emphasis id="B976FE312A21FFF4C3ACFEF84F005DEF" bold="true" box="[199,312,367,392]" pageId="4" pageNumber="164">Coloring.</emphasis>
Dorsal surfaces of body and tail and lateral part of tail grayish brown with a conspicuous light vertebral stripe, one and half to two scales wide, extending from rostral to the tail. Near the base of tail the light stripe widens assumes a greyish tonality and occupy most of its dorsal part. Vertebral stripe bordered on the sides along its entire length by dark colour, usually interspersed with irregularly disposed light ocelli, which also occur on the flanks and dorsal surfaces of the limbs. Ventral surface of head and body creamy white, dotted with small dark spots. Ventral parts of limbs and tail with the same general color but with less intense dotting. Lateral surface of the head with a short and thin light stripe extending from preocular to temporals and bordered above by a thinner dark stripe that cross ear opening, goes over the front limbs and extends to the flanks. Supralabials and infralabials light colored with scanty dark pigmentation.
</paragraph>
<paragraph id="8BBD22232A21FFF4C3ACFD234BA55F3A" blockId="4.[151,1437,151,1334]" pageId="4" pageNumber="164">
<emphasis id="B976FE312A21FFF4C3ACFD234F035EAA" bold="true" box="[199,315,692,717]" pageId="4" pageNumber="164">Variation.</emphasis>
There is no sexual variation in meristic or morphometric characters in the
<typeStatus id="54B99C812A21FFF4C7C2FD224AE15EAA" box="[1193,1241,693,717]" pageId="4" pageNumber="164">type</typeStatus>
series (
<tableCitation id="C68017982A21FFF4C65FFD234BB25EAB" box="[1332,1418,692,717]" captionStart="TABLE 1" captionStartId="4.[151,239,1382,1405]" captionTargetBox="[159,1414,1548,2025]" captionTargetPageId="4" captionText="TABLE 1. Meristic and morphometric (mm) data (mean ± one standard deviation) of the type series of Placosoma limaverdorum sp. nov." httpUri="http://table.plazi.org/id/DF7D72AB2A21FFF4C3FCFAF14FB159FA" pageId="4" pageNumber="164" tableUuid="DF7D72AB2A21FFF4C3FCFAF14FB159FA">Table 1</tableCitation>
). Likewise, sexual color dimorphism is apparently absent although females seem more dimly colored than males. One male (CHUFC L4170) showed small orange spots disposed in a irregularly longitudinal series along sides of body, complemented by similar scattered spots on limbs, ventral region and near the cloaca (
<figureCitation id="13393EA62A21FFF4C7DFFC884B2E5F5F" box="[1204,1302,799,824]" captionStart="FIGURE 3" captionStartId="6.[151,250,1849,1871]" captionTargetBox="[151,1436,1416,1827]" captionTargetId="figure@6.[151,1436,1416,1827]" captionTargetPageId="6" captionText="FIGURE 3. Color in life of two paratypes of Placosoma limaverdorum sp. nov. (A) CHUFC L 4170, adult male and (B) CHUFC L 4171, adult female. (Photos: D. M. Borges-Nojosa)." httpUri="https://zenodo.org/record/262094/files/figure.png" pageId="4" pageNumber="164">Figure 3</figureCitation>
). Preserved specimens become darker but maintain a colour pattern similar to that of the live specimens except for the intensity.
</paragraph>
</subSubSection>
<subSubSection id="C31871A82A21FFF4C3ACFCF04A20599E" pageId="4" pageNumber="164" type="distribution">
<paragraph id="8BBD22232A21FFF4C3ACFCF04D125951" blockId="4.[151,1437,151,1334]" pageId="4" pageNumber="164">
<emphasis id="B976FE312A21FFF4C3ACFCF04C665FE7" bold="true" box="[199,606,871,896]" pageId="4" pageNumber="164">Distribution and natural history.</emphasis>
The
<typeStatus id="54B99C812A21FFF4C1C9FCFF4CEA5FE7" box="[674,722,872,896]" pageId="4" pageNumber="164">type</typeStatus>
series of
<taxonomicName id="4C0259A02A21FFF4C025FCF04A4C5FE7" box="[846,1140,871,896]" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A21FFF4C025FCF04A4C5FE7" box="[846,1140,871,896]" italics="true" pageId="4" pageNumber="164">Placosoma limaverdorum</emphasis>
</taxonomicName>
indicate that the species occurs in the forested mountains of Serra de Baturité (4o05’–
<geoCoordinate id="EE3644E42A21FFF4C03BFC194DA15FC0" box="[848,921,907,935]" direction="south" orientation="latitude" pageId="4" pageNumber="164" precision="925" value="-4.6666665">4o40’S</geoCoordinate>
/38o30’–
<geoCoordinate id="EE3644E42A21FFF4C092FC184A595FC0" box="[1017,1121,907,935]" direction="west" orientation="longitude" pageId="4" pageNumber="164" precision="925" value="-39.166668">39o10’W</geoCoordinate>
), and Serra de Maranguape (3o54’–
<geoCoordinate id="EE3644E42A21FFF4C383FC224F085FA9" box="[232,304,946,974]" direction="south" orientation="latitude" pageId="4" pageNumber="164" precision="925" value="-4.05">4o03’S</geoCoordinate>
/38o32’–
<geoCoordinate id="EE3644E42A21FFF4C2FAFC214FCF5FA9" box="[401,503,946,974]" direction="west" orientation="longitude" pageId="4" pageNumber="164" precision="925" value="-38.666668">38o40’W</geoCoordinate>
) in the state of
<collectingRegion id="49C6ECC12A21FFF4C1D1FC214CC65FA9" box="[698,766,950,974]" country="Brazil" name="Ceara" pageId="4" pageNumber="164">Ceará</collectingRegion>
, northeastern
<collectingCountry id="F31562B32A21FFF4C0C8FC224DDF5FA9" box="[931,999,949,974]" name="Brazil" pageId="4" pageNumber="164">Brazil</collectingCountry>
from where it is apparently endemic (
<figureCitation id="13393EA62A21FFF4C3F5FC4E4F305F95" box="[158,264,985,1010]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1122]" captionTargetBox="[157,1430,199,1071]" captionTargetId="figure@7.[151,1436,193,1078]" captionTargetPageId="7" captionText="FIGURE 4. Distribution of Placosoma limaverdorum sp. nov.: Serra de Maranguape (circles), Serra da Aratanha (triangle) and Maciço de Baturité (squares; 1 — type locality) in the state of Ceará, Brazil." httpUri="https://zenodo.org/record/262095/files/figure.png" pageId="4" pageNumber="164">Figure 4</figureCitation>
). Another record, although without voucher (P.C.M.D. Mesquita, personal information and photo) confirm its occurrence at Serra da Aratanha (3º55’–
<geoCoordinate id="EE3644E42A21FFF4C18BFC6A4D105871" box="[736,808,1021,1046]" direction="south" orientation="latitude" pageId="4" pageNumber="164" precision="925" value="-3.9666667">3º58’S</geoCoordinate>
/38º38’–
<geoCoordinate id="EE3644E42A21FFF4C0E7FC6A4DCC5871" box="[908,1012,1021,1046]" direction="west" orientation="longitude" pageId="4" pageNumber="164" precision="925" value="-38.6">38º36’W</geoCoordinate>
), another nearby forest region. These areas, the called “brejos de altitude”, are forest islands isolated in the semiarid Caatingas.
<taxonomicName id="4C0259A02A21FFF4C674FBB64F0D5839" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="limaverdorum">
<emphasis id="B976FE312A21FFF4C674FBB64F0D5839" italics="true" pageId="4" pageNumber="164">Placosoma limaverdorum</emphasis>
</taxonomicName>
was always found inside the forest. Most specimens were spotted on the leaf litter of the primary and secondary vegetation, or in banana plantations (
<bibRefCitation id="EF935FD22A21FFF4C1ABFBFE4D7B58E5" author="Lima" box="[704,835,1129,1154]" pageId="4" pageNumber="164" refString="Lima, D. C. (2005) A Bananicultura na Area de Protecao da Serra de Maranguape-CE e suas Implicacoes no Ambiente Fisico, Humano e na Biodiversidade. Dissertacao apresentada ao Mestrado em Desenvolvimento e Meio Ambiente (PRODEMA) - UFC, Fortaleza-CE 109 pp." type="book" year="2005">Lima 2005</bibRefCitation>
), in sympatry with the gymnophthalmids
<taxonomicName id="4C0259A02A21FFF4C642FBFD4C5658C1" authority="Rodrigues &amp; Borges, 1997" authorityName="Rodrigues &amp; Borges" authorityYear="1997" class="Insecta" family="Curculionidae" genus="Leposoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="baturitensis">
<emphasis id="B976FE312A21FFF4C642FBFD4F2358C1" italics="true" pageId="4" pageNumber="164">Leposoma baturitensis</emphasis>
<bibRefCitation id="EF935FD22A21FFF4C241FB1A4C5658C1" author="Rodrigues" box="[298,622,1165,1190]" pageId="4" pageNumber="164" refString="Rodrigues, M. T. &amp; Borges, D. M. (1997). A new species of Leposoma (Squamata: Gymnophthalmidae) from a relictual forest in semiarid Northeastern Brazil. Herpetologica, 53 (1), 1 - 6." type="journal article" year="1997">Rodrigues &amp; Borges, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0259A02A21FFF4C1EAFB1A4D8F58C2" box="[641,951,1165,1190]" class="Reptilia" family="Gymnophthalmidae" genus="Colobosauroides" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="cearensis">
<emphasis id="B976FE312A21FFF4C1EAFB1A4D8F58C2" box="[641,951,1165,1190]" italics="true" pageId="4" pageNumber="164">Colobosauroides cearensis</emphasis>
</taxonomicName>
<bibRefCitation id="EF935FD22A21FFF4C0AEFB1A4B5D58C1" author="Cunha" box="[965,1381,1165,1190]" pageId="4" pageNumber="164" refString="Cunha, O. R., Lima-Verde, J. S. &amp; Lima, A. C. M. (1991) Novo genero e especie de lagarto do Estado do Ceara (Lacertilia: Teiidae). Boletim Museu Paraense Emilio Goeldi, Zoologia, 7 (2), 163 - 176." type="journal article" year="1991">
<collectingCountry id="F31562B32A21FFF4C0AEFB1A4A2958C1" box="[965,1041,1165,1190]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="164">Cunha</collectingCountry>
, Lima-Verde &amp; Lima, 1991
</bibRefCitation>
and
<taxonomicName id="4C0259A02A21FFF4C3FCFB264C2458AD" authority="Boulenger, 1887" authorityName="Boulenger" authorityYear="1887" box="[151,540,1201,1226]" class="Reptilia" family="Gymnophthalmidae" genus="Stenolepis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="ridleyi">
<emphasis id="B976FE312A21FFF4C3FCFB264F6158AD" box="[151,345,1201,1226]" italics="true" pageId="4" pageNumber="164">Stenolepis ridleyi</emphasis>
<bibRefCitation id="EF935FD22A21FFF4C234FB264C2458AD" author="Boulenger" box="[351,540,1201,1226]" pageId="4" pageNumber="164" refString="Boulenger, G. A. (1887) Description of a new genus of lizards of the Family Teiidae. Proceedings of the Zoological Society of London, 1887, 640 - 642. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1887. tb 08157. x" type="proceedings paper" year="1887">Boulenger, 1887</bibRefCitation>
</taxonomicName>
. Two specimens were found in tree trunks, one on a central part of the trunk of a thin tree, about 1.3 m high, and the other in a vertical branch, about 1.2 m from the ground, between epiphytes and lichens. Three specimens were taken from the stomach contents of the snake
<taxonomicName id="4C0259A02A21FFF4C76FFB6E4BA45975" authority="Peters, 1863" authorityName="Peters" authorityYear="1863" box="[1028,1436,1273,1298]" class="Reptilia" family="Colubridae" genus="Drymoluber" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="dichrous">
<emphasis id="B976FE312A21FFF4C76FFB6E4ACE5975" box="[1028,1270,1273,1298]" italics="true" pageId="4" pageNumber="164">Drymoluber dichrous</emphasis>
(
<bibRefCitation id="EF935FD22A21FFF4C66EFB6D4BAB5975" author="Peters" box="[1285,1427,1273,1298]" pageId="4" pageNumber="164" refString="Peters, W. (1863) Hr. W. Peters berichtet uber einige neue oder weniger bekannte Schlangenarten des zoologischen Museums zu Berlin. Monatshefte fur Akademie der Wissenschaften zu Berlin, 29, 272 - 289." type="journal article" year="1863">Peters, 1863</bibRefCitation>
)
</taxonomicName>
from Serra de Maranguape (
<bibRefCitation id="EF935FD22A21FFF4C2B9FA8A4D255951" author="Borges-Nojosa" box="[466,797,1309,1334]" pageId="4" pageNumber="164" refString="Borges-Nojosa, D. M. &amp; Lima, D. C. (2001) Dieta de Drymoluber dichrous (Peters, 1863) dos Brejos-de-Altitude do Estado do Ceara - Brasil (Serpentes, Colubridae). Boletim do Museu Nacional, New Series (Zoologia), 468, 1 - 5." type="journal article" year="2001">Borges-Nojosa &amp; Lima 2001</bibRefCitation>
).
</paragraph>
<caption id="DF7D72AB2A21FFF4C3FCFAF14FB159FA" ID-Table-UUID="DF7D72AB2A21FFF4C3FCFAF14FB159FA" httpUri="http://table.plazi.org/id/DF7D72AB2A21FFF4C3FCFAF14FB159FA" pageId="4" pageNumber="164" targetBox="[159,1414,1548,2025]" targetIsTable="true" targetPageId="4">
<paragraph id="8BBD22232A21FFF4C3FCFAF14FB159FA" blockId="4.[151,1435,1381,1437]" pageId="4" pageNumber="164">
<emphasis id="B976FE312A21FFF4C3FCFAF14F35591A" bold="true" box="[151,269,1382,1405]" pageId="4" pageNumber="164">TABLE 1.</emphasis>
Meristic and morphometric (mm) data (mean ± one standard deviation) of the type series of
<taxonomicName id="4C0259A02A21FFF4C64EFAF24F1559FA" class="Reptilia" family="Gymnophthalmidae" genus="Placosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="164" phylum="Chordata" rank="species" species="limaverdorum" status="sp. nov.">
<emphasis id="B976FE312A21FFF4C64EFAF24F1559FA" italics="true" pageId="4" pageNumber="164">Placosoma limaverdorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245434A2A21FFF4C25FFA114FB159FA" box="[308,393,1414,1437]" pageId="4" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBD22232A21FFF4C3F4FA2E4B2359A8" blockId="4.[159,1307,1465,1529]" box="[159,1307,1465,1487]" pageId="4" pageNumber="164">Characters Holotype Paratypes Total (n=13)</paragraph>
<paragraph id="8BBD22232A21FFF4C104FA744A20599E" blockId="4.[159,1307,1465,1529]" box="[623,1048,1507,1529]" pageId="4" pageNumber="164">Males (n=8) Females (n=4)</paragraph>
</subSubSection>
</treatment>
</document>
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