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<document id="4FBDC6AA613E4202CF79C89DE7DE9223" ID-DOI="10.1111/zoj.12337" ID-ISSN="0024-4082" ID-Zenodo-Dep="5359514" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630227898451" checkinUser="marcus" docAuthor="Monjaraz-Ruedas, Rodrigo &amp; Francke, Oscar F." docDate="2016" docId="03A787D0FFA4A711FCE47623FE82FD65" docLanguage="en" docName="zoj12337.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (4)" docSource="http://dx.doi.org/10.1111/zoj.12337" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Mayazomus estor Monjaraz-Ruedas &amp; Francke 2016" docType="treatment" docVersion="3" lastPageNumber="801" masterDocId="FF9EFFA8FFB4A705FFA0722EFFA5FFC0" masterDocTitle="Systematics of the genus Mayazomus (Arachnida: Schizomida): the relevance of using continuous characters and pedipalp setae patterns to schizomid phylogenetics" masterLastPageNumber="805" masterPageNumber="781" pageNumber="797" updateTime="1699192520556" updateUser="plazi">
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<mods:title id="67C0A2C101329B3B930471DCF2374E1A">Systematics of the genus Mayazomus (Arachnida: Schizomida): the relevance of using continuous characters and pedipalp setae patterns to schizomid phylogenetics</mods:title>
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<mods:namePart id="D66706D0644BFC95FD6F34BC1E4DA929">Monjaraz-Ruedas, Rodrigo</mods:namePart>
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<mods:namePart id="6ADC7EB9E9A45B054164815591EB8FA6">Francke, Oscar F.</mods:namePart>
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<mods:title id="219021EEE6DF94780B0C75287262B5AC">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="0B013D16746EA5A1625AF2DB678F5180">2016</mods:date>
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<mods:number id="DF79B550CACAABCB9123289AD07919CF">2015-11-16</mods:number>
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<mods:number id="D04CA7A8E818DD7201E5FF0047F582E5">176</mods:number>
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<mods:identifier id="5C665F0450689084D64A037F92B0BD00" type="DOI">10.1111/zoj.12337</mods:identifier>
<mods:identifier id="D9EB6C13650FC904D1CCDC6E844593D5" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03A787D0FFA4A711FCE47623FE82FD65" LSID="urn:lsid:plazi:treatment:03A787D0FFA4A711FCE47623FE82FD65" httpUri="http://treatment.plazi.org/id/03A787D0FFA4A711FCE47623FE82FD65" lastPageId="20" lastPageNumber="801" pageId="16" pageNumber="797">
<subSubSection id="C314654DFFA4A715FCE47623FB41FB83" pageId="16" pageNumber="797" type="nomenclature">
<paragraph id="8BB136C6FFA4A715FCE47623FAD5FBE3" blockId="16.[836,1392,1037,1091]" box="[836,1392,1037,1059]" pageId="16" pageNumber="797">
<heading id="D0F981AAFFA4A715FCE47623FAD5FBE3" box="[836,1392,1037,1059]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="797" reason="8">
<taxonomicName id="4C0E4D45FFA4A715FCE47623FB85FBE0" ID-CoL="3YH7L" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[836,1056,1037,1058]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="805" phylum="Arthropoda" rank="species" species="estor">
<emphasis id="B97AEAD4FFA4A715FCE47623FB85FBE0" box="[836,1056,1037,1058]" italics="true" pageId="16" pageNumber="797">MAYAZOMUS ESTOR</emphasis>
</taxonomicName>
(DE ARMAS &amp; VÍQUEZ, 2010)
</heading>
</paragraph>
<paragraph id="8BB136C6FFA4A715FC6F761EFB41FB83" blockId="16.[836,1392,1037,1091]" box="[975,1252,1069,1091]" pageId="16" pageNumber="797">
<heading id="D0F981AAFFA4A715FC6F761EFB41FB83" box="[975,1252,1069,1091]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="797" reason="4">
<emphasis id="B97AEAD4FFA4A715FC6F761EFBF3FB82" bold="true" box="[975,1110,1069,1090]" pageId="16" pageNumber="797">COMB. NOV.</emphasis>
(
<figureCitation id="13352A43FFA4A715FBC77603FB79FB83" box="[1127,1244,1069,1091]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStartId-0="17.[164,243,870,889]" captionStartId-1="18.[144,223,1730,1749]" captionStartId-2="19.[164,243,1227,1246]" captionTargetBox-0="[421,1184,198,838]" captionTargetBox-1="[306,1258,199,1695]" captionTargetBox-2="[164,1444,195,1197]" captionTargetId-0="figure-447@17.[419,1187,196,840]" captionTargetId-1="figure-0@18.[303,1263,194,1700]" captionTargetId-2="figure-302@19.[164,1444,195,1197]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="Figure 8. Mayazomus estor comb. nov. Paratype male. Habitus: A, dorsal view. Paratype female. Habitus: B, dorsal view. Scale bar = 1 mm." captionText-1="Figure 9. Mayazomus estor comb. nov. Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm." captionText-2="Figure 10. Mayazomus estor comb. nov. holotype male. Chelicera (A–C): A, mesal view. B, fixed finger. C, moveable finger. Paratype female. Spermathecae: D, dorsal view. Scale bars = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5359530" figureDoi-1="http://doi.org/10.5281/zenodo.5359532" figureDoi-2="http://doi.org/10.5281/zenodo.5359534" httpUri-0="https://zenodo.org/record/5359530/files/figure.png" httpUri-1="https://zenodo.org/record/5359532/files/figure.png" httpUri-2="https://zenodo.org/record/5359534/files/figure.png" pageId="16" pageNumber="797">FIGS 8–10</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C314654DFFA4A715FC86767DFB9BFB65" pageId="16" pageNumber="797" type="reference_group">
<paragraph id="8BB136C6FFA4A715FC86767DFB9BFB65" blockId="16.[806,1421,1107,1190]" pageId="16" pageNumber="797">
<treatmentCitationGroup id="AB1E11E8FFA4A715FC86767DFB9BFB65" pageId="16" pageNumber="797">
<taxonomicName id="4C0E4D45FFA4A715FC86767DFCE6FB47" ID-CoL="6M4MV" authority="de Armas &amp; Viquez, 2010: 12 - 13" authorityName="de Armas &amp; Viquez" authorityPageNumber="12 - 13" authorityYear="2010" class="Arachnida" family="Hubbardiidae" genus="Heteronochrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="estor">
<emphasis id="B97AEAD4FFA4A715FC86767DFBABFBA9" box="[806,1038,1107,1129]" italics="true" pageId="16" pageNumber="797">Heteronochrus estor</emphasis>
de Armas &amp; Víquez, 2010: 12– 13
</taxonomicName>
, fig. 4A–H, table II;
<treatmentCitation id="0AAF10D7FFA4A715FB8F765CFC10FB65" author="Moreno-Gonzalez JA &amp; Villarreal MO" page="74" pageId="16" pageNumber="797" year="2012">
<bibRefCitation id="EF9F4B37FFA4A715FB8F765CFC10FB65" author="Moreno-Gonzalez JA &amp; Villarreal MO" pageId="16" pageNumber="797" pagination="61 - 78" refId="ref16615" refString="Moreno-Gonzalez JA, Villarreal MO. 2012. A new genus of Hubbardiidae (Arachnida, Schizomida) from the Colombian Andes, with some taxonomic comments. Zootaxa 3560: 61 - 78." type="journal article" year="2012">Moreno-González &amp; Villarreal, 2012: 74–77</bibRefCitation>
</treatmentCitation>
, tables 2–4.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C314654DFFA4A711FC8676FBFE82FD65" lastPageId="20" lastPageNumber="801" pageId="16" pageNumber="797" type="diagnosis">
<paragraph id="8BB136C6FFA4A715FC8676FBFBA0FB2A" blockId="16.[806,1422,1237,1902]" box="[806,1029,1237,1258]" pageId="16" pageNumber="797">
<emphasis id="B97AEAD4FFA4A715FC8676FBFBA0FB2A" box="[806,1029,1237,1258]" italics="true" pageId="16" pageNumber="797">Emended diagnosis</emphasis>
</paragraph>
<paragraph id="8BB136C6FFA4A714FC8676DAFCA0FBC3" blockId="16.[806,1422,1237,1902]" lastBlockId="17.[164,779,974,1027]" lastPageId="17" lastPageNumber="798" pageId="16" pageNumber="797">
Males can be distinguished by the pedipalp trochanter having a long, pointed apical process less than 45°; the pedipalp femur with spiniform setiferous tubercles
<emphasis id="B97AEAD4FFA4A715FCD9777EFC07FAA5" box="[889,930,1360,1381]" italics="true" pageId="16" pageNumber="797">Fv1</emphasis>
and
<emphasis id="B97AEAD4FFA4A715FC7A777EFBA6FAA5" box="[986,1027,1360,1381]" italics="true" pageId="16" pageNumber="797">Fv2</emphasis>
small and close to each other; setae
<emphasis id="B97AEAD4FFA4A715FC867740FCF4FA43" box="[806,849,1390,1411]" italics="true" pageId="16" pageNumber="797">Fv1</emphasis>
located ventrally, setae
<emphasis id="B97AEAD4FFA4A715FBC67740FB2AFA43" box="[1126,1167,1390,1411]" italics="true" pageId="16" pageNumber="797">Fe1</emphasis>
small and acuminate; the apophysis on the mesal surface (FAP) small and inconspicuous; pedipalp tibia without mesal apophysis (TMA); male flagellum cordate, with a submedian depression on the dorsal surface of the flagellum (
<figureCitation id="13352A43FFA4A715FA9177C7FC99F9DD" captionStart="Figure 9" captionStartId="18.[144,223,1730,1749]" captionTargetBox="[306,1258,199,1695]" captionTargetId="figure-0@18.[303,1263,194,1700]" captionTargetPageId="18" captionText="Figure 9. Mayazomus estor comb. nov. Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5359532" httpUri="https://zenodo.org/record/5359532/files/figure.png" pageId="16" pageNumber="797">Fig. 9E– G</figureCitation>
); pedipalp trochanter of heteromorphic male with a pronounced mesal spur. Females can be distinguished by the spermathecae (
<figureCitation id="13352A43FFA4A715FB29746AFB4EF99A" box="[1161,1259,1604,1626]" captionStart="Figure 10" captionStartId="19.[164,243,1227,1246]" captionTargetBox="[164,1444,195,1197]" captionTargetId="figure-302@19.[164,1444,195,1197]" captionTargetPageId="19" captionText="Figure 10. Mayazomus estor comb. nov. holotype male. Chelicera (A–C): A, mesal view. B, fixed finger. C, moveable finger. Paratype female. Spermathecae: D, dorsal view. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5359534" httpUri="https://zenodo.org/record/5359534/files/figure.png" pageId="16" pageNumber="797">Fig. 10D</figureCitation>
) having lobes subequal in length; median lobes curved along their length; a wide chitinized arc.
<taxonomicName id="4C0E4D45FFA4A715FBD774ACFA9CF957" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[1143,1337,1666,1687]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="estor">
<emphasis id="B97AEAD4FFA4A715FBD774ACFA9CF957" box="[1143,1337,1666,1687]" italics="true" pageId="16" pageNumber="797">Mayazomus estor</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C0E4D45FFA4A715FCF7748FFC09F976" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2015" box="[855,940,1697,1718]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="yaax">
<emphasis id="B97AEAD4FFA4A715FCF7748FFC09F976" box="[855,940,1697,1718]" italics="true" pageId="16" pageNumber="797">M. yaax</emphasis>
</taxonomicName>
in the general shape of the pedipalp, having the femur less robust than the other species of the genus, and the females having a V-shaped chitinized arc;
<taxonomicName id="4C0E4D45FFA4A715FCF874D3FC17F8D2" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[856,946,1789,1810]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="estor">
<emphasis id="B97AEAD4FFA4A715FCF874D3FC17F8D2" box="[856,946,1789,1810]" italics="true" pageId="16" pageNumber="797">M. estor</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C0E4D45FFA4A715FB9374D3FB60F8D1" authorityName="Rowland" authorityYear="1975" box="[1075,1221,1788,1810]" class="Arachnida" family="Hubbardiidae" genus="Schizomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B97AEAD4FFA4A715FB9374D3FB60F8D1" box="[1075,1221,1788,1810]" italics="true" pageId="16" pageNumber="797">M. infernalis</emphasis>
</taxonomicName>
in the shape and subequal length of the spermathecae lobes, and the curvature on the median lobes. The male flagellum of
<taxonomicName id="4C0E4D45FFA4A715FC867576FC20F8AD" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[806,901,1880,1901]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="16" pageNumber="797" phylum="Arthropoda" rank="species" species="estor">
<emphasis id="B97AEAD4FFA4A715FC867576FC20F8AD" box="[806,901,1880,1901]" italics="true" pageId="16" pageNumber="797">M. estor</emphasis>
</taxonomicName>
is cordate in shape, with a single dorsal submedian depression, whereas in
<taxonomicName id="4C0E4D45FFA5A714FD8871E1FD11FC23" authorityName="Rowland" authorityYear="1975" box="[552,692,974,996]" class="Arachnida" family="Hubbardiidae" genus="Schizomus" kingdom="Animalia" order="Schizomida" pageId="17" pageNumber="798" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B97AEAD4FFA5A714FD8871E1FD11FC23" box="[552,692,974,996]" italics="true" pageId="17" pageNumber="798">M. infernalis</emphasis>
</taxonomicName>
the flagellum has two inconspicuous posterior depressions.
</paragraph>
<caption id="DF71664EFFA5A714FF047148FE04FC57" ID-DOI="http://doi.org/10.5281/zenodo.5359530" ID-Zenodo-Dep="5359530" httpUri="https://zenodo.org/record/5359530/files/figure.png" pageId="17" pageNumber="798" startId="17.[164,243,870,889]" targetBox="[421,1184,198,838]" targetPageId="17">
<paragraph id="8BB136C6FFA5A714FF047148FE04FC57" blockId="17.[164,1442,870,919]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FF047148FEA9FCB9" bold="true" box="[164,268,870,889]" pageId="17" pageNumber="798">Figure 8.</emphasis>
<taxonomicName id="4C0E4D45FFA5A714FEB77149FE6AFCB9" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[279,463,871,889]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="17" pageNumber="798" phylum="Arthropoda" rank="species" species="estor" status="comb. nov.">
<emphasis id="B97AEAD4FFA5A714FEB77149FE6AFCB9" box="[279,463,871,889]" italics="true" pageId="17" pageNumber="798">Mayazomus estor</emphasis>
</taxonomicName>
<emphasis id="B97AEAD4FFA5A714FE797149FDFDFCBA" bold="true" box="[473,600,871,890]" pageId="17" pageNumber="798">
<taxonomicNameLabel id="A24957AFFFA5A714FE797149FDFDFCBA" box="[473,600,871,890]" pageId="17" pageNumber="798" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Paratype male. Habitus: A, dorsal view. Paratype female. Habitus: B, dorsal view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BB136C6FFA5A714FF04761EFE73FB85" blockId="17.[164,779,1072,1186]" box="[164,470,1072,1093]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FF04761EFE73FB85" box="[164,470,1072,1093]" italics="true" pageId="17" pageNumber="798">Supplementary description</emphasis>
</paragraph>
<paragraph id="8BB136C6FFA5A714FF047661FDD0FB62" blockId="17.[164,779,1072,1186]" pageId="17" pageNumber="798">
The following description and meristic data for the
<typeStatus id="54B58864FFA5A714FF047643FEA0FB43" box="[164,261,1133,1155]" pageId="17" pageNumber="798" type="holotype">holotype</typeStatus>
and
<typeStatus id="54B58864FFA5A714FEE67640FE12FB43" box="[326,439,1134,1155]" pageId="17" pageNumber="798" type="paratype">paratypes</typeStatus>
supplement the original description by de Armas &amp; Víquez (2010).
</paragraph>
<paragraph id="8BB136C6FFA5A714FF0476E1FEDFF9B3" blockId="17.[164,779,1231,1651]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FF0476E1FE11FB24" box="[164,436,1231,1252]" italics="true" pageId="17" pageNumber="798">
Chelicerae (
<figureCitation id="13352A43FFA5A714FE8976E1FE03FB24" box="[297,422,1231,1252]" captionStart="Figure 10" captionStartId="19.[164,243,1227,1246]" captionTargetBox="[164,1444,195,1197]" captionTargetId="figure-302@19.[164,1444,195,1197]" captionTargetPageId="19" captionText="Figure 10. Mayazomus estor comb. nov. holotype male. Chelicera (A–C): A, mesal view. B, fixed finger. C, moveable finger. Paratype female. Spermathecae: D, dorsal view. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5359534" httpUri="https://zenodo.org/record/5359534/files/figure.png" pageId="17" pageNumber="798">Fig. 10A–C</figureCitation>
):
</emphasis>
Moveable finger: serrula with 16 teeth, guard tooth present. Fixed finger with four smaller teeth between two primary teeth. Setation: setal group formula: 1:3, 2:6, 3:4, 4:2, 5:7, 6:1. G1 (setae group 1) with three spatulate setae, covered with few spinose and small spicules starting from the middle of the shaft; G2 composed of six feathered setae subequal in length, and shorter than moveable finger length; G3 with four setae subequal in length, feathered apically, and smooth basally; G4 consisting of two setae, smooth, short, and thick, elongated at the tip; G5 with seven similarsized setae, feathered apically and longer than fixed finger; G6 with one smooth setae about half of moveable finger length.
</paragraph>
<paragraph id="8BB136C6FFA5A714FF04748EFBF4FAB2" blockId="17.[164,779,1696,1902]" lastBlockId="17.[826,1441,974,1394]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FF04748EFE03F975" box="[164,422,1696,1717]" italics="true" pageId="17" pageNumber="798">
Pedipalps (
<figureCitation id="13352A43FFA5A714FE85748EFE3DF975" box="[293,408,1696,1717]" captionStart="Figure 9" captionStartId="18.[144,223,1730,1749]" captionTargetBox="[306,1258,199,1695]" captionTargetId="figure-0@18.[303,1263,194,1700]" captionTargetPageId="18" captionText="Figure 9. Mayazomus estor comb. nov. Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5359532" httpUri="https://zenodo.org/record/5359532/files/figure.png" pageId="17" pageNumber="798">Fig. 9A–D</figureCitation>
):
</emphasis>
Robust 2.42 times longer than propeltidium length. Trochanter 2.33 times longer than high; apical process pointed and broadly triangular, less than 45°; mesal surface with a row of three setae near ventral margin, plus two setae near dorsal margin; with a pronounced medial mesal spur. Femur 1.52 times longer than high, distal margin straight, ventral margin flattened; distal margin on ectal surface with spiniform setiferous tubercles
<emphasis id="B97AEAD4FFA5A714FB8471C3FBEBFBC2" box="[1060,1102,1005,1026]" italics="true" pageId="17" pageNumber="798">Fv1</emphasis>
and
<emphasis id="B97AEAD4FFA5A714FB2C71C3FB12FBC2" box="[1164,1207,1005,1026]" italics="true" pageId="17" pageNumber="798">Fv2</emphasis>
small, close to each other;
<emphasis id="B97AEAD4FFA5A714FC247625FC08FBE0" box="[900,941,1035,1056]" italics="true" pageId="17" pageNumber="798">Fe1</emphasis>
small, an acuminate setae on ectal surface; mesal surface with a row of three ventral setae plus two dorsal spiniform setae (
<emphasis id="B97AEAD4FFA5A714FBCA7666FB12FB9D" box="[1130,1207,1096,1117]" italics="true" pageId="17" pageNumber="798">Fvr1–3</emphasis>
, setae
<emphasis id="B97AEAD4FFA5A714FAA47666FA95FB9D" box="[1284,1328,1096,1117]" italics="true" pageId="17" pageNumber="798">Fd1</emphasis>
and
<emphasis id="B97AEAD4FFA5A714FAC77666FA37FB9D" box="[1383,1426,1096,1117]" italics="true" pageId="17" pageNumber="798">Fd2</emphasis>
); distal margin of the mesal surface with a very small and inconspicuous apophysis (FAP). Patella slightly curved, inverse arc-shaped; setae
<emphasis id="B97AEAD4FFA5A714FB62768BFB78FB7A" box="[1218,1245,1189,1210]" italics="true" pageId="17" pageNumber="798">Pe</emphasis>
,
<emphasis id="B97AEAD4FFA5A714FB4D768BFAB6FB7A" box="[1261,1299,1189,1210]" italics="true" pageId="17" pageNumber="798">Pm</emphasis>
,
<emphasis id="B97AEAD4FFA5A714FA83768AFAC2FB79" box="[1315,1383,1188,1209]" italics="true" pageId="17" pageNumber="798">Pe5–3</emphasis>
, and
<emphasis id="B97AEAD4FFA5A714FC9A76EDFC2CFB18" box="[826,905,1219,1240]" italics="true" pageId="17" pageNumber="798">Pm5–3</emphasis>
acuminate, without special armature. Setae formula on tibia 3:3:4 (external row: medial row: internal row); seta
<emphasis id="B97AEAD4FFA5A714FC5A772EFB8BFAD5" box="[1018,1070,1280,1301]" italics="true" pageId="17" pageNumber="798">Tm1</emphasis>
acuminate and setae
<emphasis id="B97AEAD4FFA5A714FA85772EFAFDFAD5" box="[1317,1368,1280,1301]" italics="true" pageId="17" pageNumber="798">Tm2</emphasis>
feathered; without tibial mesal apophysis (TMA). Claw about half as long as dorsal length of tarsus; spurs asymmetrical,
<quantity id="4CF69B23FFA5A714FC0A7772FBAAFAB1" box="[938,1039,1372,1393]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="798" unit="mm" value="0.09">0.09 mm</quantity>
long.
</paragraph>
<paragraph id="8BB136C6FFA5A714FC9A779CFA07FA26" blockId="17.[826,1442,1457,1510]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FC9A779CFCDFFA07" box="[826,890,1458,1479]" italics="true" pageId="17" pageNumber="798">Legs:</emphasis>
Leg I, basitarsal-telotarsal proportions: 20:3:3:4:4:4:9. Femur IV 2.25 times longer than wide.
</paragraph>
<paragraph id="8BB136C6FFA5A714FC9A7408FBFCF8AE" blockId="17.[826,1442,1574,1902]" pageId="17" pageNumber="798">
<emphasis id="B97AEAD4FFA5A714FC9A7408FC71F9FB" box="[826,980,1574,1595]" italics="true" pageId="17" pageNumber="798">Opisthosoma:</emphasis>
Tergite I with two pairs of microsetae anteriorly plus one pair of macrosetae 2 + 1; tergite II with three pairs of microsetae anteriorly plus five macrosetae 3 + 2; tergites III–VII with one pair of dorsal setae each; tergites VIII–IX with one pair of dorsal and one pair of lateral setae each; segments X–XI telescoped with one pair of lateral setae and five ventral setae; segment XII with one dorsal and two lateral setae and three ventral pairs of setae, without posterodorsal process. Sternites with scattered setae; genital aperture without microsetae.
</paragraph>
<caption id="DF71664EFFA6A717FF3074ECFB35F8ED" ID-DOI="http://doi.org/10.5281/zenodo.5359532" ID-Zenodo-Dep="5359532" httpUri="https://zenodo.org/record/5359532/files/figure.png" pageId="18" pageNumber="799" startId="18.[144,223,1730,1749]" targetBox="[306,1258,199,1695]" targetPageId="18">
<paragraph id="8BB136C6FFA6A717FF3074ECFB35F8ED" blockId="18.[144,1421,1730,1837]" pageId="18" pageNumber="799">
<emphasis id="B97AEAD4FFA6A717FF3074ECFF5DF915" bold="true" box="[144,248,1730,1749]" pageId="18" pageNumber="799">Figure 9.</emphasis>
<taxonomicName id="4C0E4D45FFA6A717FEA374EDFE1DF915" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[259,440,1731,1749]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="18" pageNumber="799" phylum="Arthropoda" rank="species" species="estor" status="comb. nov.">
<emphasis id="B97AEAD4FFA6A717FEA374EDFE1DF915" box="[259,440,1731,1749]" italics="true" pageId="18" pageNumber="799">Mayazomus estor</emphasis>
</taxonomicName>
<emphasis id="B97AEAD4FFA6A717FE6074ECFD99F915" bold="true" box="[448,572,1730,1749]" pageId="18" pageNumber="799">
<taxonomicNameLabel id="A24957AFFFA6A717FE6074ECFD99F915" box="[448,572,1730,1749]" pageId="18" pageNumber="799" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="DF71664EFFA7A716FF0476E5FC4DFB3C" ID-DOI="http://doi.org/10.5281/zenodo.5359534" ID-Zenodo-Dep="5359534" httpUri="https://zenodo.org/record/5359534/files/figure.png" pageId="19" pageNumber="800" startId="19.[164,243,1227,1246]" targetBox="[164,1444,195,1197]" targetPageId="19">
<paragraph id="8BB136C6FFA7A716FF0476E5FC4DFB3C" blockId="19.[164,1441,1227,1276]" pageId="19" pageNumber="800">
<emphasis id="B97AEAD4FFA7A716FF0476E5FEBCFB1E" bold="true" box="[164,281,1227,1246]" pageId="19" pageNumber="800">Figure 10.</emphasis>
<taxonomicName id="4C0E4D45FFA7A716FE8476E2FE79FB1E" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2016" baseAuthorityName="Monjaraz-Ruedas &amp; Francke" baseAuthorityYear="2016" box="[292,476,1228,1246]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="19" pageNumber="800" phylum="Arthropoda" rank="species" species="estor" status="comb. nov.">
<emphasis id="B97AEAD4FFA7A716FE8476E2FE79FB1E" box="[292,476,1228,1246]" italics="true" pageId="19" pageNumber="800">Mayazomus estor</emphasis>
</taxonomicName>
<emphasis id="B97AEAD4FFA7A716FE4676E5FDC0FB1E" bold="true" box="[486,613,1227,1246]" pageId="19" pageNumber="800">
<taxonomicNameLabel id="A24957AFFFA7A716FE4676E5FDC0FB1E" box="[486,613,1227,1246]" pageId="19" pageNumber="800" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
holotype male. Chelicera (A–C): A, mesal view. B, fixed finger. C, moveable finger. Paratype female. Spermathecae: D, dorsal view. Scale bars = 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BB136C6FFA7A716FF047701FE05F9B6" blockId="19.[164,779,1327,1655]" pageId="19" pageNumber="800">
<emphasis id="B97AEAD4FFA7A716FF047701FE38FA84" box="[164,413,1327,1348]" italics="true" pageId="19" pageNumber="800">
Flagellum (
<figureCitation id="13352A43FFA7A716FE827701FE35FA84" box="[290,400,1327,1348]" captionStart="Figure 9" captionStartId="18.[144,223,1730,1749]" captionTargetBox="[306,1258,199,1695]" captionTargetId="figure-0@18.[303,1263,194,1700]" captionTargetPageId="18" captionText="Figure 9. Mayazomus estor comb. nov. Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5359532" httpUri="https://zenodo.org/record/5359532/files/figure.png" pageId="19" pageNumber="800">Fig. 9E–G</figureCitation>
):
</emphasis>
Dorsoventrally flattened, cordate in shape; 2.08 times longer than wide, 3.00 times longer than pedicel length; with a single dorsosubmedian depression. Setation:
<emphasis id="B97AEAD4FFA7A716FE2477A5FE1FFA60" box="[388,442,1419,1440]" italics="true" pageId="19" pageNumber="800">Dm1</emphasis>
situated over the bulb base;
<emphasis id="B97AEAD4FFA7A716FF047787FF7EFA7E" box="[164,219,1449,1470]" italics="true" pageId="19" pageNumber="800">Dm4</emphasis>
situated medially;
<emphasis id="B97AEAD4FFA7A716FE197787FE47FA7E" box="[441,482,1449,1470]" italics="true" pageId="19" pageNumber="800">Dl1</emphasis>
at same level as
<emphasis id="B97AEAD4FFA7A716FD0C7787FD76FA7E" box="[684,723,1449,1470]" italics="true" pageId="19" pageNumber="800">Vl1</emphasis>
;
<emphasis id="B97AEAD4FFA7A716FD427787FCAEFA7E" box="[738,779,1449,1470]" italics="true" pageId="19" pageNumber="800">Dl3</emphasis>
posteriorly to
<emphasis id="B97AEAD4FFA7A716FEEA77E6FED4FA1D" box="[330,369,1480,1501]" italics="true" pageId="19" pageNumber="800">Vl2</emphasis>
; pair
<emphasis id="B97AEAD4FFA7A716FE1C77E6FE54FA1D" box="[444,497,1480,1501]" italics="true" pageId="19" pageNumber="800">Vm2</emphasis>
present; seta
<emphasis id="B97AEAD4FFA7A716FD3D77E6FD77FA1D" box="[669,722,1480,1501]" italics="true" pageId="19" pageNumber="800">Vm1</emphasis>
posteriorly to
<emphasis id="B97AEAD4FFA7A716FE8177C8FEF3FA3B" box="[289,342,1510,1531]" italics="true" pageId="19" pageNumber="800">Vm2</emphasis>
;
<emphasis id="B97AEAD4FFA7A716FEC677C9FE28FA3C" box="[358,397,1511,1532]" italics="true" pageId="19" pageNumber="800">Vm</emphasis>
5 posterior to
<emphasis id="B97AEAD4FFA7A716FD9377C8FDFFFA3B" box="[563,602,1510,1531]" italics="true" pageId="19" pageNumber="800">Vl1</emphasis>
. With one pair of antero-dorsal microsetae between
<emphasis id="B97AEAD4FFA7A716FDC0742BFD32F9DA" box="[608,663,1541,1562]" italics="true" pageId="19" pageNumber="800">Dm1</emphasis>
and
<emphasis id="B97AEAD4FFA7A716FD7A742BFCA6F9DA" box="[730,771,1541,1562]" italics="true" pageId="19" pageNumber="800">Dl1</emphasis>
, nearest to
<emphasis id="B97AEAD4FFA7A716FE84740DFEE8F9F8" box="[292,333,1571,1592]" italics="true" pageId="19" pageNumber="800">Dl1</emphasis>
; one pair of antero-lateral microsetae on flagellar pedicel; and with two pairs of microsetae between
<emphasis id="B97AEAD4FFA7A716FEAC744FFE96F9B6" box="[268,307,1633,1654]" italics="true" pageId="19" pageNumber="800">Vl1</emphasis>
and
<emphasis id="B97AEAD4FFA7A716FED2744FFE3CF9B6" box="[370,409,1633,1654]" italics="true" pageId="19" pageNumber="800">Vl2</emphasis>
.
</paragraph>
<paragraph id="8BB136C6FFA7A716FF04748EFC0AF955" blockId="19.[164,779,1696,1902]" lastBlockId="19.[826,1442,1327,1685]" pageId="19" pageNumber="800">
<emphasis id="B97AEAD4FFA7A716FF04748EFED3F975" box="[164,374,1696,1717]" italics="true" pageId="19" pageNumber="800">
Female (
<typeStatus id="54B58864FFA7A716FEA5748EFECDF975" box="[261,360,1696,1717]" pageId="19" pageNumber="800" type="paratype">paratype</typeStatus>
):
</emphasis>
Similar to male, except in pedipalp development and without the projected spur on the trochanter, pedipalp 1.88 times longer than propeltidium length, pedipalps similar to the homeomorphic male (
<figureCitation id="13352A43FFA7A716FF0C7535FEA5F8F1" box="[172,256,1819,1841]" captionStart="Figure 9" captionStartId="18.[144,223,1730,1749]" captionTargetBox="[306,1258,199,1695]" captionTargetId="figure-0@18.[303,1263,194,1700]" captionTargetPageId="18" captionText="Figure 9. Mayazomus estor comb. nov. Holotype homeomorphic male. A, pedipalp ectal view. Female. B, pedipalp ectal view. Paratype heteromorphic male. Pedipalp (C, D): C, ectal view. D, mesal view. Holotype male flagellum (E–G): E, dorsal view. F, ventral view. G, lateral view. Abbreviations: Fe, femur ectal; Fv, femur ventral; AP, apical process; Dm, dorsal median; Dl, dorsal lateral; Vm, ventral median; Vl, ventral lateral. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5359532" httpUri="https://zenodo.org/record/5359532/files/figure.png" pageId="19" pageNumber="800">Fig. 9B</figureCitation>
); setation as on male. Flagellum with three annuli. Setation: seta
<emphasis id="B97AEAD4FFA7A716FE3D7517FE60F88E" box="[413,453,1849,1870]" italics="true" pageId="19" pageNumber="800">Dl1</emphasis>
reduced and anterior to
<emphasis id="B97AEAD4FFA7A716FD7E7517FCA1F88E" box="[734,772,1849,1870]" italics="true" pageId="19" pageNumber="800">Vl1</emphasis>
;
<emphasis id="B97AEAD4FFA7A716FF047576FF68F8AD" box="[164,205,1880,1901]" italics="true" pageId="19" pageNumber="800">Dl3</emphasis>
posterior to
<emphasis id="B97AEAD4FFA7A716FEC67576FE28F8AD" box="[358,397,1880,1901]" italics="true" pageId="19" pageNumber="800">Vl2</emphasis>
;
<emphasis id="B97AEAD4FFA7A716FE3E7576FE76F8AD" box="[414,467,1880,1901]" italics="true" pageId="19" pageNumber="800">Vm2</emphasis>
present and reduced; seta
<emphasis id="B97AEAD4FFA7A716FC9A7701FCCBFA84" box="[826,878,1327,1348]" italics="true" pageId="19" pageNumber="800">Vm1</emphasis>
at same level as
<emphasis id="B97AEAD4FFA7A716FB8A7701FBF8FA84" box="[1066,1117,1327,1348]" italics="true" pageId="19" pageNumber="800">Vm2</emphasis>
. Segment III with one lateral pair of microsetae near to
<emphasis id="B97AEAD4FFA7A716FBCA7763FB05FAA2" box="[1130,1184,1357,1378]" italics="true" pageId="19" pageNumber="800">Vm4</emphasis>
. Segment IV with one pair of distolateral microsetae close to
<emphasis id="B97AEAD4FFA7A716FB757742FAA6FA41" box="[1237,1283,1388,1409]" italics="true" pageId="19" pageNumber="800">Dl3.</emphasis>
Spermathecae (
<figureCitation id="13352A43FFA7A716FCE277A4FC01FA61" box="[834,932,1418,1441]" captionStart="Figure 10" captionStartId="19.[164,243,1227,1246]" captionTargetBox="[164,1444,195,1197]" captionTargetId="figure-302@19.[164,1444,195,1197]" captionTargetPageId="19" captionText="Figure 10. Mayazomus estor comb. nov. holotype male. Chelicera (A–C): A, mesal view. B, fixed finger. C, moveable finger. Paratype female. Spermathecae: D, dorsal view. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5359534" httpUri="https://zenodo.org/record/5359534/files/figure.png" pageId="19" pageNumber="800">Fig. 10D</figureCitation>
) with four lobes; median pair curved along its length, subequal in length; lateral pair curved outwardly, both with duct openings. Gonopod long, wide. Chitinized arch V-shaped, posterior branch wide basally, with irregular margins, lateral tip not sclerotized. Chelicera: serrula with 18 teeth. Setal group formula 1:3, 2:6, 3:4, 4:2, 5:7, 6:1. Setal group 3 with two setae small and two larger, both feathered. Fixed finger with five teeth.
</paragraph>
<paragraph id="8BB136C6FFA7A716FC9A7491FC01F914" blockId="19.[826,1441,1727,1902]" box="[826,932,1727,1748]" pageId="19" pageNumber="800">
<emphasis id="B97AEAD4FFA7A716FC9A7491FC01F914" box="[826,932,1727,1748]" italics="true" pageId="19" pageNumber="800">Variation</emphasis>
</paragraph>
<paragraph id="8BB136C6FFA7A711FC9A74F3FEC1FE0F" blockId="19.[826,1441,1727,1902]" lastBlockId="20.[144,759,197,464]" lastPageId="20" lastPageNumber="801" pageId="19" pageNumber="800">
The homeomorphic male differs from the heteromorphic male mainly in the pedipalp size (total length of the pedipalp in heteromorphic male 2.42, and in homeomorphic male 1.8 times the length of the propeltidium); trochanter of the pedipalp without the spur project- ed; femur with setae
<emphasis id="B97AEAD4FFA0A711FE2572EBFE15FF1A" box="[389,432,197,218]" italics="true" pageId="20" pageNumber="801">Fv1</emphasis>
and
<emphasis id="B97AEAD4FFA0A711FE4B72EBFDB2FF1A" box="[491,535,197,218]" italics="true" pageId="20" pageNumber="801">Fv2</emphasis>
less developed; the development of the femoral apophysis is very poor in both
<typeStatus id="54B58864FFA0A711FF67732DFEA4FED8" box="[199,257,259,280]" pageId="20" pageNumber="801">types</typeStatus>
of male. However, based on the morphometric ratios, the development of the femoral apophysis, and the curvature of the patella, the male described by de Armas &amp; Víquez (2010) (and revised in this work) as a heteromorphic male is probably a pseudoheteromorphic male, as was reported for
<taxonomicName id="4C0E4D45FFA0A711FE0E73B2FDA8FE70" authorityName="Monjaraz-Ruedas &amp; Francke" authorityYear="2015" box="[430,525,411,433]" class="Arachnida" family="Hubbardiidae" genus="Mayazomus" kingdom="Animalia" order="Schizomida" pageId="20" pageNumber="801" phylum="Arthropoda" rank="species" species="loobil">
<emphasis id="B97AEAD4FFA0A711FE0E73B2FDA8FE70" box="[430,525,411,433]" italics="true" pageId="20" pageNumber="801">M. loobil</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9F4B37FFA0A711FD9673B5FEC5FE0F" author="Monjaraz-Ruedas R &amp; Francke OF" pageId="20" pageNumber="801" pagination="451 - 490" refId="ref16524" refString="Monjaraz-Ruedas R, Francke OF. 2015. Taxonomic revision of the genus Mayazomus Reddell &amp; Cokendolpher, 1995 (Schizomida: Hubbardiidae), with description of five new species from Chiapas, Mexico. Zootaxa 3915: 451 - 490." type="journal article" year="2015">Monjaraz-Ruedas &amp; Francke (2015)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB136C6FFA0A711FF3073D9FEBEFDCC" blockId="20.[144,549,503,555]" box="[144,283,503,524]" pageId="20" pageNumber="801">
<emphasis id="B97AEAD4FFA0A711FF3073D9FEBEFDCC" box="[144,283,503,524]" italics="true" pageId="20" pageNumber="801">Distribution</emphasis>
</paragraph>
<paragraph id="8BB136C6FFA0A711FF307038FD80FDEB" blockId="20.[144,549,503,555]" box="[144,549,533,555]" pageId="20" pageNumber="801">
Known only from the
<typeStatus id="54B58864FFA0A711FE327038FE66FDEB" box="[402,451,534,555]" pageId="20" pageNumber="801">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BB136C6FFA0A711FF30707DFEE4FDA8" blockId="20.[144,759,595,677]" box="[144,321,595,616]" pageId="20" pageNumber="801">
<emphasis id="B97AEAD4FFA0A711FF30707DFEE4FDA8" box="[144,321,595,616]" italics="true" pageId="20" pageNumber="801">Natural history</emphasis>
</paragraph>
<paragraph id="8BB136C6FFA0A711FF30705FFE82FD65" blockId="20.[144,759,595,677]" pageId="20" pageNumber="801">
The specimens were collected sympatric with
<taxonomicName id="4C0E4D45FFA0A711FDDF705FFE85FD65" authorityName="Chamberlin" authorityYear="1922" class="Arachnida" family="Hubbardiidae" genus="Stenochrus" kingdom="Animalia" order="Schizomida" pageId="20" pageNumber="801" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="B97AEAD4FFA0A711FDDF705FFE85FD65" italics="true" pageId="20" pageNumber="801">Stenochrus portoricensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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