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158 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.468.8506" ID-GBIF-Dataset="df3f2d10-3b5e-480f-936c-368e01ffd7b1" ID-PMC="PMC4296520" ID-Pensoft-Pub="1313-2970-468-1" ID-PubMed="25610341" ID-ZBK="AE5AB793FDC74DCD8A47AE96A141E2AD" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-468-1" ModsDocOrigin="ZooKeys 468" ModsDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451244902156" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael" docDate="2014" docId="3B46ABA37131983A5372D98A176605FC" docLanguage="en" docName="ZooKeys 468: 1-83" docOrigin="ZooKeys 468" docSource="http://dx.doi.org/10.3897/zookeys.468.8506" docTitle="Exocelina sandaunensis Shaverdo & Balke, sp. n." docType="treatment" docUuid="0133AA56-7AA3-413B-B58D-89FE64CCF29E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="26" masterDocId="FFE9FFA5117AFFD17452FFB5FF94FFE3" masterDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="24" updateTime="1668159774671" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panjaitan, Rawati</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menufandu, Herlina</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>468</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.468.8506</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.468.8506</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-468-1</mods:identifier>
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<mods:identifier type="ZBK">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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<mods:identifier type="ZooBank">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152057385" LSID="urn:lsid:zoobank.org:act:0133AA56-7AA3-413B-B58D-89FE64CCF29E" httpUri="http://treatment.plazi.org/id/3B46ABA37131983A5372D98A176605FC" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
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<subSubSection pageId="23" pageNumber="24" type="multiple">
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<paragraph pageId="23" pageNumber="24">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="24" type="nomenclature">
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<paragraph pageId="23" pageNumber="24">
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17.
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<taxonomicName LSID="http://zoobank.org/0133AA56-7AA3-413B-B58D-89FE64CCF29E" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina sandaunensis" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sandaunensis">Exocelina sandaunensis Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="23" pageNumber="24">sp. n.</taxonomicNameLabel>
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Figs 13, 41
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="24" type="type locality">
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<paragraph pageId="23" pageNumber="24">Type locality.</paragraph>
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<paragraph pageId="23" pageNumber="24">
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Papua New Guinea: Sandaun Province, Sokamin,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-4.8475">04°50.85'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="141.63116">141°37.87'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="type material">
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<paragraph pageId="23" pageNumber="24">Type material.</paragraph>
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<paragraph pageId="24" pageNumber="25">
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<pageBreakToken pageId="24" pageNumber="25" start="start">Holotype</pageBreakToken>
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: male "Papua New Guinea: Sandaun, Sokamin4, 1200m, 19.x.2003,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.847417">4 50.845S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="141.63109">141 37.865E</geoCoordinate>
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, K. Sagata (WB 102)" (ZSM). Paratypes: 4 males with the same label as the holotype, one male additionally with a label "DNA M. Balke 666" (NHMW, ZSM). 3 males, 4 females "Papua New Guinea: Sandaun, Sokamin4, 1200m, 19.x.2003,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.847417">4 50.845S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="141.63109">141 37.865E</geoCoordinate>
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, K. Sagata (WB 100)", one male additionally with a label "DNA M. Balke 682" (NHMW, ZSM). 1 male "Papua New Guinea: Sandaun, MekilW100, 1718m, 14.xi.2003,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.8106165">4 48.637S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="141.6499">141 38.994E</geoCoordinate>
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, K. Sagata (WB 19)" (ZSM). 4 males "Papua New Guinea: Sandaun, MekilK [sic!], 1718m, 14.x.2003,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.8123665">4 48.742S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="141.65125">141 39.075E</geoCoordinate>
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, K. Sagata (WB 106)", two males additionally with labels "DNA M. Balke 672" and "DNA M. Balke 681" (NHMW, ZSM). 3 males "Papua New Guinea: Sandaun, Ofektaman, 820m, 17.x.2008, 5.04.113S 141.35.841E, Ibalim (PNG 190)", two males additionally with green labels "DNA M.Balke 3720", "DNA M.Balke 3721" (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="additional material">
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<paragraph pageId="24" pageNumber="25">Additional material.</paragraph>
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<paragraph pageId="24" pageNumber="25">
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2 females with the same label as the holotype (ZSM), these females might belong to two species:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina sandaunensis" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sandaunensis">Exocelina sandaunensis</taxonomicName>
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sp. n. and a species of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina rivulus" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rivulus">Exocelina rivulus</taxonomicName>
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group. 2 females "Papua New Guinea: Sandaun, MekilK [sic!], 1718m, 14.x.2003,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.8123665">4 48.742S</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="141.65125">141 39.075E</geoCoordinate>
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, K. Sagata (WB 106)" (ZSM), these females might belong to two species:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina sandaunensis" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sandaunensis">Exocelina sandaunensis</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ketembang" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ketembang">Exocelina ketembang</taxonomicName>
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Balke, 1998. 7 females "Papua New Guinea: Sandaun, Ofektaman, 820m, 17.x.2008, 5.04.113S 141.35.841E, Ibalim (PNG 190)" (ZSM), these females might belong to three species:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina sandaunensis" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sandaunensis">Exocelina sandaunensis</taxonomicName>
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sp. n.,
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina aipomek" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="aipomek">Exocelina aipomek</taxonomicName>
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Balke, 1998, and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ketembang" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ketembang">Exocelina ketembang</taxonomicName>
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Balke, 1998.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="diagnosis">
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<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
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<paragraph pageId="24" pageNumber="25">
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Beetle medium-sized, dark brown to piceous, slightly submatt; pronotum with lateral bead; male antennomeres 3-5 evidently enlarged, slightly rounded, antennomeres 3, 4 almost equal in size, antennomere 5 slightly smaller, with external margin almost straight, antennomere 6 somewhat enlarged; male protarsomere 4 with large, slender, evidently curved anterolateral hook-like seta; median lobe broad, with very weak submedian constriction in ventral view and thin apex in lateral view, apex with small lateral setae; paramere with notch on dorsal side and subdistal part elongate, with numerous, long, thick, curved at apex setae. This species is similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina simbaiarea" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="simbaiarea">Exocelina simbaiarea</taxonomicName>
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sp. n.,
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina tariensis" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tariensis">Exocelina tariensis</taxonomicName>
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sp. n., and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina jimiensis" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="jimiensis">Exocelina jimiensis</taxonomicName>
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sp. n., see differences under their diagnoses.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
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<paragraph pageId="24" pageNumber="25">Description.</paragraph>
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<paragraph pageId="24" pageNumber="25">Size and shape: Beetle medium-sized (TL-H 3.5-4.0 mm, TL 3.9-4.5 mm, MW 1.85-2.15 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsal surface more or less uniform dark brown to piceous, slightly paler on clypeus, vertex, pronotal sides, and along elytral suture; head appendages and legs yellowish red, legs reddish brown distally (Fig. 41). Teneral specimens paler.</paragraph>
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<paragraph pageId="24" pageNumber="25">Surface sculpture: Head with dense punctation (spaces between punctures 1-3 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures equal or smaller than diameter of cells of microreticulation. Pronotum with finer, sparser, and more evenly distributed punctation than on head. Elytra with very sparse and fine punctation. Pronotum and elytra with relatively weakly impressed microreticulation, dorsal surface slightly submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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<pageBreakToken pageId="25" pageNumber="26" start="start">Structures</pageBreakToken>
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: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process and neck of prosternal process with distinct ridge, smooth and not rounded anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 slightly truncate apically.
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</paragraph>
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<paragraph pageId="25" pageNumber="26">Male: Antennomeres 3-5 evidently enlarged, slightly rounded, antennomeres 3, 4 almost equal in size, antennomere 5 slightly smaller, with external margin almost straight, antennomere 6 somewhat enlarged; (Fig. 13A). Protarsomere 4 with large, slender, evidently curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 15 and posterior row of 3 short setae (Fig. 13B). Abdominal ventrite 6 with 7-10 lateral striae on each side. Median lobe broad, with very weak submedian constriction in ventral view and thin apex in lateral view, apex with small lateral setae (Figs 13C, D). Paramere with notch on dorsal side and subdistal part elongate, with numerous, long, thick, curved at apex setae (Fig. 13E).</paragraph>
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<paragraph pageId="25" pageNumber="26">Holotype: TL-H 3.75 mm, TL 4.25 mm, MW 2 mm.</paragraph>
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<paragraph pageId="25" pageNumber="26">Female: Antennae simple, abdominal ventrite 6 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="distribution">
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<paragraph pageId="25" pageNumber="26">Distribution.</paragraph>
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<paragraph pageId="25" pageNumber="26">Papua New Guinea. This species is known from Sandaun Province (Fig. 53).</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="etymology">
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<paragraph pageId="25" pageNumber="26">Etymology.</paragraph>
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<paragraph pageId="25" pageNumber="26">The species is named after Sandaun Province where it was collected. The name is an adjective in the nominative singular.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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