<documentid="DF333A5C8B3CAC6F2F8180A9814B0BD7"ID-CLB-Dataset="302884"ID-DOI="10.5852/ejt.2024.950.2643"ID-GBIF-Dataset="81037ee2-a646-42f0-beb2-c315dec5320c"ID-ISSN="2118-9773"ID-Zenodo-Dep="13691805"ID-ZooBank="ACFB8240-3FDC-43B8-8200-236BFE00AD94"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="juliana"IM.treatmentCitations_approvedBy="juliana"IM.treatments_approvedBy="juliana"checkinTime="1725525331103"checkinUser="plazi"docAuthor="Onuferko, Thomas M. & Rightmyer, Molly G."docDate="2024"docId="038F531FFFF3FFB93008FB86FC44FDED"docLanguage="en"docName="EJT.2024.950.1-106.pdf"docOrigin="European Journal of Taxonomy 950 (1)"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2643/12195"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Triepeolus Robertson 1901"docType="key"docVersion="4"lastPageNumber="18"masterDocId="FFB62B67FFF5FFA83244FFE9FF88FFEF"masterDocTitle="A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae)"masterLastPageNumber="106"masterPageNumber="1"pageNumber="7"updateTime="1725986112318"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="3A6AC2F928525962B51320A648BD68A2">A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae)</mods:title>
<mods:affiliationid="44A532E56518414AB2BA54EFE9C1628E">Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, ON K 1 P 6 P 4, Canada and University of Toronto Scarborough, 1265 Military Trail, Toronto, ON M 1 C 1 A 4, Canada.</mods:affiliation>
<mods:affiliationid="F81D14CCDE45242484DAD3E95898F0DC">Department of Entomology, San Diego Natural History Museum, 1788 El Prado, San Diego, CA 92101, USA.</mods:affiliation>
group are easily recognized by the well-defined trapezoidal or triangular pseudopygidial area, which has reflective (golden/coppery to silvery) setae, and concave apical margin of T5 (
<figureCitationid="131DFE8CFFF3FFAE3008FACCFD1AFAD0"box="[588,658,1317,1343]"captionStart="Fig"captionStartId="5.[189,232,1963,1989]"captionTargetBox="[211,1376,1177,1890]"captionTargetId="figure-276@5.[189,1398,1139,1926]"captionTargetPageId="5"captionText="Fig. 1. Female T. tuberculifer Onuferko, Rightmyer & Roig-Alsina, 2024 illustrating the mesosomal and metasomal bands of tomentum commonly present in Triepeolus Robertson, 1901."figureDoi="http://doi.org/10.5281/zenodo.13691807"httpUri="https://zenodo.org/record/13691807/files/figure.png"pageId="6"pageNumber="7">Fig. 1</figureCitation>
; see also photographs in species treatments below, but note the reflective setae are not clearly visible in some images of the pseudopygidial area due to diffused lighting). Females of three other
<bibRefCitationid="EFB79FF8FFF3FFAE36F9F9F5FAE1F9D9"author="Cresson E. T."box="[1213,1385,1564,1590]"pageId="6"pageNumber="7"pagination="61 - 136"refId="ref73927"refString="Cresson E. T. 1878. Descriptions of new North American Hymenoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society 7: 61 - 136. https: // doi. org / 10.2307 / 25076368"type="journal article"year="1878">Cresson, 1878</bibRefCitation>
)
</taxonomicName>
, the pseudopygidial area of the female has golden to silvery setae but is more or less rectangular in shape (
<figureCitationid="131DFE8CFFF3FFAE3281F98BFEA9F993"box="[197,289,1634,1660]"captionStart="Fig"captionStartId="7.[189,232,1297,1323]"captionTargetBox="[218,1370,820,1244]"captionTargetId="figure-210@7.[218,1370,820,1259]"captionTargetPageId="7"captionText="Fig. 2. Pseudopygidial area of female, dorsal view. A. Triepeolus dacotensis (Stevens, 1919) (BOLD sample ID: CCDB-32640 E03; RSKM RSKM_ENT_E-186200). B. T. distinctus (Cresson, 1878) (OKBS OKBS.POL.314)."figureDoi="http://doi.org/10.5281/zenodo.13691809"httpUri="https://zenodo.org/record/13691809/files/figure.png"pageId="6"pageNumber="7">Fig. 2B</figureCitation>
group are not easily recognized as no single morphological feature or unique combination of features has been identified that can be considered diagnostic, not in the external morphology or terminalia.
<bibRefCitationid="EFB79FF8FFF3FFAE33BCF941FD42F92D"author="Rightmyer M. G."box="[504,714,1704,1730]"pageId="6"pageNumber="7"pagination="1 - 51"refId="ref75792"refString="Rightmyer M. G. 2004. Phylogeny and classification of the parasitic bee tribe Epeolini (Hymenoptera: Apidae, Nomadinae). Scientific Papers, Natural History Museum, The University of Kansas 33: 1 - 51."type="journal article"year="2004">Rightmyer (2004)</bibRefCitation>
<bibRefCitationid="EFB79FF8FFF3FFAE3025F925FD70F909"author="Rozen J. G. Jr"box="[609,760,1740,1766]"pageId="6"pageNumber="7"pagination="1 - 18"refId="ref76133"refString="Rozen J. G. Jr 1989. Two new species and the redescription of another species of the cleptoparasitic bee genus Triepeolus with notes on their immature stages (Anthophoridae: Nomadinae). American Museum Novitates 2956: 1 - 18. Available from https: // www. biodiversitylibrary. org / page / 62309460 [accessed 25 Oct. 2023]."type="journal article"year="1989">Rozen, 1989</bibRefCitation>
</taxonomicName>
, among the 21 morphological characters pertaining to the male S7, S8, or genitalia assessed for 11 markedly different species in the genus. Although several morphological features have been identified as common to males of the
species groups (see Supp. file 3 for the complete list). In lieu of a diagnosis, a fully illustrated key is presented to enable the identification of males of the
1. Any of the following (alone or in combination): clypeus with well-defined glabrous midline, which may extend its entire length (
<figureCitationid="131DFE8CFFF2FFAF3005FEB3FD28FE9B"box="[577,672,346,372]"captionStart="Fig"captionStartId="7.[189,232,1928,1954]"captionTargetBox="[218,1370,1453,1877]"captionTargetId="figure-212@7.[218,1370,1453,1892]"captionTargetPageId="7"captionText="Fig. 3. Head of male, frontal view. A. Triepeolus michiganensis Mitchell, 1962 (CNC 719805), which has a strong glabrous clypeal midline (blue arrow). B. T. obliteratus Graenicher, 1911 (WRME WRME_ EJH02370), which does not have a glabrous clypeal midline."figureDoi="http://doi.org/10.5281/zenodo.13691811"httpUri="https://zenodo.org/record/13691811/files/figure.png"pageId="7"pageNumber="8">Fig. 3A</figureCitation>
); F2 shorter than F1 (
<figureCitationid="131DFE8CFFF2FFAF31DBFEB3FC76FE9B"box="[927,1022,346,372]"captionStart="Fig"captionStartId="8.[189,232,757,783]"captionTargetBox="[218,1370,275,699]"captionTargetId="figure-19@8.[218,1370,275,714]"captionTargetPageId="8"captionText="Fig. 4. Right antenna (basal portion) of male. A. Triepeolus loomisorum Rozen, 1989 (PCYU), showing the longer side of F1 noticeably longer than F2. B. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28), showing the longer side of F1 and F2 subequal in length. Blue lines indicate the greatest length of F1."figureDoi="http://doi.org/10.5281/zenodo.13691813"httpUri="https://zenodo.org/record/13691813/files/figure.png"pageId="7"pageNumber="8">Fig. 4A</figureCitation>
); mesoscutellum to some extent reddish brown (
<figureCitationid="131DFE8CFFF2FFAF33E0FE94FD8BFE77"box="[420,515,381,408]"captionStart="Fig"captionStartId="8.[189,232,1928,1954]"captionTargetBox="[218,1370,1018,1882]"captionTargetId="figure-78@8.[218,1370,1018,1887]"captionTargetPageId="8"captionText="Fig. 5. Axillae and mesoscutellum of male, dorsal view. A. Triepeolus intrepidus (Smith, 1879) (PCYU PCYU-SD09:612). B. T. roni Genaro, 1999 (USNM M.G.R. Database No. 5922). C. T. hirsutus sp. nov., allotype (PCYU). D. T. brittaini Cockerell, 1931 (BOLD sample ID: 05-NS-0718; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13691815"httpUri="https://zenodo.org/record/13691815/files/figure.png"pageId="7"pageNumber="8">Fig. 5A</figureCitation>
); mesopleuron ventrolaterally with dense, long (≥ ½ MOD), erect/suberect, simple or minutely branched setae (
<figureCitationid="131DFE8CFFF2FFAF30CEFE48FD60FE55"box="[650,744,416,443]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="7"pageNumber="8">Fig. 6A</figureCitation>
); propodeal triangle mostly covered in dense, appressed pale setae (
<figureCitationid="131DFE8CFFF2FFAF3330FE2DFE50FE31"box="[372,472,452,478]"captionStart="Fig"captionStartId="9.[189,231,1928,1954]"captionTargetBox="[218,1370,1441,1865]"captionTargetId="figure-98@9.[218,1370,1441,1881]"captionTargetPageId="9"captionText="Fig. 7. Mesosoma of male, posterior view. A. Triepeolus eliseae Rightmyer, 2017 (INHS 356997), in which the propodeal triangle is mostly covered in dense, appressed pale setae.B. T. obliteratus Graenicher, 1911 (WRME WRME_EJH02370), in which the propodeal triangle is mostly to entirely glabrous."figureDoi="http://doi.org/10.5281/zenodo.13691819"httpUri="https://zenodo.org/record/13691819/files/figure.png"pageId="7"pageNumber="8">Fig. 7A</figureCitation>
); and/or pygidial plate with well-defined basal or subapical transverse ridge (
<figureCitationid="131DFE8CFFF2FFAF32B7FE0EFEC7FDEE"box="[243,335,487,513]"captionStart="Fig"captionStartId="10.[189,232,765,791]"captionTargetBox="[218,1370,276,700]"captionTargetId="figure-19@10.[218,1370,276,716]"captionTargetPageId="10"captionText="Fig. 8. Pygidial plate of male, dorsal view. A. Triepeolus rufithorax Graenicher, 1928 (PCYU), which has a well-defined transverse ridge (blue arrow). B. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28), which does not have a transverse ridge."figureDoi="http://doi.org/10.5281/zenodo.13691821"httpUri="https://zenodo.org/record/13691821/files/figure.png"pageId="7"pageNumber="8">Fig. 8A</figureCitation>
) ................................................................... most other
– The following in combination: clypeus without glabrous midline or rarely with ill-defined glabrous midline restricted to upper half (
<figureCitationid="131DFE8CFFF2FFAF3022FDC4FD49FDA7"box="[614,705,557,584]"captionStart="Fig"captionStartId="7.[189,232,1928,1954]"captionTargetBox="[218,1370,1453,1877]"captionTargetId="figure-212@7.[218,1370,1453,1892]"captionTargetPageId="7"captionText="Fig. 3. Head of male, frontal view. A. Triepeolus michiganensis Mitchell, 1962 (CNC 719805), which has a strong glabrous clypeal midline (blue arrow). B. T. obliteratus Graenicher, 1911 (WRME WRME_ EJH02370), which does not have a glabrous clypeal midline."figureDoi="http://doi.org/10.5281/zenodo.13691811"httpUri="https://zenodo.org/record/13691811/files/figure.png"pageId="7"pageNumber="8">Fig. 3B</figureCitation>
); F2 as long as or longer than F1 (
<figureCitationid="131DFE8CFFF2FFAF3610FDC4FB39FDA7"box="[1108,1201,557,584]"captionStart="Fig"captionStartId="8.[189,232,757,783]"captionTargetBox="[218,1370,275,699]"captionTargetId="figure-19@8.[218,1370,275,714]"captionTargetPageId="8"captionText="Fig. 4. Right antenna (basal portion) of male. A. Triepeolus loomisorum Rozen, 1989 (PCYU), showing the longer side of F1 noticeably longer than F2. B. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28), showing the longer side of F1 and F2 subequal in length. Blue lines indicate the greatest length of F1."figureDoi="http://doi.org/10.5281/zenodo.13691813"httpUri="https://zenodo.org/record/13691813/files/figure.png"pageId="7"pageNumber="8">Fig. 4B</figureCitation>
); mesoscutellum black (
<figureCitationid="131DFE8CFFF2FFAF337CFDB8FE3FFD84"box="[312,439,592,619]"captionStart="Fig"captionStartId="8.[189,232,1928,1954]"captionTargetBox="[218,1370,1018,1882]"captionTargetId="figure-78@8.[218,1370,1018,1887]"captionTargetPageId="8"captionText="Fig. 5. Axillae and mesoscutellum of male, dorsal view. A. Triepeolus intrepidus (Smith, 1879) (PCYU PCYU-SD09:612). B. T. roni Genaro, 1999 (USNM M.G.R. Database No. 5922). C. T. hirsutus sp. nov., allotype (PCYU). D. T. brittaini Cockerell, 1931 (BOLD sample ID: 05-NS-0718; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13691815"httpUri="https://zenodo.org/record/13691815/files/figure.png"pageId="7"pageNumber="8">Fig. 5B–D</figureCitation>
); mesopleuron ventrolaterally at most with sparse, long (≥ ½ MOD), erect/suberect, simple setae (
<figureCitationid="131DFE8CFFF2FFAF33D5FD9DFE67FD61"box="[401,495,628,654]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="7"pageNumber="8">Fig. 6B</figureCitation>
) in addition to usual appressed, branched setae or only short (<½ MOD), appressed setae (
<figureCitationid="131DFE8CFFF2FFAF33F2FD7EFDB3FD5E"box="[438,571,663,689]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="7"pageNumber="8">Fig. 6C–D</figureCitation>
); propodeal triangle mostly to entirely glabrous, with any pale setae usually restricted to small lateral patches (
<figureCitationid="131DFE8CFFF2FFAF30A6FD53FCC8FD3B"box="[738,832,698,724]"captionStart="Fig"captionStartId="9.[189,231,1928,1954]"captionTargetBox="[218,1370,1441,1865]"captionTargetId="figure-98@9.[218,1370,1441,1881]"captionTargetPageId="9"captionText="Fig. 7. Mesosoma of male, posterior view. A. Triepeolus eliseae Rightmyer, 2017 (INHS 356997), in which the propodeal triangle is mostly covered in dense, appressed pale setae.B. T. obliteratus Graenicher, 1911 (WRME WRME_EJH02370), in which the propodeal triangle is mostly to entirely glabrous."figureDoi="http://doi.org/10.5281/zenodo.13691819"httpUri="https://zenodo.org/record/13691819/files/figure.png"pageId="7"pageNumber="8">Fig. 7B</figureCitation>
); and pygidial plate without transverse ridge or rarely with ill-defined basal transverse ridge (
<figureCitationid="131DFE8CFFF2FFAF3145FD34FCD3FD17"box="[769,859,733,760]"captionStart="Fig"captionStartId="10.[189,232,765,791]"captionTargetBox="[218,1370,276,700]"captionTargetId="figure-19@10.[218,1370,276,716]"captionTargetPageId="10"captionText="Fig. 8. Pygidial plate of male, dorsal view. A. Triepeolus rufithorax Graenicher, 1928 (PCYU), which has a well-defined transverse ridge (blue arrow). B. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28), which does not have a transverse ridge."figureDoi="http://doi.org/10.5281/zenodo.13691821"httpUri="https://zenodo.org/record/13691821/files/figure.png"pageId="7"pageNumber="8">Fig. 8B</figureCitation>
<bibRefCitationid="EFB79FF8FFF2FFAF36F8FADCFAE6FAA0"author="Cresson E. T."box="[1212,1390,1333,1359]"pageId="7"pageNumber="8"pagination="61 - 136"refId="ref73927"refString="Cresson E. T. 1878. Descriptions of new North American Hymenoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society 7: 61 - 136. https: // doi. org / 10.2307 / 25076368"type="journal article"year="1878">Cresson, 1878</bibRefCitation>
<bibRefCitationid="EFB79FF8FFF2FFAF31FDF861FBEFF84D"author="Mitchell T. B."box="[953,1127,1928,1954]"pageId="7"pageNumber="8"pagination="1 - 557"refId="ref75083"refString="Mitchell T. B. 1962. Bees of the eastern United States. Volume II. North Carolina Agricultural Experiment Station Technical Bulletin 152: 1 - 557."type="journal article"year="1962">Mitchell, 1962</bibRefCitation>
</taxonomicName>
(CNC 719805), which has a strong glabrous clypeal midline (blue arrow).
<bibRefCitationid="EFB79FF8FFF2FFAF3197F842FB14F82A"author="Graenicher S."box="[979,1180,1963,1989]"pageId="7"pageNumber="8"pagination="221 - 249"refId="ref74330"refString="Graenicher S. 1911. Bees of northwestern Wisconsin. Bulletin of the Public Museum of the City of Milwaukee 1: 221 - 249."type="journal article"year="1911">Graenicher, 1911</bibRefCitation>
</taxonomicName>
(WRME WRME_ EJH02370), which does not have a glabrous clypeal midline.
<bibRefCitationid="EFB79FF8FFFDFFA0364EFD1FFB15FCE0"author="Rozen J. G. Jr"box="[1034,1181,757,784]"pageId="8"pageNumber="9"pagination="1 - 18"refId="ref76133"refString="Rozen J. G. Jr 1989. Two new species and the redescription of another species of the cleptoparasitic bee genus Triepeolus with notes on their immature stages (Anthophoridae: Nomadinae). American Museum Novitates 2956: 1 - 18. Available from https: // www. biodiversitylibrary. org / page / 62309460 [accessed 25 Oct. 2023]."type="journal article"year="1989">Rozen, 1989</bibRefCitation>
</taxonomicName>
(PCYU), showing the longer side of F1 noticeably longer than F2.
<taxonomicNameid="4C26998AFFFDFFA0315FFCF0FC61FCDD"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[795,1001,792,818]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="8"pageNumber="9"phylum="Arthropoda"rank="species"species="oblongimacula"status="sp. nov.">
, allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28), showing the longer side of F1 and F2 subequal in length. Blue lines indicate the greatest length of F1.
<bibRefCitationid="EFB79FF8FFFDFFA03052F842FD3DF82A"author="Genaro J. A."box="[534,693,1963,1989]"pageId="8"pageNumber="9"pagination="215 - 220"refId="ref74229"refString="Genaro J. A. 1999. Revision del genero Triepeolus en Cuba (Hymenoptera: Apidae), con descripcion de dos especies nuevas. Caribbean Journal of Science 35: 215 - 220."type="journal article"year="1999">Genaro, 1999</bibRefCitation>
<taxonomicNameid="4C26998AFFFDFFA036D2F845FA98F82A"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[1174,1296,1963,1989]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="8"pageNumber="9"phylum="Arthropoda"rank="species"species="hirsutus"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFFCFFA1366AFB78FB76FB44"author="Rightmyer M. G."box="[1070,1278,1169,1195]"pageId="9"pageNumber="10"pagination="1 - 170"refId="ref75868"refString="Rightmyer M. G. 2008. A review of the cleptoparasitic bee genus Triepeolus (Hymenoptera: Apidae). - Part I. Zootaxa 1710: 1 - 170. https: // doi. org / 10.11646 / zootaxa. 1710.1.1"type="journal article"year="2008">Rightmyer, 2008</bibRefCitation>
</taxonomicName>
(CSUC), which ventrolaterally has dense, long (≥ ½ MOD), erect/suberect, minutely branched setae (blue arrow).
<taxonomicNameid="4C26998AFFFCFFA132A4FB31FE1BFB1D"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[224,403,1240,1266]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="9"pageNumber="10"phylum="Arthropoda"rank="species"species="parahirsutus"status="sp. nov.">
, allotype (ZMB 21261), which ventrolaterally has sparse, long (≥ ½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow).
<bibRefCitationid="EFB79FF8FFFCFFA132F9FAF7FE0AFAD7"author="Rightmyer M. G."box="[189,386,1310,1336]"pageId="9"pageNumber="10"pagination="1 - 170"refId="ref75868"refString="Rightmyer M. G. 2008. A review of the cleptoparasitic bee genus Triepeolus (Hymenoptera: Apidae). - Part I. Zootaxa 1710: 1 - 170. https: // doi. org / 10.11646 / zootaxa. 1710.1.1"type="journal article"year="2008">Rightmyer, 2008</bibRefCitation>
<taxonomicNameid="4C26998AFFFCFFA13031FAF6FCB9FAD7"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[629,817,1310,1336]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="9"pageNumber="10"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFFCFFA136ABF842FF7CF807"author="Graenicher S."pageId="9"pageNumber="10"pagination="221 - 249"refId="ref74330"refString="Graenicher S. 1911. Bees of northwestern Wisconsin. Bulletin of the Public Museum of the City of Milwaukee 1: 221 - 249."type="journal article"year="1911">Graenicher, 1911</bibRefCitation>
</taxonomicName>
(WRME WRME_EJH02370), in which the propodeal triangle is mostly to entirely glabrous.
<taxonomicNameid="4C26998AFFFFFFA2315BFCC8FC64FCD5"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[799,1004,800,826]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="10"pageNumber="11"phylum="Arthropoda"rank="species"species="oblongimacula"status="sp. nov.">
<taxonomicNameid="4C26998AFFFFFFA23620F88EFD46F84A"authority="Onuferko, Rightmyer & Roig-Alsina, 2024"authorityName="Onuferko, Rightmyer & Roig-Alsina"authorityYear="2024"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="10"pageNumber="11"phylum="Arthropoda"rank="species"species="tuberculifer">
<bibRefCitationid="EFB79FF8FFFFFFA232F9F862FD46F84A"author="Onuferko T. M. & Rightmyer M. G. & Melo G. A. R. & Roig-Alsina A."box="[189,718,1931,1957]"pageId="10"pageNumber="11"pagination="1 - 50"refId="ref75439"refString="Onuferko T. M., Rightmyer M. G., Melo G. A. R. & Roig-Alsina A. 2024. A revision of the South American species of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae). European Journal of Taxonomy 931: 1 - 50. https: // doi. org / 10.5852 / ejt. 2024.931.2505"type="journal article"year="2024">Onuferko, Rightmyer & Roig-Alsina, 2024</bibRefCitation>
<bibRefCitationid="EFB79FF8FFFFFFA23104F847FB83F827"author="Graenicher S."box="[832,1035,1966,1992]"pageId="10"pageNumber="11"pagination="221 - 249"refId="ref74330"refString="Graenicher S. 1911. Bees of northwestern Wisconsin. Bulletin of the Public Museum of the City of Milwaukee 1: 221 - 249."type="journal article"year="1911">Graenicher, 1911</bibRefCitation>
<taxonomicNameid="4C26998AFFFFFFA232A1F83BFE3EF804"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[229,438,2001,2027]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="10"pageNumber="11"phylum="Arthropoda"rank="species"species="oblongimacula"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFFFFFA233DDF81DFDDCF7E1"author="Cockerell T. D. A."box="[409,596,2036,2062]"pageId="10"pageNumber="11"pagination="1 - 16"refId="ref73439"refString="Cockerell T. D. A. 1921. The epeoline bees of the American Museum Rocky Mountain expeditions. American Museum Novitates 23: 1 - 16. Available from http: // hdl. handle. net / 2246 / 3249 [accessed 25 Oct. 2023]."type="journal article"year="1921">Cockerell, 1921</bibRefCitation>
<figureCitationid="131DFE8CFFFEFFA33160FEE0FC08FECC"box="[804,896,265,291]"captionStart="Fig"captionStartId="10.[189,232,1895,1921]"captionTargetBox="[218,1370,952,1817]"captionTargetId="figure-63@10.[218,1370,952,1832]"captionTargetPageId="10"captionText="Fig. 9. Right pro- and mesotrochanters of male (anterior facing right). A. Triepeolus tuberculifer Onuferko, Rightmyer & Roig-Alsina, 2024, allotype (FSCA M.G.R. Database No. 3704), showing distinct protuberances (blue arrows). B. T. obliteratus Graenicher, 1911 (WRME WRME_EJH02370). C. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28). D. T. rhododontus Cockerell, 1921 (BOLD sample ID: CMNTO_139; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13742575"httpUri="https://zenodo.org/record/13742575/files/figure.png"pageId="11"pageNumber="12">Fig. 9A</figureCitation>
<taxonomicNameid="4C26998AFFFEFFA333AEFEC5FB12FEA9"authority="Onuferko, Rightmyer & Roig-Alsina, 2024"authorityName="Onuferko, Rightmyer & Roig-Alsina"authorityYear="2024"box="[490,1178,300,327]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="11"pageNumber="12"phylum="Arthropoda"rank="species"species="tuberculifer">
<bibRefCitationid="EFB79FF8FFFEFFA330DBFEC5FB12FEA9"author="Onuferko T. M. & Rightmyer M. G. & Melo G. A. R. & Roig-Alsina A."box="[671,1178,300,326]"pageId="11"pageNumber="12"pagination="1 - 50"refId="ref75439"refString="Onuferko T. M., Rightmyer M. G., Melo G. A. R. & Roig-Alsina A. 2024. A revision of the South American species of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae). European Journal of Taxonomy 931: 1 - 50. https: // doi. org / 10.5852 / ejt. 2024.931.2505"type="journal article"year="2024">Onuferko, Rightmyer & Roig-Alsina, 2024</bibRefCitation>
<figureCitationid="131DFE8CFFFEFFA33099FEA6FCD3FE86"box="[733,859,335,361]"captionStart="Fig"captionStartId="10.[189,232,1895,1921]"captionTargetBox="[218,1370,952,1817]"captionTargetId="figure-63@10.[218,1370,952,1832]"captionTargetPageId="10"captionText="Fig. 9. Right pro- and mesotrochanters of male (anterior facing right). A. Triepeolus tuberculifer Onuferko, Rightmyer & Roig-Alsina, 2024, allotype (FSCA M.G.R. Database No. 3704), showing distinct protuberances (blue arrows). B. T. obliteratus Graenicher, 1911 (WRME WRME_EJH02370). C. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28). D. T. rhododontus Cockerell, 1921 (BOLD sample ID: CMNTO_139; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13742575"httpUri="https://zenodo.org/record/13742575/files/figure.png"pageId="11"pageNumber="12">Fig. 9B–D</figureCitation>
3. Both S4 with apical/subapical fringe of dense, long (> 1 MOD), curved, brown setae and S5 with short, straight subapical setae (
<figureCitationid="131DFE8CFFFEFFA33012FE5DFD48FE21"box="[598,704,436,462]"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="11"pageNumber="12">Fig. 10A</figureCitation>
– S4 with apical/subapical fringe of dense, long (> 1 MOD), curved, coppery to silvery setae (
<figureCitationid="131DFE8CFFFEFFA33746FE3EFE88FDFA"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="11"pageNumber="12">Fig. 10B– D</figureCitation>
) and/or S5 with long, curved subapical setae (
<figureCitationid="131DFE8CFFFEFFA3315BFE13FC25FDFA"box="[799,941,506,533]"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="11"pageNumber="12">Fig. 10C–D</figureCitation>
4. T1 discal patch trapezoidal with semicircular anteromedial projection, apical transverse band conspicuously narrowed sublaterally at posterior corner of discal patch (
<figureCitationid="131DFE8CFFFEFFA33671FDB6FB28FD96"box="[1077,1184,607,633]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="11"pageNumber="12">Fig. 11A</figureCitation>
). Axilla extending well beyond midlength of mesoscutellum; tip distinctly pointed and hooked (i.e., concave along medial margin), mesally unattached to mesoscutellum for approximately half medial length of axilla (
<figureCitationid="131DFE8CFFFEFFA332B6FD20FEC3FD0C"box="[242,331,712,739]"captionStart="Fig"captionStartId="8.[189,232,1928,1954]"captionTargetBox="[218,1370,1018,1882]"captionTargetId="figure-78@8.[218,1370,1018,1887]"captionTargetPageId="8"captionText="Fig. 5. Axillae and mesoscutellum of male, dorsal view. A. Triepeolus intrepidus (Smith, 1879) (PCYU PCYU-SD09:612). B. T. roni Genaro, 1999 (USNM M.G.R. Database No. 5922). C. T. hirsutus sp. nov., allotype (PCYU). D. T. brittaini Cockerell, 1931 (BOLD sample ID: 05-NS-0718; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13691815"httpUri="https://zenodo.org/record/13691815/files/figure.png"pageId="11"pageNumber="12">Fig. 5B</figureCitation>
<bibRefCitationid="EFB79FF8FFFEFFA331BEFD20FB12FD0C"author="Genaro J. A."box="[1018,1178,713,739]"pageId="11"pageNumber="12"pagination="215 - 220"refId="ref74229"refString="Genaro J. A. 1999. Revision del genero Triepeolus en Cuba (Hymenoptera: Apidae), con descripcion de dos especies nuevas. Caribbean Journal of Science 35: 215 - 220."type="journal article"year="1999">Genaro, 1999</bibRefCitation>
<keyLeadid="30D7413CFFFEFFA33285FD05FAFFFCE8"box="[193,1399,748,775]"pageId="11"pageNumber="12">– Mesosoma and metasoma without the combination of features listed above [other regions] .......... 5</keyLead>
<bibRefCitationid="EFB79FF8FFFEFFA3301DF8A8FCABF8B3"author="Rightmyer M. G."box="[601,803,1857,1884]"pageId="11"pageNumber="12"pagination="1 - 170"refId="ref75868"refString="Rightmyer M. G. 2008. A review of the cleptoparasitic bee genus Triepeolus (Hymenoptera: Apidae). - Part I. Zootaxa 1710: 1 - 170. https: // doi. org / 10.11646 / zootaxa. 1710.1.1"type="journal article"year="2008">Rightmyer, 2008</bibRefCitation>
</taxonomicName>
(RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow).
<taxonomicNameid="4C26998AFFFEFFA3330CF845FD8CF82A"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[328,516,1963,1989]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="11"pageNumber="12"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
5. T1 with basal band of pale tomentum arched, on each side (or each if medially interrupted) continuous with (and indistinguishable from) lateral longitudinal band, such that discal patch triangular, semicircular, or crescent-shaped (
<figureCitationid="131DFE8CFFF9FFA43030FEA6FD76FE86"box="[628,766,335,361]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="12"pageNumber="13">Fig. 11B–C</figureCitation>
<bibRefCitationid="EFB79FF8FFF9FFA43693F8F7FAFFF8D7"author="Genaro J. A."box="[1239,1399,1822,1848]"pageId="12"pageNumber="13"pagination="215 - 220"refId="ref74229"refString="Genaro J. A. 1999. Revision del genero Triepeolus en Cuba (Hymenoptera: Apidae), con descripcion de dos especies nuevas. Caribbean Journal of Science 35: 215 - 220."type="journal article"year="1999">Genaro, 1999</bibRefCitation>
<taxonomicNameid="4C26998AFFF9FFA436F8F8ABFAFFF8B3"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[1212,1399,1858,1884]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="12"pageNumber="13"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
<taxonomicNameid="4C26998AFFF9FFA43166F88FFC55F890"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[802,989,1893,1919]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="12"pageNumber="13"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFF9FFA433CCF861FDAAF84D"author="Rozen J. G. Jr"box="[392,546,1928,1954]"pageId="12"pageNumber="13"pagination="1 - 18"refId="ref76133"refString="Rozen J. G. Jr 1989. Two new species and the redescription of another species of the cleptoparasitic bee genus Triepeolus with notes on their immature stages (Anthophoridae: Nomadinae). American Museum Novitates 2956: 1 - 18. Available from https: // www. biodiversitylibrary. org / page / 62309460 [accessed 25 Oct. 2023]."type="journal article"year="1989">Rozen, 1989</bibRefCitation>
(BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band.
<figureCitationid="131DFE8CFFF8FFA532F9FD30FEC5FD1C"box="[189,333,729,755]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">
<bibRefCitationid="EFB79FF8FFF8FFA53079FD15FD60FCF9"author="Cresson E. T."box="[573,744,764,790]"pageId="13"pageNumber="14"pagination="61 - 136"refId="ref73927"refString="Cresson E. T. 1878. Descriptions of new North American Hymenoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society 7: 61 - 136. https: // doi. org / 10.2307 / 25076368"type="journal article"year="1878">Cresson, 1878</bibRefCitation>
– Not as above. T1 basal and apical transverse bands (if present, distinct, and joined laterally) with abrupt transition between basal band and lateral longitudinal band (
<figureCitationid="131DFE8CFFF8FFA531B7FC80FBF4FC6C"box="[1011,1148,873,899]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11D–E</figureCitation>
) (at least if transverse bands not effectively mirror images, as in
<figureCitationid="131DFE8CFFF8FFA530BCFC65FC0FFC49"box="[760,903,908,934]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F–H</figureCitation>
), such that discal patch quadrangular or transversely oblong or elliptic (
<figureCitationid="131DFE8CFFF8FFA5301EFC46FD6FFC25"box="[602,743,943,970]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11D–H</figureCitation>
6. Any of the following (alone or in combination): T1 discal patch forming trapezoid or triangle with concave or straight anterolateral sides (basal/lateral band (or patch) of off-white to yellow tomentum on each side (or each) mesally convex or straight) (
<figureCitationid="131DFE8CFFF8FFA53126FBDEFC58FBBE"box="[866,976,1079,1105]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11B</figureCitation>
); T1–T4 with medially interrupted apical transverse bands (
<figureCitationid="131DFE8CFFF8FFA5304EFBB3FDFDFB9A"box="[522,629,1114,1141]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11B</figureCitation>
); and/or S3 with long, curved subapical setae (
<figureCitationid="131DFE8CFFF8FFA536DDFBB3FA8AFB9A"box="[1177,1282,1114,1141]"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="13"pageNumber="14">Fig. 10C</figureCitation>
) ............ ............................................................................................ other
– The following in combination: T1 discal patch semicircular or crescent-shaped (basal/lateral band (or patch) of yellow tomentum on each side (or each) mesally concave) (
<figureCitationid="131DFE8CFFF8FFA53652FB2DFB08FB31"box="[1046,1152,1220,1246]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11C</figureCitation>
); T3–T4 and usually also T2 with complete apical transverse bands (
<figureCitationid="131DFE8CFFF8FFA53155FB0EFCF2FAEE"box="[785,890,1255,1281]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11C</figureCitation>
); and S3 with short, straight subapical setae (
<figureCitationid="131DFE8CFFF8FFA532B6FAE3FED5FACA"box="[242,349,1290,1317]"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="13"pageNumber="14">Fig. 10D</figureCitation>
7. The following in combination: legs to some extent orange (
<figureCitationid="131DFE8CFFF8FFA531F0FAA5FB9EFA89"box="[948,1046,1356,1382]"captionStart="Fig"captionStartId="7.[189,232,1928,1954]"captionTargetBox="[218,1370,1453,1877]"captionTargetId="figure-212@7.[218,1370,1453,1892]"captionTargetPageId="7"captionText="Fig. 3. Head of male, frontal view. A. Triepeolus michiganensis Mitchell, 1962 (CNC 719805), which has a strong glabrous clypeal midline (blue arrow). B. T. obliteratus Graenicher, 1911 (WRME WRME_ EJH02370), which does not have a glabrous clypeal midline."figureDoi="http://doi.org/10.5281/zenodo.13691811"httpUri="https://zenodo.org/record/13691811/files/figure.png"pageId="13"pageNumber="14">Figs 3B</figureCitation>
,
<figureCitationid="131DFE8CFFF8FFA53660FAA5FBC1FA89"box="[1060,1097,1356,1382]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="13"pageNumber="14">6B</figureCitation>
<figureCitationid="131DFE8CFFF8FFA53613FAA5FB28FA89"box="[1111,1184,1356,1382]"captionStart="Fig"captionStartId="10.[189,232,1895,1921]"captionTargetBox="[218,1370,952,1817]"captionTargetId="figure-63@10.[218,1370,952,1832]"captionTargetPageId="10"captionText="Fig. 9. Right pro- and mesotrochanters of male (anterior facing right). A. Triepeolus tuberculifer Onuferko, Rightmyer & Roig-Alsina, 2024, allotype (FSCA M.G.R. Database No. 3704), showing distinct protuberances (blue arrows). B. T. obliteratus Graenicher, 1911 (WRME WRME_EJH02370). C. T. oblongimacula sp. nov., allotype (BOLD sample ID: CMNTO_298; PCYU PCYU-SD09:28). D. T. rhododontus Cockerell, 1921 (BOLD sample ID: CMNTO_139; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13742575"httpUri="https://zenodo.org/record/13742575/files/figure.png"pageId="13"pageNumber="14">9B–D</figureCitation>
<figureCitationid="131DFE8CFFF8FFA536EAFAA5FB68FA89"box="[1198,1248,1356,1382]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">11E</figureCitation>
); axilla with tip acute and distinctly pointed (
<figureCitationid="131DFE8CFFF8FFA5302FFA86FD40FA65"box="[619,712,1391,1418]"captionStart="Fig"captionStartId="8.[189,232,1928,1954]"captionTargetBox="[218,1370,1018,1882]"captionTargetId="figure-78@8.[218,1370,1018,1887]"captionTargetPageId="8"captionText="Fig. 5. Axillae and mesoscutellum of male, dorsal view. A. Triepeolus intrepidus (Smith, 1879) (PCYU PCYU-SD09:612). B. T. roni Genaro, 1999 (USNM M.G.R. Database No. 5922). C. T. hirsutus sp. nov., allotype (PCYU). D. T. brittaini Cockerell, 1931 (BOLD sample ID: 05-NS-0718; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13691815"httpUri="https://zenodo.org/record/13691815/files/figure.png"pageId="13"pageNumber="14">Fig. 5C</figureCitation>
); T1 basal band distinctly interrupted medially or notched posteromedially (
<figureCitationid="131DFE8CFFF8FFA533FFFA7BFDAEFA42"box="[443,550,1426,1453]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11E</figureCitation>
); T2 with complete or medially narrowly interrupted apical transverse band with pair of basomedially strongly convergent anterolateral extensions (except sometimes in
<bibRefCitationid="EFB79FF8FFF8FFA53383FA30FD0EFA1C"author="Cockerell T. D. A."box="[455,646,1497,1523]"pageId="13"pageNumber="14"pagination="1 - 16"refId="ref73439"refString="Cockerell T. D. A. 1921. The epeoline bees of the American Museum Rocky Mountain expeditions. American Museum Novitates 23: 1 - 16. Available from http: // hdl. handle. net / 2246 / 3249 [accessed 25 Oct. 2023]."type="journal article"year="1921">Cockerell, 1921</bibRefCitation>
</taxonomicName>
) (
<figureCitationid="131DFE8CFFF8FFA530E4FA30FC83FA1C"box="[672,779,1497,1523]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11E</figureCitation>
); and T3–T4 with complete apical transverse bands (
<figureCitationid="131DFE8CFFF8FFA532B1FA15FED3F9F9"box="[245,347,1532,1558]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11E</figureCitation>
– Any of the following (alone or in combination): legs entirely dark brown/black (
<figureCitationid="131DFE8CFFF8FFA536D1F9F6FAA9F9D6"box="[1173,1313,1567,1593]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F–G</figureCitation>
); axilla with tip right angled or obtuse and somewhat blunt if not broadly rounded (
<figureCitationid="131DFE8CFFF8FFA5361AF9ABFB34F9B2"box="[1118,1212,1602,1629]"captionStart="Fig"captionStartId="8.[189,232,1928,1954]"captionTargetBox="[218,1370,1018,1882]"captionTargetId="figure-78@8.[218,1370,1018,1887]"captionTargetPageId="8"captionText="Fig. 5. Axillae and mesoscutellum of male, dorsal view. A. Triepeolus intrepidus (Smith, 1879) (PCYU PCYU-SD09:612). B. T. roni Genaro, 1999 (USNM M.G.R. Database No. 5922). C. T. hirsutus sp. nov., allotype (PCYU). D. T. brittaini Cockerell, 1931 (BOLD sample ID: 05-NS-0718; PCYU)."figureDoi="http://doi.org/10.5281/zenodo.13691815"httpUri="https://zenodo.org/record/13691815/files/figure.png"pageId="13"pageNumber="14">Fig. 5D</figureCitation>
); T1 basal band (if present) not distinctly interrupted medially or notched posteromedially (
<figureCitationid="131DFE8CFFF8FFA5361DF98FFB4DF96F"box="[1113,1221,1638,1664]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11G</figureCitation>
<figureCitationid="131DFE8CFFF8FFA53106F960FC21F94C"box="[834,937,1673,1699]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F</figureCitation>
); and/or that of T2 (if present) without anterolateral extensions (
<figureCitationid="131DFE8CFFF8FFA53056F945FDF2F929"box="[530,634,1708,1734]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11H</figureCitation>
) or with pair of basomedially weakly convergent, perpendicular, or basolaterally divergent anterolateral extensions (
<figureCitationid="131DFE8CFFF8FFA53107F926FC44F906"box="[835,972,1743,1769]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F–G</figureCitation>
8. Any of the following (alone or in combination): legs entirely dark brown/black (
<figureCitationid="131DFE8CFFF8FFA5369CF8F8FAE1F8C4"box="[1240,1385,1809,1835]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F–G</figureCitation>
); mesopleuron densely punctate (i ≤1d) (
<figureCitationid="131DFE8CFFF8FFA53084F8DDFCA8F8A1"box="[704,800,1844,1870]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="13"pageNumber="14">Fig. 6C</figureCitation>
); T1 basal (
<figureCitationid="131DFE8CFFF8FFA531F5F8DDFB95F8A1"box="[945,1053,1844,1870]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11G</figureCitation>
) and/or apical (
<figureCitationid="131DFE8CFFF8FFA5369AF8DDFAE6F8A1"box="[1246,1390,1844,1870]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11G–H</figureCitation>
<figureCitationid="131DFE8CFFF8FFA532B7F893FED1F87B"box="[243,345,1914,1940]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F</figureCitation>
); and/or that of T2 with pair of well-defined anterolateral extensions (
<figureCitationid="131DFE8CFFF8FFA536C8F893FA9DF87B"box="[1164,1301,1914,1940]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="13"pageNumber="14">Fig. 11F–G</figureCitation>
) .......... ............................................................................................ other
– The following in combination: legs to some extent orange (
<figureCitationid="131DFE8CFFFBFFA631FDFEE0FB95FECC"box="[953,1053,265,291]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="14"pageNumber="15">Figs 6D</figureCitation>
,
<figureCitationid="131DFE8CFFFBFFA63668FEE0FBE9FECC"box="[1068,1121,265,291]"captionStart="Fig"captionStartId="11.[189,232,1787,1813]"captionTargetBox="[218,1370,871,1735]"captionTargetId="figure-183@11.[218,1370,871,1741]"captionTargetPageId="11"captionText="Fig. 10. Metasoma of male, lateral view. A. Triepeolus simplex Robertson, 1903 (BOLD sample ID: QUBS0291G11; PCYU) showing S4 with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrow). B. T. diffusus Rightmyer, 2008 (RAM pmae00146019) showing S4 with apical/ subapical fringe of dense, long (> 1 MOD), curved setae (blue arrow). C. T. rufoclypeus (Fox, 1891) (PCYU) showing S3–S5 each with apical/subapical fringe of dense, long (>1 MOD), curved setae (blue arrows). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271) showing S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved setae (blue arrows)."figureDoi="http://doi.org/10.5281/zenodo.13691825"httpUri="https://zenodo.org/record/13691825/files/figure.png"pageId="14"pageNumber="15">10D</figureCitation>
,
<figureCitationid="131DFE8CFFFBFFA6362BFEE0FB4FFECC"box="[1135,1223,265,291]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="14"pageNumber="15">11C–D</figureCitation>
<figureCitationid="131DFE8CFFFBFFA630F3FEC5FC94FEA9"box="[695,796,300,326]"captionStart="Fig"captionStartId="9.[189,232,1169,1195]"captionTargetBox="[218,1370,265,1128]"captionTargetId="figure-12@9.[218,1370,264,1132]"captionTargetPageId="9"captionText="Fig. 6. Mesosoma of male, lateral view. A. Triepeolus melanarius Rightmyer, 2008 (CSUC), which ventrolaterally has dense, long (≥½ MOD), erect/suberect, minutely branched setae (blue arrow). B. T. parahirsutus sp. nov., allotype (ZMB 21261), which ventrolaterally has sparse, long (≥½ MOD), erect/suberect, simple setae among the usual appressed, branched setae (blue arrow). C. T. joliae Rightmyer, 2008 (CNC 1801918). D. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271), which does not have dense, long (≥ ½ MOD), erect/suberect setae."figureDoi="http://doi.org/10.5281/zenodo.13691817"httpUri="https://zenodo.org/record/13691817/files/figure.png"pageId="14"pageNumber="15">Fig. 6D</figureCitation>
); T1 with medially interrupted basal and apical transverse bands (
<figureCitationid="131DFE8CFFFBFFA633FCFEA6FDAAFE86"box="[440,546,335,361]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="14"pageNumber="15">Fig. 11D</figureCitation>
); T2 with complete (
<figureCitationid="131DFE8CFFFBFFA63149FEA6FCFAFE86"box="[781,882,335,361]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="14"pageNumber="15">Fig. 11C</figureCitation>
) or medially narrowly interrupted (
<figureCitationid="131DFE8CFFFBFFA63743FEA6FAE5FE86"box="[1287,1389,335,361]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="14"pageNumber="15">Fig. 11D</figureCitation>
) apical transverse band without well-defined anterolateral extensions; T3–T4 with complete apical transverse bands (
<figureCitationid="131DFE8CFFFBFFA633F9FE7FFDC1FE5F"box="[445,585,406,432]"captionStart="Fig"captionStartId="12.[189,232,1822,1848]"captionTargetBox="[218,1370,471,1775]"captionTargetId="figure-56@12.[218,1370,471,1784]"captionText="Fig. 11A–F (continued on next page). Metasoma of male, dorsal view. A. Triepeolus roni Genaro, 1999 (USNM M.G.R. Database No. 5922). B. T. buchwaldi (Friese, 1908) (INHS 387483). C. T. eumeniformis sp. nov., paratype (ECOSUR ECO-TA-E-41271). D. T. eumeniformis sp. nov., paratype (CNC 1801914). E. T. kathrynae Rozen, 1989 (RSKM RSKM_ENT_E-168244). F. T. tristis (Smith, 1854) (BOLD sample ID: CCDB-30345 C09; PCYU). Blue lines indicate an abrupt (A, D–E) or seamless (B–C) transition between the basal band and lateral longitudinal band."figureDoi="http://doi.org/10.5281/zenodo.13737904"httpUri="https://zenodo.org/record/13737904/files/figure.png"pageId="14"pageNumber="15">Fig. 11C–D</figureCitation>
<figureCitationid="131DFE8CFFFBFFA6363AFDEFFB63FDCF"box="[1150,1259,518,544]"captionStart="Fig"captionStartId="14.[189,232,1892,1918]"captionTargetBox="[189,1398,620,1829]"captionTargetId="figure-102@14.[189,1398,620,1846]"captionText="Fig. 12A–D (continued on next page). Distribution maps. A. All species in the Triepeolus simplex group. B. T. apache sp. nov. C. T. eumeniformis sp. nov. D. T. hirsutus sp. nov. Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens)."figureDoi="http://doi.org/10.5281/zenodo.13737908"httpUri="https://zenodo.org/record/13737908/files/figure.png"pageId="14"pageNumber="15">Fig. 12A</figureCitation>
<taxonomicNameid="4C26998AFFFBFFA63375F860FE2AF84D"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[305,418,1928,1954]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="apache"status="sp. nov.">
<taxonomicNameid="4C26998AFFFBFFA63068F860FD61F84D"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[556,745,1928,1954]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
<taxonomicNameid="4C26998AFFFBFFA63137F860FC66F84D"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[883,1006,1928,1954]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="hirsutus"status="sp. nov.">
Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens).
<taxonomicNameid="4C26998AFFFAFFA732F9FEB3FE1DFE9B"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,405,346,372]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="apache"status="sp. nov.">
<taxonomicNameid="4C26998AFFFAFFA732F9FE94FE68FE78"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,480,381,407]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="eumeniformis"status="sp. nov.">
<taxonomicNameid="4C26998AFFFAFFA732F9FE48FE17FE54"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,415,417,443]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="hirsutus"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFFAFFA733F9FE2DFDD9FE31"author="Rozen J. G. Jr"box="[445,593,452,478]"pageId="15"pageNumber="16"pagination="1 - 18"refId="ref76133"refString="Rozen J. G. Jr 1989. Two new species and the redescription of another species of the cleptoparasitic bee genus Triepeolus with notes on their immature stages (Anthophoridae: Nomadinae). American Museum Novitates 2956: 1 - 18. Available from https: // www. biodiversitylibrary. org / page / 62309460 [accessed 25 Oct. 2023]."type="journal article"year="1989">Rozen, 1989</bibRefCitation>
<bibRefCitationid="EFB79FF8FFFAFFA7338FFE0EFDFEFDEE"author="Cresson E. T."box="[459,630,487,513]"pageId="15"pageNumber="16"pagination="61 - 136"refId="ref73927"refString="Cresson E. T. 1878. Descriptions of new North American Hymenoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society 7: 61 - 136. https: // doi. org / 10.2307 / 25076368"type="journal article"year="1878">Cresson, 1878</bibRefCitation>
<figureCitationid="131DFE8CFFFAFFA732F9F8A8FEC4F8B4"box="[189,332,1857,1883]"captionStart="Fig"captionStartId="14.[189,232,1892,1918]"captionTargetBox="[189,1398,620,1829]"captionTargetId="figure-102@14.[189,1398,620,1846]"captionText="Fig. 12A–D (continued on next page). Distribution maps. A. All species in the Triepeolus simplex group. B. T. apache sp. nov. C. T. eumeniformis sp. nov. D. T. hirsutus sp. nov. Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens)."figureDoi="http://doi.org/10.5281/zenodo.13737908"httpUri="https://zenodo.org/record/13737908/files/figure.png"pageId="15"pageNumber="16">
<bibRefCitationid="EFB79FF8FFFAFFA732F9F88CFEDCF890"author="Rozen J. G. Jr"box="[189,340,1893,1919]"pageId="15"pageNumber="16"pagination="1 - 18"refId="ref76133"refString="Rozen J. G. Jr 1989. Two new species and the redescription of another species of the cleptoparasitic bee genus Triepeolus with notes on their immature stages (Anthophoridae: Nomadinae). American Museum Novitates 2956: 1 - 18. Available from https: // www. biodiversitylibrary. org / page / 62309460 [accessed 25 Oct. 2023]."type="journal article"year="1989">Rozen, 1989</bibRefCitation>
<bibRefCitationid="EFB79FF8FFFAFFA73062F88CFD5BF890"author="Cresson E. T."box="[550,723,1893,1919]"pageId="15"pageNumber="16"pagination="61 - 136"refId="ref73927"refString="Cresson E. T. 1878. Descriptions of new North American Hymenoptera in the collection of the American Entomological Society. Transactions of the American Entomological Society 7: 61 - 136. https: // doi. org / 10.2307 / 25076368"type="journal article"year="1878">Cresson, 1878</bibRefCitation>
<bibRefCitationid="EFB79FF8FFFAFFA731EAF88CFBF3F890"author="Graenicher S."box="[942,1147,1893,1919]"pageId="15"pageNumber="16"pagination="221 - 249"refId="ref74330"refString="Graenicher S. 1911. Bees of northwestern Wisconsin. Bulletin of the Public Museum of the City of Milwaukee 1: 221 - 249."type="journal article"year="1911">Graenicher, 1911</bibRefCitation>
<taxonomicNameid="4C26998AFFFAFFA736EEF88FFAFFF890"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[1194,1399,1893,1919]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="15"pageNumber="16"phylum="Arthropoda"rank="species"species="oblongimacula"status="sp. nov.">
Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens).
<bibRefCitationid="EFB79FF8FFE5FFB83381FEDEFD00FEBE"author="Holmberg E. L."box="[453,648,311,337]"pageId="16"pageNumber="17"pagination="272 - 286"refId="ref74749"refString="Holmberg E. L. 1886. Sobre apidos nomadas de la Republica Argentina. Anales de la Sociedad Cientifica Argentina 22: 272 - 286."type="journal article"year="1886">Holmberg, 1886</bibRefCitation>
<bibRefCitationid="EFB79FF8FFE5FFB83383FEB2FD19FE9A"author="Graenicher S."box="[455,657,347,373]"pageId="16"pageNumber="17"pagination="221 - 249"refId="ref74330"refString="Graenicher S. 1911. Bees of northwestern Wisconsin. Bulletin of the Public Museum of the City of Milwaukee 1: 221 - 249."type="journal article"year="1911">Graenicher, 1911</bibRefCitation>
<taxonomicNameid="4C26998AFFE5FFB832F9FE97FE78FE77"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,496,382,408]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="oblongimacula"status="sp. nov.">
<taxonomicNameid="4C26998AFFE5FFB832F9FE48FE5EFE54"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,470,417,443]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="parahirsutus"status="sp. nov.">
<taxonomicNameid="4C26998AFFE5FFB832F9FE2DFE7BFE31"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,499,452,478]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="paucipunctatus"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFE5FFB8339FFE0EFD1FFDED"author="Cockerell T. D. A."box="[475,663,487,514]"pageId="16"pageNumber="17"pagination="1 - 16"refId="ref73439"refString="Cockerell T. D. A. 1921. The epeoline bees of the American Museum Rocky Mountain expeditions. American Museum Novitates 23: 1 - 16. Available from http: // hdl. handle. net / 2246 / 3249 [accessed 25 Oct. 2023]."type="journal article"year="1921">Cockerell, 1921</bibRefCitation>
<figureCitationid="131DFE8CFFE5FFB832F9F8A8FEB6F8B4"box="[189,318,1857,1883]"captionStart="Fig"captionStartId="14.[189,232,1892,1918]"captionTargetBox="[189,1398,620,1829]"captionTargetId="figure-102@14.[189,1398,620,1846]"captionText="Fig. 12A–D (continued on next page). Distribution maps. A. All species in the Triepeolus simplex group. B. T. apache sp. nov. C. T. eumeniformis sp. nov. D. T. hirsutus sp. nov. Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens)."figureDoi="http://doi.org/10.5281/zenodo.13737908"httpUri="https://zenodo.org/record/13737908/files/figure.png"pageId="16"pageNumber="17">
<taxonomicNameid="4C26998AFFE5FFB8361BF8A8FAFFF8B4"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[1119,1399,1857,1883]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="parahirsutus"status="sp. nov.">
<taxonomicNameid="4C26998AFFE5FFB8330CF88FFD93F890"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[328,539,1894,1919]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="paucipunctatus"status="sp. nov.">
<bibRefCitationid="EFB79FF8FFE5FFB8313DF88CFBB3F890"author="Cockerell T. D. A."box="[889,1083,1893,1919]"pageId="16"pageNumber="17"pagination="1 - 16"refId="ref73439"refString="Cockerell T. D. A. 1921. The epeoline bees of the American Museum Rocky Mountain expeditions. American Museum Novitates 23: 1 - 16. Available from http: // hdl. handle. net / 2246 / 3249 [accessed 25 Oct. 2023]."type="journal article"year="1921">Cockerell, 1921</bibRefCitation>
<bibRefCitationid="EFB79FF8FFE5FFB83688F88CFAFBF890"author="Genaro J. A."box="[1228,1395,1893,1919]"pageId="16"pageNumber="17"pagination="215 - 220"refId="ref74229"refString="Genaro J. A. 1999. Revision del genero Triepeolus en Cuba (Hymenoptera: Apidae), con descripcion de dos especies nuevas. Caribbean Journal of Science 35: 215 - 220."type="journal article"year="1999">Genaro, 1999</bibRefCitation>
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. Approximate geographic ranges are indicated in orange and were estimated from occurrence records known to the authors (yellow and blue circles, with the latter indicating localities of DNA barcoded specimens).
<bibRefCitationid="EFB79FF8FFE4FFB93333FED1FD9FFEBD"author="Genaro J. A."box="[375,535,312,338]"pageId="17"pageNumber="18"pagination="215 - 220"refId="ref74229"refString="Genaro J. A. 1999. Revision del genero Triepeolus en Cuba (Hymenoptera: Apidae), con descripcion de dos especies nuevas. Caribbean Journal of Science 35: 215 - 220."type="journal article"year="1999">Genaro, 1999</bibRefCitation>
<taxonomicNameid="4C26998AFFE4FFB932F9FEB2FCACFE9A"authority="Cockerell & Sandhouse, 1924"authorityName="Cockerell & Sandhouse"authorityYear="1924"box="[189,804,347,373]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="17"pageNumber="18"phylum="Arthropoda"rank="species"species="saturninus">
<bibRefCitationid="EFB79FF8FFE4FFB93387FEB2FCACFE9A"author="Cockerell T. D. A. & Sandhouse G."box="[451,804,347,373]"pageId="17"pageNumber="18"pagination="305 - 324"refId="ref73761"refString="Cockerell T. D. A. & Sandhouse G. 1924. Parasitic bees (Epeolinae and Melectinae) in the collection of the California Academy of Sciences. Proceedings of the California Academy of Sciences, Fourth Series 13: 305 - 324. Available from https: // www. biodiversitylibrary. org / page / 3137831 [accessed 25 Oct. 2023]."type="book chapter"year="1924">Cockerell & Sandhouse, 1924</bibRefCitation>
<bibRefCitationid="EFB79FF8FFE4FFB93389FE97FD04FE77"author="Cockerell T. D. A."box="[461,652,382,408]"pageId="17"pageNumber="18"pagination="361 - 364"refId="ref73278"refString="Cockerell T. D. A. 1900. Notes on New Mexico bees. The Canadian Entomologist 32: 361 - 364. https: // doi. org / 10.4039 / Ent 32361 - 12"type="journal article"year="1900">Cockerell, 1900</bibRefCitation>
<taxonomicNameid="4C26998AFFE4FFB932F9FE48FE25FE54"authorityName="Onuferko & Rightmyer"authorityYear="2024"box="[189,429,417,443]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="17"pageNumber="18"phylum="Arthropoda"rank="species"species="shoshone"status="sp. nov.">
<taxonomicNameid="4C26998AFFE4FFB932F9FE0EFC44FDED"authority="Onuferko, Rightmyer & Roig-Alsina, 2024"authorityName="Onuferko, Rightmyer & Roig-Alsina"authorityYear="2024"box="[189,972,487,514]"class="Insecta"family="Apidae"genus="Triepeolus"kingdom="Animalia"order="Hymenoptera"pageId="17"pageNumber="18"phylum="Arthropoda"rank="species"species="tuberculifer">
<bibRefCitationid="EFB79FF8FFE4FFB93395FE0EFC44FDED"author="Onuferko T. M. & Rightmyer M. G. & Melo G. A. R. & Roig-Alsina A."box="[465,972,487,514]"pageId="17"pageNumber="18"pagination="1 - 50"refId="ref75439"refString="Onuferko T. M., Rightmyer M. G., Melo G. A. R. & Roig-Alsina A. 2024. A revision of the South American species of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae). European Journal of Taxonomy 931: 1 - 50. https: // doi. org / 10.5852 / ejt. 2024.931.2505"type="journal article"year="2024">Onuferko, Rightmyer & Roig-Alsina, 2024</bibRefCitation>
