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114 lines
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<document id="4488CEC960C7E75D61A2D5543959EF59" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFA1FFC67EFCFD8FF08ADC7D" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis obscuripennis Dominguez, New combination" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="25" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="6015D84D973A88F8FA337711154EF42F">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:namePart id="8BDA4A8A11792A4F6D4919DE3089B21D">Molineri, Carlos</mods:namePart>
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<mods:typeOfResource id="C550B4C92B0571ECB42C61AB7382B912">text</mods:typeOfResource>
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<mods:title id="68F5C73C341272A7A8E78E3DAEFBDAF8">Zootaxa</mods:title>
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<mods:part id="D63A39876B2A761759DE3908BA59649F">
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<mods:date id="70CD0528C7D7CB8DFB2D0ECAECABEEBA">2010</mods:date>
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<mods:detail id="4AEA17BBB1F4C8CDED9E476A718428DD" type="volume">
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<mods:number id="328DB57DC46BEFC3F4DD35EC9FEFE877">2481</mods:number>
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<mods:end id="F6496B0640C79401F4F69C94E2B62B50">36</mods:end>
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<mods:classification id="E8B6521E0ABC53FAE7A1C20C25591B0B">journal article</mods:classification>
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<mods:identifier id="F43C0FE5E991684274FD2D07ACDDB031" type="DOI">10.5281/zenodo.195431</mods:identifier>
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<mods:identifier id="8B6A5854BFB8E0EDE59F9CCF3917C56F" type="GBIF-Dataset">2df8bd5a-e125-42aa-aa6f-ae3e78b83929</mods:identifier>
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<mods:identifier id="7B378759642950B53CEB7FC5FACEC203" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="464B39732C7731599FC2480ECADE8525" type="Zenodo-Dep">195431</mods:identifier>
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<treatment id="03F39207FFA1FFC67EFCFD8FF08ADC7D" ID-DOI="http://doi.org/10.5281/zenodo.5690583" ID-GBIF-Taxon="119386672" ID-Zenodo-Dep="5690583" LSID="urn:lsid:plazi:treatment:03F39207FFA1FFC67EFCFD8FF08ADC7D" httpUri="http://treatment.plazi.org/id/03F39207FFA1FFC67EFCFD8FF08ADC7D" lastPageNumber="25" pageId="24" pageNumber="25">
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<subSubSection id="C340709AFFA1FFC67EFCFD8FF64BDB26" box="[151,832,525,551]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BE52311FFA1FFC67EFCFD8FF64BDB26" blockId="24.[151,832,525,551]" box="[151,832,525,551]" pageId="24" pageNumber="25">
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<heading id="D0AD947DFFA1FFC67EFCFD8FF64BDB26" bold="true" box="[151,832,525,551]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<emphasis id="B92EFF03FFA1FFC67EFCFD8FF64BDB26" bold="true" box="[151,832,525,551]" pageId="24" pageNumber="25">
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<taxonomicName id="4C5A5892FFA1FFC67EFCFD8FF755DB26" ID-CoL="7CJ8C" authority="Dominguez" authorityName="Dominguez" box="[151,606,525,551]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="obscuripennis" status="comb. nov.">
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<emphasis id="B92EFF03FFA1FFC67EFCFD8FF4BEDB26" bold="true" box="[151,437,525,551]" italics="true" pageId="24" pageNumber="25">Tortopsis obscuripennis</emphasis>
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(Domínguez)
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFA1FFC67C0EFD8FF64BDB26" box="[613,832,525,551]" pageId="24" pageNumber="25">New combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFA1FFC67EFCFDD7F65BDB6D" box="[151,848,597,620]" pageId="24" pageNumber="25" type="reference_group">
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<paragraph id="8BE52311FFA1FFC67EFCFDD7F65BDB6D" blockId="24.[151,848,597,620]" box="[151,848,597,620]" pageId="24" pageNumber="25">
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<treatmentCitationGroup id="AB4A043FFFA1FFC67EFCFDD7F65BDB6D" box="[151,848,597,620]" pageId="24" pageNumber="25">
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<taxonomicName id="4C5A5892FFA1FFC67EFCFDD7F770DB6D" authority="Dominguez 1985: 71" authorityName="Dominguez" authorityPageNumber="71" authorityYear="1985" box="[151,635,597,620]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="obscuripennis">
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<emphasis id="B92EFF03FFA1FFC67EFCFDD7F49ADB6D" box="[151,401,597,620]" italics="true" pageId="24" pageNumber="25">Tortopus obscuripennis</emphasis>
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<treatmentCitation id="0AFB0500FFA1FFC67FFCFDD7F770DB6D" author="Dominguez" box="[407,635,597,620]" page="71" pageId="24" pageNumber="25" year="1985">
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<bibRefCitation id="EFCB5EE0FFA1FFC67FFCFDD7F75FDB6D" author="Dominguez" box="[407,596,597,620]" pageId="24" pageNumber="34" refString="Dominguez, E. (1985) El genero Tortopus Needham y Murphy (Ephemeroptera: Polymitarcyidae) en la Argentina. Physis, B 43, 69 - 72." type="journal article" year="1985">Domínguez 1985</bibRefCitation>
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: 71
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AFB0500FFA1FFC67CECFDD7F647DB6D" author="Molineri" box="[647,844,597,620]" page="10" pageId="24" pageNumber="25" year="2008">
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<bibRefCitation id="EFCB5EE0FFA1FFC67CECFDD7F62FDB6D" author="Molineri" box="[647,804,597,620]" pageId="24" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
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: 10
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFA1FFC67EFCFD18F08ADC7D" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="8BE52311FFA1FFC67EFCFD18F09EDBDB" blockId="24.[151,1437,666,1404]" pageId="24" pageNumber="25">
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Material (
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<collectionCode id="ED4BBBD4FFA1FFC67F61FD18F44DDBB5" box="[266,326,666,692]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="24" pageNumber="25">IML</collectionCode>
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):
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<typeStatus id="54E19DB3FFA1FFC67F31FD18F4CBDBB5" box="[346,448,666,692]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
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and 18
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<typeStatus id="54E19DB3FFA1FFC67C74FD18F784DBB5" box="[543,655,666,692]" pageId="24" pageNumber="25" type="paratype">paratypes</typeStatus>
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female imagos from
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<collectingCountry id="F34D6381FFA1FFC67DE0FD18F10FDBB5" box="[907,1028,666,692]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
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: Salta, Aguas Blancas, Estancia El Arrayazal,
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<date id="FFE405D1FFA1FFC67F72FD42F4AADBDB" box="[281,417,704,730]" pageId="24" pageNumber="25" value="1981-12-09">9-XII-1981</date>
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, Domínguez col. Other material from
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<collectingCountry id="F34D6381FFA1FFC67D0AFD42F6DEDBDB" box="[865,981,704,730]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
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and
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<collectingCountry id="F34D6381FFA1FFC67A64FD42F16EDBDB" box="[1039,1125,704,730]" name="Bolivia" pageId="24" pageNumber="25">Bolivia</collectingCountry>
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listed in
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<bibRefCitation id="EFCB5EE0FFA1FFC67ABAFD42F09ADBDB" author="Molineri" box="[1233,1425,704,730]" pageId="24" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri (2008)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE52311FFA1FFC67EADFD67F646DD06" blockId="24.[151,1437,666,1404]" pageId="24" pageNumber="25">
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Female imago. Head black between ocelli, with gray anastomosed marks on occiput (some times very slightly marked). Pronotum with anterior ring widely shaded black, and posterior ring shaded grayish black on central area, lateral to this shading there is an isolated narrow gray mark, and also shaded near anterolateral corner. Wings with very dark veins; vein R3 and short intercalary before it absent in fore wings but some females present a weak vein in zig-zag formed by coalescence of cross veins, with a length of 2 cells. Abdominal terga with thin black medial line, surrounded by a paler grayish band, and more laterally with wide submedian dark bands. Sternum VIII with relatively small C-shaped sockets, with indistinct margin (not sclerotized or pigmented), depression subcircular (
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<figureCitation id="13613F94FFA1FFC67C8CFC6FF634DD06" box="[743,831,1005,1031]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="24" pageNumber="25">Fig. 48</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE52311FFA1FFC67EADFB90F74ADD53" blockId="24.[151,1437,666,1404]" pageId="24" pageNumber="25">Egg. Suboval, no attachment structures. Length, 390–410 µm; width, 340–350. The chorionic cover is sculptured with a hexagonal pattern.</paragraph>
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<paragraph id="8BE52311FFA1FFC67EADFBDFF08ADC7D" blockId="24.[151,1437,666,1404]" pageId="24" pageNumber="25">
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Discussion and diagnosis. This species, originally described from female adults (
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<bibRefCitation id="EFCB5EE0FFA1FFC67AC9FBDFF07EDD76" author="Dominguez" box="[1186,1397,1117,1143]" pageId="24" pageNumber="34" refString="Dominguez, E. (1985) El genero Tortopus Needham y Murphy (Ephemeroptera: Polymitarcyidae) en la Argentina. Physis, B 43, 69 - 72." type="journal article" year="1985">Domínguez 1985</bibRefCitation>
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) is currently known from all the stages (
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<bibRefCitation id="EFCB5EE0FFA1FFC67C35FB00F618DD9D" author="Molineri" box="[606,787,1154,1180]" pageId="24" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
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) and has been recorded from
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<collectingCountry id="F34D6381FFA1FFC67AE8FB00F1F6DD9D" box="[1155,1277,1154,1180]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
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and
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<collectingCountry id="F34D6381FFA1FFC67B51FB00F092DD9D" box="[1338,1433,1154,1180]" name="Bolivia" pageId="24" pageNumber="25">Bolivia</collectingCountry>
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.
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<taxonomicName id="4C5A5892FFA1FFC67EFCFB2AF4A1DDC0" box="[151,426,1192,1217]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="obscuripennis">
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<emphasis id="B92EFF03FFA1FFC67EFCFB2AF4A1DDC0" box="[151,426,1192,1217]" italics="true" pageId="24" pageNumber="25">Tortopus obscuripennis</emphasis>
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</taxonomicName>
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can be separated from all other species of the genus by: 1) fore wing length 15.0 mm (male),
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<quantity id="4CA28EF4FFA1FFC67E9EFB4FF4A9DDE6" box="[245,418,1229,1255]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="24" pageNumber="25" unit="mm" value="20.0" valueMax="20.5" valueMin="19.5">19.5–20.5 mm</quantity>
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(female); 2) relatively dark wings (membrane translucent), all veins shaded gray; 3) parastyli long and moderately curved; 4) penes abruptly widening in distal half; 5) female parastylus receptors C–shaped (
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<figureCitation id="13613F94FFA1FFC67F76FA9AF47DDC33" box="[285,374,1304,1330]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="24" pageNumber="25">Fig. 48</figureCitation>
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); 6) head shaded black among ocelli, occiput with a finely reticulated grayish pattern; 7) female pronotum with anterior ring widely shaded black, and posterior ring shaded grayish black on central area, lateral to this shading there is an isolated narrow gray mark, and also shaded near anterolateral corner.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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