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<document id="792E78831A6FC4B731071164BD7208AF" ID-DOI="10.5281/zenodo.186202" ID-GBIF-Dataset="0b071cfc-13c8-49bd-95c4-b3d0c6d77657" ID-ISSN="1175-5326" ID-Zenodo-Dep="186202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047799636" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2009" docId="8D5C1307FF8DDF0FAAF99BFDFE6CA4AB" docLanguage="en" docName="zt02027p054.pdf" docOrigin="Zootaxa 2027" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crotonia ovata Olszanowski 2000" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="71656B7FFF8BDF06AA6E9F38FFABA62F" masterDocTitle="New species of Crotonia (Acari: Oribatida) from Tasmania Rainforest, and the habitat preferences of Crotoniidae" masterLastPageNumber="54" masterPageNumber="43" pageNumber="49" updateTime="1698577644603" updateUser="plazi">
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<mods:title id="3A4153788A24560A08E1AE7AF7EE1E07">New species of Crotonia (Acari: Oribatida) from Tasmania Rainforest, and the habitat preferences of Crotoniidae</mods:title>
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<mods:namePart id="C2AD219EC9DBE83DA1E7DE22B4817B81">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="12F0E18A3744D3A7A11108ACFDD9DA0E">text</mods:typeOfResource>
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<mods:date id="001E2D18E5AC30177E0DDFADBAA9F237">2009</mods:date>
<mods:detail id="1E81D735B5E3AA3CC6DCB8EE1096C2EF" type="volume">
<mods:number id="B8000AFDCBE98E748FFD543BCD603552">2027</mods:number>
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<mods:classification id="D485544ED9A633240BBBDCB8DDB34B04">journal article</mods:classification>
<mods:identifier id="66D70202A635FEAC6F2887F260443673" type="DOI">10.5281/zenodo.186202</mods:identifier>
<mods:identifier id="F7666223637186D129BB56A22756D2CE" type="GBIF-Dataset">0b071cfc-13c8-49bd-95c4-b3d0c6d77657</mods:identifier>
<mods:identifier id="C239023A2D6AD20CB4E51DEC924941F2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CA9FFB8394ABDB4FE33B03B2AD514245" type="Zenodo-Dep">186202</mods:identifier>
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<treatment id="8D5C1307FF8DDF0FAAF99BFDFE6CA4AB" ID-DOI="http://doi.org/10.5281/zenodo.6221932" ID-GBIF-Taxon="119611213" ID-Zenodo-Dep="6221932" LSID="urn:lsid:plazi:treatment:8D5C1307FF8DDF0FAAF99BFDFE6CA4AB" httpUri="http://treatment.plazi.org/id/8D5C1307FF8DDF0FAAF99BFDFE6CA4AB" lastPageId="9" lastPageNumber="52" pageId="6" pageNumber="49">
<subSubSection id="4DEFF19AFF8DDF00AAF99BFDFEB3A32D" pageId="6" pageNumber="49" type="nomenclature">
<paragraph id="054AA211FF8DDF00AAF99BFDFDEBA2F0" blockId="6.[151,576,1221,1282]" box="[151,576,1221,1247]" pageId="6" pageNumber="49">
<heading id="5E02157DFF8DDF00AAF99BFDFDEBA2F0" bold="true" box="[151,576,1221,1247]" fontSize="11" level="1" pageId="6" pageNumber="49" reason="1">
<emphasis id="37817E03FF8DDF00AAF99BFDFDEBA2F0" bold="true" box="[151,576,1221,1247]" pageId="6" pageNumber="49">
<taxonomicName id="C2F5D992FF8DDF00AAF99BFDFD90A2F0" ID-CoL="ZRHG" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[151,571,1221,1247]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF8DDF00AAF99BFDFEE0A2F0" bold="true" box="[151,331,1221,1247]" italics="true" pageId="6" pageNumber="49">Crotonia ovata</emphasis>
<bibRefCitation id="6164DFE0FF8DDF00AB3C9BFDFD90A2F0" author="Olszanowski" box="[338,571,1221,1247]" pageId="6" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski, 2000</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="054AA211FF8DDF00AAF99BD0FEB3A32D" blockId="6.[151,576,1221,1282]" box="[151,280,1256,1282]" pageId="6" pageNumber="49">
(
<figureCitation id="9DCEBE94FF8DDF00AAF19BD0FF59A32D" box="[159,242,1256,1282]" captionStart="FIGURE 3" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[185,1402,482,1711]" captionTargetId="figure@7.[780,1416,495,1703]" captionTargetPageId="7" captionText="FIGURE 3. Crotonia ovata Olszanowski, 2000, male a) dorsal; b) ventral." httpUri="https://zenodo.org/record/186205/files/figure.png" pageId="6" pageNumber="49">Figs. 3</figureCitation>
,
<figureCitation id="9DCEBE94FF8DDF00AA919BD0FEA4A32D" box="[255,271,1256,1282]" captionStart="FIGURE 4" captionStartId="8.[151,255,1533,1557]" captionTargetBox="[248,1314,643,1457]" captionTargetId="figure@8.[235,1320,640,1113]" captionTargetPageId="8" captionText="FIGURE 4. Crotonia ovata Olszanowski, 2000, a) male, lateral view; b) caudal region of a female specimen from Mt. Victoria showing artefactual alteration in morphology (cf. text for details)." httpUri="https://zenodo.org/record/186206/files/figure.png" pageId="6" pageNumber="49">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4DEFF19AFF8DDF00AAF99A0DFD35A343" pageId="6" pageNumber="49" type="reference_group">
<paragraph id="054AA211FF8DDF00AAF99A0DFDC8A363" blockId="6.[151,670,1333,1388]" box="[151,611,1333,1356]" pageId="6" pageNumber="49">
<taxonomicName id="C2F5D992FF8DDF00AAF99A0DFDA4A363" ID-CoL="ZRHG" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[151,527,1333,1356]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF8DDF00AAF99A0DFE93A363" box="[151,312,1333,1356]" italics="true" pageId="6" pageNumber="49">Crotonia ovata</emphasis>
<bibRefCitation id="6164DFE0FF8DDF00AB519A0DFDA4A363" author="Olszanowski" box="[319,527,1333,1356]" pageId="6" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski, 2000</bibRefCitation>
</taxonomicName>
: p. 240.
</paragraph>
<paragraph id="054AA211FF8DDF00AAF99A6DFD35A343" blockId="6.[151,670,1333,1388]" box="[151,670,1365,1388]" pageId="6" pageNumber="49">
<taxonomicName id="C2F5D992FF8DDF00AAF99A6DFD96A343" authority="Lochynska, 2008" authorityName="Lochynska" authorityYear="2008" box="[151,573,1365,1388]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF8DDF00AAF99A6DFED1A343" box="[151,378,1365,1388]" italics="true" pageId="6" pageNumber="49">Holonothrus ryszardi</emphasis>
<bibRefCitation id="6164DFE0FF8DDF00ABEF9A6DFD96A343" author="Lochynska" box="[385,573,1365,1388]" pageId="6" pageNumber="54" refString="Lochynska, M. (2008 b) The ontogenetic description of two Tasmanian crotoniid mites (Acari: Oribatida: Crotoniidae). International Journal of Acarology, 34, 123 - 142." type="journal article" year="2008" yearSuffix="b">Łochyńska, 2008</bibRefCitation>
</taxonomicName>
b: p. 123.
</paragraph>
</subSubSection>
<subSubSection id="4DEFF19AFF8DDF01AAF99AA4FEF0A7B3" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="description">
<paragraph id="054AA211FF8DDF00AAF99AA4FF70A399" blockId="6.[151,1437,1436,2006]" box="[151,219,1436,1462]" pageId="6" pageNumber="49">Male.</paragraph>
<paragraph id="054AA211FF8DDF00AAF99AFAFAE8A07F" blockId="6.[151,1437,1436,2006]" pageId="6" pageNumber="49">
<emphasis id="37817E03FF8DDF00AAF99AFAFE9AA3F3" bold="true" box="[151,305,1474,1500]" pageId="6" pageNumber="49">Dimensions:</emphasis>
mean body length of males (n = 6) 1060 (range
<date id="714B84D1FF8DDF00A90F9AFAFC43A3F3" box="[865,1000,1474,1500]" pageId="6" pageNumber="49" value="1011" valueMax="1148">1011–1148</date>
); mean breadth 436 (range 396–488); mean length of females (n = 6); 1267 (range
<date id="714B84D1FF8DDF00A8C39AD1FC9CA02C" box="[685,823,1513,1539]" pageId="6" pageNumber="49" value="1203" valueMax="1356">1203–1356</date>
); mean breadth 488 (range 450–602); mean ratio of length of prodorsum to total length (both sexes): 0.32. The following supplementary description is based on the male, with details of artefactual change to the caudal region (
<figureCitation id="9DCEBE94FF8DDF00A9E0990EFC78A07F" box="[910,979,1590,1616]" captionStart="FIGURE 4" captionStartId="8.[151,255,1533,1557]" captionTargetBox="[248,1314,643,1457]" captionTargetId="figure@8.[235,1320,640,1113]" captionTargetPageId="8" captionText="FIGURE 4. Crotonia ovata Olszanowski, 2000, a) male, lateral view; b) caudal region of a female specimen from Mt. Victoria showing artefactual alteration in morphology (cf. text for details)." httpUri="https://zenodo.org/record/186206/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
b) based on a female specimen.
</paragraph>
<paragraph id="054AA211FF8DDF00AAAB9965FCE2A0C4" blockId="6.[151,1437,1436,2006]" pageId="6" pageNumber="49">
<emphasis id="37817E03FF8DDF00AAAB9965FEE1A059" box="[197,330,1629,1654]" italics="true" pageId="6" pageNumber="49">Prodorsum</emphasis>
: as illustrated by
<bibRefCitation id="6164DFE0FF8DDF00A84C9965FCB8A058" author="Olszanowski" box="[546,787,1629,1655]" pageId="6" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski (2000)</bibRefCitation>
except the trans-bothridial ridge may be less rounded (
<figureCitation id="9DCEBE94FF8DDF00AACE99BCFF40A0B1" box="[160,235,1668,1694]" captionStart="FIGURE 3" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[185,1402,482,1711]" captionTargetId="figure@7.[780,1416,495,1703]" captionTargetPageId="7" captionText="FIGURE 3. Crotonia ovata Olszanowski, 2000, male a) dorsal; b) ventral." httpUri="https://zenodo.org/record/186205/files/figure.png" pageId="6" pageNumber="49">Fig. 3</figureCitation>
a); rostrum with well-developed naso, the microsculpture of the region posterior to the bothridia consists of sparse scattered tubercles, and interlamellar setae extend beyond apices of rostral setae. Bases of lamellar setal apophyses conspicuously expanded laterally.
</paragraph>
<paragraph id="054AA211FF8DDF01AAAB99C0FD35A6F7" blockId="6.[151,1437,1436,2006]" lastBlockId="7.[151,1437,152,412]" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
<emphasis id="37817E03FF8DDF00AAAB99C0FEE3A13E" box="[197,328,1784,1809]" italics="true" pageId="6" pageNumber="49">Notogaster</emphasis>
: ratio of length to breadth 1.16. Anterior notogaster not simple as illustrated by
<bibRefCitation id="6164DFE0FF8DDF00AF6A99C0FF4CA117" author="Olszanowski" pageId="6" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski (2000)</bibRefCitation>
, but with pre-notogastral shield, separated from notogastral shield by transverse hyaline strip on which tubercles of setae
<emphasis id="37817E03FF8DDF00AB1D987DFE2BA171" box="[371,384,1861,1886]" italics="true" pageId="6" pageNumber="49">c</emphasis>
1 are positioned, visible on well-cleared specimens (
<figureCitation id="9DCEBE94FF8DDF00A994987DFBEFA170" box="[1018,1092,1861,1887]" captionStart="FIGURE 3" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[185,1402,482,1711]" captionTargetId="figure@7.[780,1416,495,1703]" captionTargetPageId="7" captionText="FIGURE 3. Crotonia ovata Olszanowski, 2000, male a) dorsal; b) ventral." httpUri="https://zenodo.org/record/186205/files/figure.png" pageId="6" pageNumber="49">Fig. 3</figureCitation>
a). Microsculpture of caudal region and that of the lateral strips consists of tubercles not alveloli. Median of the strips is a narrow zone of smooth cuticle. The centrodorsal region is microporose. The suprapleural suture is not fully visible in dorsal view, being positioned ventral of the lateral margin of the notogastral shield in lateral view (
<figureCitation id="9DCEBE94FF8DDF00AE8F9884FA81A1F9" box="[1249,1322,1980,2006]" captionStart="FIGURE 4" captionStartId="8.[151,255,1533,1557]" captionTargetBox="[248,1314,643,1457]" captionTargetId="figure@8.[235,1320,640,1113]" captionTargetPageId="8" captionText="FIGURE 4. Crotonia ovata Olszanowski, 2000, a) male, lateral view; b) caudal region of a female specimen from Mt. Victoria showing artefactual alteration in morphology (cf. text for details)." httpUri="https://zenodo.org/record/186206/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
a). Below the suprapleural suture, the microsculpture of the cuticle consists of scattered tubercles, and the caudal margin of the notogaster descends almost vertically.
</paragraph>
<paragraph id="054AA211FF8CDF01AAAB9FDDFEF0A7B3" blockId="7.[151,1437,152,412]" pageId="7" pageNumber="50">
Some specimens have undergone shrinkage caused by post-mortem muscle contraction. In these, there is an artefactual change in the morphology of the caudal notogastral region (
<figureCitation id="9DCEBE94FF8CDF01AE769E34FBC9A709" box="[1048,1122,268,294]" captionStart="FIGURE 4" captionStartId="8.[151,255,1533,1557]" captionTargetBox="[248,1314,643,1457]" captionTargetId="figure@8.[235,1320,640,1113]" captionTargetPageId="8" captionText="FIGURE 4. Crotonia ovata Olszanowski, 2000, a) male, lateral view; b) caudal region of a female specimen from Mt. Victoria showing artefactual alteration in morphology (cf. text for details)." httpUri="https://zenodo.org/record/186206/files/figure.png" pageId="7" pageNumber="50">Fig. 4</figureCitation>
b). The caudal notogastral region becomes narrowed and concave, the margins curled dorsally and there is a consequent relocation of the apophyses of setae
<emphasis id="37817E03FF8CDF01ABF39E61FE07A75D" box="[413,428,345,370]" italics="true" pageId="7" pageNumber="50">h</emphasis>
1 and
<emphasis id="37817E03FF8CDF01AB959E61FDA1A75D" box="[507,522,345,370]" italics="true" pageId="7" pageNumber="50">h</emphasis>
2 to positions whereby they point dorsally rather than posteriorly or posteriolaterally.
</paragraph>
</subSubSection>
<caption id="518AF299FF8CDF01AAF999EAFC1EA0C6" box="[151,949,1746,1770]" httpUri="https://zenodo.org/record/186205/files/figure.png" pageId="7" pageNumber="50" targetBox="[185,1402,482,1711]" targetPageId="7">
<paragraph id="054AA211FF8CDF01AAF999EAFC1EA0C6" blockId="7.[151,949,1746,1770]" box="[151,949,1746,1770]" pageId="7" pageNumber="50">
<emphasis id="37817E03FF8CDF01AAF999EAFEB2A0C5" bold="true" box="[151,281,1746,1770]" pageId="7" pageNumber="50">FIGURE 3.</emphasis>
<taxonomicName id="C2F5D992FF8CDF01AB4E99EAFD3CA0C6" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[288,663,1746,1769]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF8CDF01AB4E99EAFED4A0C6" box="[288,383,1746,1769]" italics="true" pageId="7" pageNumber="50">Crotonia</emphasis>
<emphasis id="37817E03FF8CDF01ABE899EAFE6AA0C6" box="[390,449,1746,1769]" italics="true" pageId="7" pageNumber="50">ovata</emphasis>
Olszanowski, 2000
</taxonomicName>
, male a) dorsal; b) ventral.
</paragraph>
</caption>
<subSubSection id="4DEFF19AFF8CDF01AAAB9802FCE7A1C0" pageId="7" pageNumber="50" type="materials_examined">
<paragraph id="054AA211FF8CDF01AAAB9802FCE7A1C0" blockId="7.[151,1438,1850,2031]" pageId="7" pageNumber="50">
<emphasis id="37817E03FF8CDF01AAAB9802FEBDA17C" box="[197,278,1850,1875]" italics="true" pageId="7" pageNumber="50">Ven t er</emphasis>
: epimeral microsculpture punctate, but with tuberculate margins. Sparsely tuberculate microsculpture in perigenital region and on adanal plates (
<figureCitation id="9DCEBE94FF8CDF01A92E9859FC2DA154" box="[832,902,1889,1915]" captionStart="FIGURE 3" captionStartId="7.[151,255,1746,1770]" captionTargetBox="[185,1402,482,1711]" captionTargetId="figure@7.[780,1416,495,1703]" captionTargetPageId="7" captionText="FIGURE 3. Crotonia ovata Olszanowski, 2000, male a) dorsal; b) ventral." httpUri="https://zenodo.org/record/186205/files/figure.png" pageId="7" pageNumber="50">Fig. 3</figureCitation>
b). Epimeral setal formula 3-1-3-3, seta
<emphasis id="37817E03FF8CDF01AF329859FADCA155" box="[1372,1399,1889,1914]" italics="true" pageId="7" pageNumber="50">3c</emphasis>
on well-developed tubercle (these setae covered in debris and not visible on the
<typeStatus id="DA4E1CB3FF8CDF01AE7298B0FB28A18D" box="[1052,1155,1928,1954]" pageId="7" pageNumber="50" type="holotype">holotype</typeStatus>
specimen).
<emphasis id="37817E03FF8CDF01AAAB9896FE10A1E7" bold="true" box="[197,443,1966,1992]" pageId="7" pageNumber="50">Material examined:</emphasis>
<typeStatus id="DA4E1CB3FF8CDF01ABAD9896FD81A1E7" box="[451,554,1966,1992]" pageId="7" pageNumber="50" type="holotype">holotype</typeStatus>
female, Hellyer River Gorge (in
<collectionCode id="63E43AD4FF8CDF01A9C09896FC50A1E7" LSID="urn:lsid:biocol.org:col:32981" box="[942,1019,1966,1992]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="50">ANIC</collectionCode>
). Non-type material: six males, two females from Mt. Michael; four females from Mt. Victoria.
</paragraph>
</subSubSection>
<subSubSection id="4DEFF19AFF83DF0FAAA89FAFFB84A418" lastPageId="9" lastPageNumber="52" pageId="8" pageNumber="51" type="discussion">
<paragraph id="054AA211FF83DF0EAAA89FAFFEA7A6D0" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="51">
<emphasis id="37817E03FF83DF0EAAA89FAFFE94A69E" bold="true" box="[198,319,151,177]" pageId="8" pageNumber="51">Remarks.</emphasis>
The above description, of the male of
<taxonomicName id="C2F5D992FF83DF0EA9439FA0FC3DA69E" box="[813,918,152,177]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF83DF0EA9439FA0FC3DA69E" box="[813,918,152,177]" italics="true" pageId="8" pageNumber="51">C. ovata</emphasis>
</taxonomicName>
, is intended to supplement Olszanowski’s (2000) description of the female. There is no significant morphological difference between the sexes apart from size.
</paragraph>
<paragraph id="054AA211FF83DF0EAAAB9E33FAF9A412" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="51">
<bibRefCitation id="6164DFE0FF83DF0EAAAB9E33FE06A709" author="Lochynska" box="[197,429,267,294]" pageId="8" pageNumber="54" refString="Lochynska, M. (2008 b) The ontogenetic description of two Tasmanian crotoniid mites (Acari: Oribatida: Crotoniidae). International Journal of Acarology, 34, 123 - 142." type="journal article" year="2008" yearSuffix="b">Łochyńska (2008b)</bibRefCitation>
described
<taxonomicName id="C2F5D992FF83DF0EA85E9E34FC84A70A" box="[560,815,268,293]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF83DF0EA85E9E34FC84A70A" box="[560,815,268,293]" italics="true" pageId="8" pageNumber="51">Holonothrus ryszardi</emphasis>
</taxonomicName>
and placed the species in
<taxonomicName id="C2F5D992FF83DF0EAE059E34FAAAA70A" box="[1131,1281,268,293]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAE059E34FAAAA70A" box="[1131,1281,268,293]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
even though the notogastral setation was tri-deficient, i.e. missing setae
<emphasis id="37817E03FF83DF0EA9329E0AFCC2A764" box="[860,873,306,331]" italics="true" pageId="8" pageNumber="51">c</emphasis>
2,
<emphasis id="37817E03FF83DF0EA9EC9E0AFC3AA764" box="[898,913,306,331]" italics="true" pageId="8" pageNumber="51">d</emphasis>
1 and
<emphasis id="37817E03FF83DF0EA9BA9E0AFC4AA764" box="[980,993,306,331]" italics="true" pageId="8" pageNumber="51">e</emphasis>
1.
<taxonomicName id="C2F5D992FF83DF0EA9979E0AFED2A759" authority="Norton &amp; Olszanowski, 1989" authorityName="Norton &amp; Olszanowski" authorityYear="1989" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="virungensis">
<emphasis id="37817E03FF83DF0EA9979E0AFA8BA764" box="[1017,1312,306,331]" italics="true" pageId="8" pageNumber="51">Holonothrus virungensis</emphasis>
<bibRefCitation id="6164DFE0FF83DF0EAF469E0AFED2A759" author="Norton" pageId="8" pageNumber="54" refString="Norton, R. A. &amp; Olszanowski, Z. (1989) A new Holonothrus (Oribatida: Crotoniioidea) from Zaire, with notes on the distribution of crotoniid mites. Revue Zoologique Africaine, 103, 405 - 412." type="journal article" year="1989">Norton &amp; Olszanowski, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C2F5D992FF83DF0EABEA9E64FD62A759" authority="Kuty, 2006" authorityName="Kuty" authorityYear="2006" box="[388,713,348,374]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="37817E03FF83DF0EABEA9E64FDE9A75A" box="[388,578,348,373]" italics="true" pageId="8" pageNumber="51">H. ecuadorensis</emphasis>
<bibRefCitation id="6164DFE0FF83DF0EA8269E64FD62A759" author="Kuty" box="[584,713,348,374]" pageId="8" pageNumber="54" refString="Kuty, M. (2006) A new species of the genus Holonothrus from Ecuador (Acari: Oribatida: Crotonioidea). Genus, 17, 307 - 310." type="journal article" year="2006">Kuty, 2006</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C2F5D992FF83DF0EA96F9E64FBCAA759" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[769,1121,348,374]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="nortoni">
<emphasis id="37817E03FF83DF0EA96F9E64FCD0A75A" box="[769,891,348,373]" italics="true" pageId="8" pageNumber="51">H. nortoni</emphasis>
<bibRefCitation id="6164DFE0FF83DF0EA9EF9E64FBCAA759" author="Olszanowski" box="[897,1121,348,374]" pageId="8" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski, 2000</bibRefCitation>
</taxonomicName>
lack
<emphasis id="37817E03FF83DF0EAEF09E64FB06A75A" box="[1182,1197,348,373]" italics="true" pageId="8" pageNumber="51">d</emphasis>
1 and
<emphasis id="37817E03FF83DF0EAE839E64FB51A75A" box="[1261,1274,348,373]" italics="true" pageId="8" pageNumber="51">e</emphasis>
1, but no other members of the genus are known to be trideficient (i.e. also lack
<emphasis id="37817E03FF83DF0EA9819EBEFC57A7B0" box="[1007,1020,390,415]" italics="true" pageId="8" pageNumber="51">c</emphasis>
2). The placement of
<taxonomicName id="C2F5D992FF83DF0EAF7F9EBEFAD0A7B0" box="[1297,1403,390,415]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF83DF0EAF7F9EBEFAD0A7B0" box="[1297,1403,390,415]" italics="true" pageId="8" pageNumber="51">ryszardi</emphasis>
</taxonomicName>
in
<taxonomicName id="C2F5D992FF83DF0EAAF99E97FE84A7E7" box="[151,303,431,456]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAAF99E97FE84A7E7" box="[151,303,431,456]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
appears to be predicated on the presence of two pairs of anal setae, mentioned by
<bibRefCitation id="6164DFE0FF83DF0EAF779E97FF5DA7DF" author="Lochynska" pageId="8" pageNumber="54" refString="Lochynska, M. (2008 b) The ontogenetic description of two Tasmanian crotoniid mites (Acari: Oribatida: Crotoniidae). International Journal of Acarology, 34, 123 - 142." type="journal article" year="2008" yearSuffix="b">Łochyńska (2008b)</bibRefCitation>
in the description and illustrated in her Fig. 9. However, the scanning electron micrograph of the ventral surface (her Fig. 25) clearly shows three pairs of anal setae, characteristic of
<taxonomicName id="C2F5D992FF83DF0EAED09EC5FA87A439" box="[1214,1324,509,534]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAED09EC5FA87A439" box="[1214,1324,509,534]" italics="true" pageId="8" pageNumber="51">Crotonia</emphasis>
</taxonomicName>
. The trideficient notogastral setation is also consistent with this species being a member of the genus
<taxonomicName id="C2F5D992FF83DF0EAE8C9D1BFAE0A413" box="[1250,1355,547,572]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAE8C9D1BFAE0A413" box="[1250,1355,547,572]" italics="true" pageId="8" pageNumber="51">Crotonia</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="518AF299FF83DF0EAAF99AC5FC1BA018" httpUri="https://zenodo.org/record/186206/files/figure.png" pageId="8" pageNumber="51" targetBox="[248,1314,643,1457]" targetPageId="8">
<paragraph id="054AA211FF83DF0EAAF99AC5FC1BA018" blockId="8.[151,1436,1533,1591]" pageId="8" pageNumber="51">
<emphasis id="37817E03FF83DF0EAAF99AC5FEB8A03A" bold="true" box="[151,275,1533,1557]" pageId="8" pageNumber="51">FIGURE 4</emphasis>
.
<taxonomicName id="C2F5D992FF83DF0EAB4C9AC6FD36A03A" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[290,669,1534,1557]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF83DF0EAB4C9AC6FE6EA03A" box="[290,453,1534,1557]" italics="true" pageId="8" pageNumber="51">Crotonia ovata</emphasis>
Olszanowski, 2000
</taxonomicName>
, a) male, lateral view; b) caudal region of a female specimen from Mt. Victoria showing artefactual alteration in morphology (cf. text for details).
</paragraph>
</caption>
<paragraph id="054AA211FF83DF0EAAAB995EFDE0A1F3" blockId="8.[151,1437,1637,2012]" pageId="8" pageNumber="51">
The immatures described by
<bibRefCitation id="6164DFE0FF83DF0EA871995DFCADA0AF" author="Lochynska" box="[543,774,1637,1664]" pageId="8" pageNumber="54" refString="Lochynska, M. (2008 b) The ontogenetic description of two Tasmanian crotoniid mites (Acari: Oribatida: Crotoniidae). International Journal of Acarology, 34, 123 - 142." type="journal article" year="2008" yearSuffix="b">Łochyńska (2008b)</bibRefCitation>
and assigned to
<taxonomicName id="C2F5D992FF83DF0EA9A1995EFBFFA050" box="[975,1108,1638,1663]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF83DF0EA9A1995EFBFFA050" box="[975,1108,1638,1663]" italics="true" pageId="8" pageNumber="51">H. ryszardi</emphasis>
</taxonomicName>
have a few character states consistent with immatures of
<taxonomicName id="C2F5D992FF83DF0EA86599B5FDD1A089" box="[523,634,1677,1702]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EA86599B5FDD1A089" box="[523,634,1677,1702]" italics="true" pageId="8" pageNumber="51">Crotonia</emphasis>
</taxonomicName>
, including the deutonymphal epimeral setal formula of 3-1-2-2, consistent with that for
<taxonomicName id="C2F5D992FF83DF0EABD0998BFC8BA0E2" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" box="[446,800,1715,1741]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF83DF0EABD0998BFD8CA0E3" box="[446,551,1715,1740]" italics="true" pageId="8" pageNumber="51">C. ovata</emphasis>
<bibRefCitation id="6164DFE0FF83DF0EA85E998BFC8BA0E2" author="Olszanowski" box="[560,800,1715,1741]" pageId="8" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski, 2000</bibRefCitation>
</taxonomicName>
, rather than 4-1/2-2/3-3 for those
<taxonomicName id="C2F5D992FF83DF0EAEA8998BFACAA0E3" box="[1222,1377,1715,1740]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAEA8998BFACAA0E3" box="[1222,1377,1715,1740]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
spp. described to date. Also the lateral notogastral setae emerge from small tubercles or apophyses that develop at the posterior ends of punctate cuticular plaques, whereas in
<taxonomicName id="C2F5D992FF83DF0EA9199839FBBBA135" box="[887,1040,1793,1818]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EA9199839FBBBA135" box="[887,1040,1793,1818]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
these setae emerge from alveoli typically positioned in the centre or anteriolateral portion of the plaques. The apparent presence of four setae in epimere I in the tritonymph is more typical of
<taxonomicName id="C2F5D992FF83DF0EA88B9876FCD0A148" box="[741,891,1870,1895]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EA88B9876FCD0A148" box="[741,891,1870,1895]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
, but this may be erroneous. It is important to note that the immatures were from samples that contain as many as four species of
<taxonomicName id="C2F5D992FF83DF0EAE1C984DFAADA1A1" box="[1138,1286,1909,1934]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAE1C984DFAADA1A1" box="[1138,1286,1909,1934]" italics="true" pageId="8" pageNumber="51">Holonothrus</emphasis>
</taxonomicName>
and three of
<taxonomicName id="C2F5D992FF83DF0EAAF998A3FEADA19B" box="[151,262,1947,1972]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF83DF0EAAF998A3FEADA19B" box="[151,262,1947,1972]" italics="true" pageId="8" pageNumber="51">Crotonia</emphasis>
</taxonomicName>
(including those described herein). Under these circumstances, matching immatures to adults correctly is less than straightforward.
</paragraph>
<paragraph id="054AA211FF82DF0FAAAB9FA0FB84A418" blockId="9.[151,1437,152,644]" pageId="9" pageNumber="52">
If one accepts the placement of
<taxonomicName id="C2F5D992FF82DF0FA82A9FA0FD61A69E" box="[580,714,152,177]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF82DF0FA82A9FA0FD61A69E" box="[580,714,152,177]" italics="true" pageId="9" pageNumber="52">H. ryszardi</emphasis>
</taxonomicName>
within
<taxonomicName id="C2F5D992FF82DF0FA9489FA0FC3BA69E" box="[806,912,152,177]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF82DF0FA9489FA0FC3BA69E" box="[806,912,152,177]" italics="true" pageId="9" pageNumber="52">Crotonia</emphasis>
</taxonomicName>
, then the next question is its differentiation from previously-described species. In fact, it shows striking similarity to
<taxonomicName id="C2F5D992FF82DF0FA9939F87FBCAA6F7" box="[1021,1121,191,216]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF82DF0FA9939F87FBCAA6F7" box="[1021,1121,191,216]" italics="true" pageId="9" pageNumber="52">C. ovata</emphasis>
</taxonomicName>
in the following character states: 1) the prodorsal ridges are the same length, curved and strongly divergent apically; 2) the transbothridial ridge is well-developed and distinctively bi-lobed; 3) the lengths and relative positions of notogastral setae of the
<emphasis id="37817E03FF82DF0FABC59E0AFE13A764" box="[427,440,306,331]" italics="true" pageId="9" pageNumber="52">c</emphasis>
series,
<emphasis id="37817E03FF82DF0FA87E9E0AFDB4A764" box="[528,543,306,331]" italics="true" pageId="9" pageNumber="52">d</emphasis>
2,
<emphasis id="37817E03FF82DF0FA85B9E0AFDE9A764" box="[565,578,306,331]" italics="true" pageId="9" pageNumber="52">e</emphasis>
2 and
<emphasis id="37817E03FF82DF0FA8EC9E0AFD21A764" box="[642,650,306,331]" italics="true" pageId="9" pageNumber="52">f</emphasis>
2 are almost an exact match; 4) the apices of the lamellar apophyses do not extend beyond the rostrum. This is an unusual feature in
<taxonomicName id="C2F5D992FF82DF0FA9E79E64FC59A75A" box="[905,1010,348,373]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF82DF0FA9E79E64FC59A75A" box="[905,1010,348,373]" italics="true" pageId="9" pageNumber="52">Crotonia</emphasis>
</taxonomicName>
(even accounting for the artefactual effect of any tilt of the specimen) and is confined to some members of the
<taxonomicName id="C2F5D992FF82DF0FAE5F9EBBFB16A7B3" box="[1073,1213,387,412]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="cophinaria">
<emphasis id="37817E03FF82DF0FAE5F9EBBFB16A7B3" box="[1073,1213,387,412]" italics="true" pageId="9" pageNumber="52">Cophinaria</emphasis>
</taxonomicName>
species group (
<taxonomicName id="C2F5D992FF82DF0FAFEE9EBBFEB2A7ED" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="cophinaria">
<emphasis id="37817E03FF82DF0FAFEE9EBBFEB2A7ED" italics="true" pageId="9" pageNumber="52">C. cophinaria</emphasis>
</taxonomicName>
,
<taxonomicName id="C2F5D992FF82DF0FAB469E91FE63A7ED" box="[296,456,425,450]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="lanceolata">
<emphasis id="37817E03FF82DF0FAB469E91FE63A7ED" box="[296,456,425,450]" italics="true" pageId="9" pageNumber="52">C. lanceolata</emphasis>
</taxonomicName>
,
<taxonomicName id="C2F5D992FF82DF0FABB99E91FDC1A7ED" box="[471,618,425,450]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="perforata">
<emphasis id="37817E03FF82DF0FABB99E91FDC1A7ED" box="[471,618,425,450]" italics="true" pageId="9" pageNumber="52">C. perforata</emphasis>
</taxonomicName>
,
<taxonomicName id="C2F5D992FF82DF0FA8179E91FCA6A7ED" box="[633,781,425,450]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="brassicae">
<emphasis id="37817E03FF82DF0FA8179E91FCA6A7ED" box="[633,781,425,450]" italics="true" pageId="9" pageNumber="52">C. brassicae</emphasis>
</taxonomicName>
and
<taxonomicName id="C2F5D992FF82DF0FA9279E91FC76A7ED" box="[841,989,425,450]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="37817E03FF82DF0FA9279E91FC76A7ED" box="[841,989,425,450]" italics="true" pageId="9" pageNumber="52">C. reticulata</emphasis>
</taxonomicName>
). In
<taxonomicName id="C2F5D992FF82DF0FAE769E91FB06A7ED" box="[1048,1197,425,450]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="37817E03FF82DF0FAE769E91FB06A7ED" box="[1048,1197,425,450]" italics="true" pageId="9" pageNumber="52">Holonothrus</emphasis>
</taxonomicName>
it is unusual for the lamellar apophyses to extend as far as the rostrum. Typically, they extend no more than half the distance between their bases and the rostrum (exceptions being
<taxonomicName id="C2F5D992FF82DF0FA9749ECFFBC8A43E" authority="Kuty, 2006" authorityName="Kuty" authorityYear="2006" box="[794,1123,503,529]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="37817E03FF82DF0FA9749ECFFC7CA43F" box="[794,983,503,528]" italics="true" pageId="9" pageNumber="52">H. ecuadorensis</emphasis>
<bibRefCitation id="6164DFE0FF82DF0FA9B09ECFFBC8A43E" author="Kuty" box="[990,1123,503,529]" pageId="9" pageNumber="54" refString="Kuty, M. (2006) A new species of the genus Holonothrus from Ecuador (Acari: Oribatida: Crotonioidea). Genus, 17, 307 - 310." type="journal article" year="2006">Kuty, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C2F5D992FF82DF0FAE019ECFFF79A418" authority="Lochynska, 2008" authorityName="Lochynska" authorityYear="2008" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="glaesarius">
<emphasis id="37817E03FF82DF0FAE019ECFFAA6A43F" box="[1135,1293,503,528]" italics="true" pageId="9" pageNumber="52">H. glaesarius</emphasis>
Łochyńska, 2008
</taxonomicName>
,
<taxonomicName id="C2F5D992FF82DF0FAA8E9D26FDEDA418" authority="Olszanowski, 1997" authorityName="Olszanowski" authorityYear="1997" box="[224,582,541,567]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="37817E03FF82DF0FAA8E9D26FEF6A418" box="[224,349,542,567]" italics="true" pageId="9" pageNumber="52">H. gracilis</emphasis>
<bibRefCitation id="6164DFE0FF82DF0FAB0B9D25FDEDA418" author="Olszanowski" box="[357,582,541,567]" pageId="9" pageNumber="54" refString="Olszanowski, Z. (1997) On the genus Holonothrus (Acari: Oribatida) in the Australian region. Acarologia, 37, 385 - 402." type="journal article" year="1997">Olszanowski, 1997</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="C2F5D992FF82DF0FA8109D26FB89A418" authority="Olszanowski, 1999" authorityName="Olszanowski" authorityYear="1999" box="[638,1058,541,567]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="venetiolanus">
<emphasis id="37817E03FF82DF0FA8109D26FC9DA418" box="[638,822,542,567]" italics="true" pageId="9" pageNumber="52">H. venetiolanus</emphasis>
<bibRefCitation id="6164DFE0FF82DF0FA9539D25FB89A418" author="Olszanowski" box="[829,1058,541,567]" pageId="9" pageNumber="54" refString="Olszanowski, Z. (1999) Three new species of Holonothrus from the Neotropical region (Acari: Oribatida). Journal of Natural History, 33, 233 - 253." type="journal article" year="1999">Olszanowski, 1999</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4DEFF19AFF82DF0FAAAB9D7CFE6CA4AB" pageId="9" pageNumber="52" type="reference_group">
<paragraph id="054AA211FF82DF0FAAAB9D7CFE6CA4AB" blockId="9.[151,1437,152,644]" pageId="9" pageNumber="52">
<taxonomicName id="C2F5D992FF82DF0FAAAB9D7CFD31A471" authority="Lochynska, 2008" authorityName="Lochynska" authorityYear="2008" box="[197,666,579,606]" class="Arachnida" family="Crotoniidae" genus="Holonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="37817E03FF82DF0FAAAB9D7CFE6FA472" box="[197,452,580,605]" italics="true" pageId="9" pageNumber="52">Holonothrus ryszardi</emphasis>
Łochyńska, 2008
</taxonomicName>
is hereby designated the junior subjective synonym of
<taxonomicName id="C2F5D992FF82DF0FAF5C9D7CFE68A4AB" ID-CoL="ZRHG" authority="Olszanowski, 2000" authorityName="Olszanowski" authorityYear="2000" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="37817E03FF82DF0FAF5C9D7CFF73A4AB" italics="true" pageId="9" pageNumber="52">Crotonia ovata</emphasis>
<bibRefCitation id="6164DFE0FF82DF0FAAB19D52FE68A4AB" author="Olszanowski" box="[223,451,618,644]" pageId="9" pageNumber="54" refString="Olszanowski, Z. (2000) Two new Australian species of Crotonia (Acari: Oribatida), with new records of Crotonioidea from the Australian Region. Acta Zoologica Academiae Scientiarum Hungaricae, 46, 239 - 248." type="journal article" year="2000">Olszanowski, 2000</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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