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178 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.231.3687" ID-GBIF-Dataset="9f36ba97-970f-40b6-ab7b-339c1f4cc0ef" ID-PMC="PMC3494023" ID-Pensoft-Pub="1313-2970-231-1" ID-PubMed="23204912" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-231-1" ModsDocOrigin="ZooKeys 231" ModsDocTitle="A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae)" checkinTime="1451248656124" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2012" docId="CB211462989A60F57B3F5123BFF17408" docLanguage="en" docName="ZooKeys 231: 1-116" docOrigin="ZooKeys 231" docSource="http://dx.doi.org/10.3897/zookeys.231.3687" docTitle="Polytrichophora setigera" docType="treatment" docVersion="3" lastPageNumber="63" masterDocId="FFE9FFC5E821FFA5FFA88C0071260670" masterDocTitle="A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae)" masterLastPageNumber="116" masterPageNumber="1" pageNumber="60" updateTime="1668154588673" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>231</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>116</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.231.3687</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.231.3687</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-231-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152037404" LSID="urn:lsid:plazi:treatment:CB211462989A60F57B3F5123BFF17408" httpUri="http://treatment.plazi.org/id/CB211462989A60F57B3F5123BFF17408" lastPageId="62" lastPageNumber="63" pageId="59" pageNumber="60">
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<subSubSection pageId="59" pageNumber="60" type="nomenclature">
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName LSID="http://species-id.net/wiki/Polytrichophora_setigera" authority="Cresson" class="Insecta" family="Ephydridae" genus="Polytrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polytrichophora setigera" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="setigera">Polytrichophora setigera (Cresson)</taxonomicName>
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Figs 104108
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</paragraph>
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</subSubSection>
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<subSubSection pageId="59" pageNumber="60" type="reference_group">
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName class="Insecta" family="Ephydridae" genus="Discocerina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocerina setigera" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="setigera">Discocerina setigera</taxonomicName>
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<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Entomological News" pageId="71" pageNumber="72" pagination="147 - 152" title="Descriptions of new genera and species of the dipterous family Ephydridae. III." volume="27" year="1916">Cresson 1916</bibRefCitation>
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: 148.
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</paragraph>
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName class="Insecta" family="Ephydridae" genus="Polytrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polytrichophora setigera" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="setigera">Polytrichophora setigera</taxonomicName>
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.
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<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Transactions of the American Entomological Society" pageId="71" pageNumber="72" pagination="101 - 128" title="Synopses of North American Ephydridae (Diptera) I. The subfamily Psilopinae, with descriptions of new species." volume="68" year="1942">Cresson 1942</bibRefCitation>
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: 114 [generic combination, review of Nearctic fauna].
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<bibRefCitation pageId="59" pageNumber="60">Wirth 1965</bibRefCitation>
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: 740 [Nearctic catalog].
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<bibRefCitation author="Mathis, WN" journalOrPublisher="Memoirs on Entomology, International" pageId="71" pageNumber="72" title="A world catalog of the shore flies (Diptera: Ephydridae)." volume="4" year="1995">Mathis and Zatwarnicki 1995</bibRefCitation>
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: 186 [world catalog].
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</paragraph>
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName class="Insecta" family="Ephydridae" genus="Polytrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polytrichophora orbitalis" order="Diptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="orbitalis">Polytrichophora orbitalis</taxonomicName>
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of authors, not Loew [misidentification].
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<bibRefCitation author="Zatwarnicki, T" journalOrPublisher="Annales Zoologici" pageId="71" pageNumber="72" pagination="5 - 51" title="A generic classification of the tribe Discocerinini (Diptera: Ephydridae)." volume="51" year="2001">Zatwarnicki and Mathis 2001</bibRefCitation>
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: 49 [illustration of male terminalia from Jackson County, Oregon].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="diagnosis">
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<paragraph pageId="59" pageNumber="60">Diagnosis.</paragraph>
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<paragraph pageId="59" pageNumber="60">This species is distinguished from congeners by the following combination of characters: Small shore flies, body length 1.60-1.95 mm.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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<pageBreakToken pageId="60" pageNumber="61" start="start">Head</pageBreakToken>
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: Frons largely dull to faintly subshiny, brown with moderate investment of gray microtomentum, anterior margin mostly concolorous, some specimens faintly orange; fronto-orbits narrow, gray to whitish gray. Antenna mostly yellow; basal flagellomere tannish yellow apicodorsally; arista with 5 dorsal rays. Face at narrowest point about equal to combined length of pedicel and basal flagellomere; face more densely microtomentose than frons, microtomentum with faintly shiny, mostly gray to whitish gray; parafacial and gena becoming slightly more whitish to silvery white than face; parafacial becoming wider ventrally with moderate ventral dilation; gena moderately high, height less than combined length of pedicel and basal flagellomere; gena-to-eye ratio 0.13-0.16.
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</paragraph>
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<paragraph pageId="60" pageNumber="61">Thorax: Mesonotum mostly dull to faintly subshiny, moderately densely microtomentose, mostly tan to brown, becoming gray laterally and anteriorly; pleural area mostly gray. Anterior margin of wing lacking spine-like setae; costal vein ratio 0.72-0.77; M vein ratio 0.48-0.55. Legs, including most of coxae, yellow; forefemur lacking a row of 9-10 short, stout setulae along apical half of anteroventral surface; forefemur with a row of 4-5, moderately well-developed, evenly spaced posteroventral setae on apical half, each seta less than width of femur; tibiae mostly gray, only apices yellowish; basal tarsomeres yellow, apical 1-2 becoming slightly darker, mostly tannish yellow.</paragraph>
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<paragraph pageId="60" pageNumber="61">Abdomen: Tergites subshiny with faint metallic tannish to dark gray luster medially, laterally gray; tergite 5 of male pointed, lacking distinctive row of setulae apically. Male terminalia (Figs 104-107): Epandrium in posterior view (Fig. 104) generally elliptical, more or less uniformly setulose, although slightly clumped dorsally and ventrally, dorsal margin broadly rounded, both ventral extensions pointed, narrowly connected dorsally, in lateral view (Fig. 105) with ventral apex acutely pointed; cercus relatively small but conspicuous; aedeagus tubular, in lateral view (Fig. 107) slightly tapered toward apex to bluntly rounded apex, in ventral view (Fig. 106) narrowly elongate with lateral margins very shallowly sinuous; phallapodeme in lateral view (Fig. 107) roughly triangular with extended keel bluntly rounded apically, in ventral view (Fig. 106) clavate with basal margin bluntly rounded; gonite in lateral view (Fig. 107) narrowly elongate, rounded angulate, angle obtuse, in ventral view (Fig. 106) conspicuously obtusely angulate, narrow, almost parallel sided; hypandrium in lateral view (Fig. 107) elongate, with lateral, irregular extension, in ventral view (Fig. 106) more or less as a rounded square with posterior, lateral arms.</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type material">
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<paragraph pageId="60" pageNumber="61">Type material.</paragraph>
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<paragraph pageId="60" pageNumber="61">The holotype male is labeled "MesaGrande[,] SonomaCo[,] V 1908 Cal[ifornia]/JPBaumberger Collector/♂/Holo-TYPE Discocerina SETIGERA E. T. Cresson Jr [magenta; species and generic names handwritten]. The holotype is double mounted (minute pin in rectangular card), is in good condition, and is deposited in the ANSP (6101).</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type locality">
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<paragraph pageId="60" pageNumber="61">Type locality.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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United States. California. Sonoma: Mesa Grande (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="32.943333">32°56.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-117.185">117°11.1'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="other specimens examined">
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<paragraph pageId="60" pageNumber="61">Other specimens examined.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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Nearctic. UNITED STATES. CALIFORNIA. Alameda: Berkeley Hills (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="37.883335">37°53'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-122.238335">122°14.3'W</geoCoordinate>
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), 11 Apr 1908 (3♂; ANSP). San Francisco: San Francisco (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.788334">37°47.3'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-122.495">122°29.7'W</geoCoordinate>
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), 27 May 1908, F. E. Blaisdell (1♂; ANSP).
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</paragraph>
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<paragraph pageId="61" pageNumber="62">
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<pageBreakToken pageId="61" pageNumber="62" start="start">OREGON</pageBreakToken>
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. Harney: Page Springs (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="42.783333">42°47'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-118.85">118°51'W</geoCoordinate>
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; 1300 m), 6 Aug 2005, D. and W. N. Mathis (6♂, 2♀; USNM); Pike Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.575">42°34.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-118.528336">118°31.7'W</geoCoordinate>
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; 1320 m), 7 Aug 2005, D. and W. N. Mathis (2♂, 2♀; USNM). Jackson: Rogue Elk Park, Rogue River, (ca. 8 km NE Shady Cove;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.656666">42°39.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-122.77167">122°46.3'W</geoCoordinate>
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), 28 Jul 1974, P. H. Arnaud, Jr. (1♂; USNM). Union: North Powder (3 km N;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="45.05">45°03'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-117.92">117°55.2'W</geoCoordinate>
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), 13 Jul 1988, W. N. Mathis (1♀; USNM).
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</paragraph>
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<paragraph pageId="61" pageNumber="62">
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WASHINGTON. Asotin: Snake River at Grande Ronde River (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="48.08">48°04.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-116.98167">116°58.9'W</geoCoordinate>
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;), 9 May 1915, A. L. Melander (1♂; ANSP). Franklin: Columbia River (NW Pasco;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.358334">46°21.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.25833">119°15.5'W</geoCoordinate>
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), 29 Jul 1998, D. and W. N. Mathis (1♂; USNM); Pasco (3.2 km E;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.25">46°15'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-119.11667">119°07'W</geoCoordinate>
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), 8 Aug 1975, W. N. Mathis (1♀; USNM); Ringold (NW Pasco:
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.506668">46°30.4'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.255">119°15.3'W</geoCoordinate>
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), 2-29 Jul 1988, 1998, D. and W. N. Mathis (8♂, 1♀; USNM); Sacajawea State Park (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.2">46°12'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.04">119°02.4'W</geoCoordinate>
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; 100 m), 12 Jul 1988, D. and W. N. Mathis (1♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="distribution">
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<paragraph pageId="61" pageNumber="62">Distribution</paragraph>
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<paragraph lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
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(Fig. 108). Nearctic: United States (California, Oregon, Washington)
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<pageBreakToken pageId="62" pageNumber="63" start="start">.</pageBreakToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="62" pageNumber="63" type="remarks">
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<paragraph pageId="62" pageNumber="63">Remarks.</paragraph>
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<paragraph pageId="62" pageNumber="63">Among congeners of the orbitalis group, this species is distinguished by characters of the male terminalia as follows: Area between anterior arms of the hypandrium membranous; the ventral margin of the epandrium is gradually tapered to apex; the hypandrium is large and more heavily sclerotized (Figs 104-107).</paragraph>
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<caption pageId="62" pageNumber="63">
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<paragraph pageId="62" pageNumber="63">
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Figures 104-107. Illustration of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Polytrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polytrichophora setigera" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="setigera">Polytrichophora setigera</taxonomicName>
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(Cresson) (male) (Idaho. Chatcolet) 104 epandrium and cerci, posterior view 105 same, lateral view 106 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 107 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="62" pageNumber="63">
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<paragraph pageId="62" pageNumber="63">
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Figure 108. Distribution of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Polytrichophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polytrichophora setigera" order="Diptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="setigera">Polytrichophora setigera</taxonomicName>
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(Cresson).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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