treatments-xml/data/03/A7/87/03A78782FFB8E46180F44A2FFE41FE24.xml

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<document id="64D6B9882A41D52994844B68901DC1E4" ID="10.11646/zootaxa.4242.2.1SLASH10449" ID-CLB-Dataset="36308" ID-DOI="10.11646/zootaxa.4242.2.1" ID-GBIF-Dataset="35c61ca1-c858-4f10-baa2-b4b2b72d1454" ID-ISSN="1175-5326" ID-Zenodo-Dep="376356" ID-ZooBank="88B94383-F912-4BBD-B9F0-5642002C496D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489133894790" checkinUser="plazi" docAuthor="Gordon, Dennis P. &amp; Taylor, Paul D." docDate="2017" docId="03A78782FFB8E46180F44A2FFE41FE24" docLanguage="en" docName="zootaxa.4242.2.1SLASH10449.pdf" docOrigin="Zootaxa 4242 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Olisthella alma Gordon &amp; Taylor, 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="221" masterDocId="FF9EFFFAFFAAE47580634F56FFC6FFF4" masterDocTitle="Resolving the status of Pyriporoides and Daisyella (Bryozoa: Cheilostomata), with the systematics of some additional taxa of Calloporoidea having an ooecial heterozooid" masterLastPageNumber="232" masterPageNumber="201" pageNumber="219" updateTime="1698437807781" updateUser="ExternalLinkService">
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<mods:title id="3E34032BF98345822D0354C2672D75D4">Resolving the status of Pyriporoides and Daisyella (Bryozoa: Cheilostomata), with the systematics of some additional taxa of Calloporoidea having an ooecial heterozooid</mods:title>
</mods:titleInfo>
<mods:name id="8632A629F8A8BF9CF6FACFD3D43C2E90" type="personal">
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<mods:namePart id="9069F4E024D67CFDAEDE58F66AB6CD46">Gordon, Dennis P.</mods:namePart>
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<mods:namePart id="41044106A7A583CC7ADF0AAC1C5F8977">Taylor, Paul D.</mods:namePart>
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<mods:title id="AA119BC5ACA9756BF482BF3098463D8F">Zootaxa</mods:title>
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<mods:part id="C399C3F93A729147F1E4FDFA1D3B9279">
<mods:date id="88D5CCA293044C79A9EDB112C697F7EA">2017</mods:date>
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<mods:number id="379776B8FFB104FF10980B5549532EC4">4242</mods:number>
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<mods:identifier id="8437F38BF8A111F58ADD57D2494C1E77" type="DOI">10.11646/zootaxa.4242.2.1</mods:identifier>
<mods:identifier id="A2B16C65534C1070A05EBA902A6AAA73" type="GBIF-Dataset">35c61ca1-c858-4f10-baa2-b4b2b72d1454</mods:identifier>
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<treatment id="03A78782FFB8E46180F44A2FFE41FE24" ID-DOI="http://doi.org/10.5281/zenodo.6043886" ID-GBIF-Taxon="127843517" ID-Zenodo-Dep="6043886" LSID="urn:lsid:plazi:treatment:03A78782FFB8E46180F44A2FFE41FE24" httpUri="http://treatment.plazi.org/id/03A78782FFB8E46180F44A2FFE41FE24" lastPageId="20" lastPageNumber="221" pageId="18" pageNumber="219">
<subSubSection id="C314651FFFB8E46780F44A2FFF3EFA40" pageId="18" pageNumber="219" type="nomenclature">
<paragraph id="8BB13694FFB8E46780F44A2FFE5FFA67" blockId="18.[151,409,1401,1461]" box="[151,409,1401,1427]" pageId="18" pageNumber="219">
<heading id="D0F981F8FFB8E46780F44A2FFE5FFA67" bold="true" box="[151,409,1401,1427]" fontSize="11" level="1" pageId="18" pageNumber="219" reason="1">
<emphasis id="B97AEA86FFB8E46780F44A2FFE5FFA67" bold="true" box="[151,409,1401,1427]" pageId="18" pageNumber="219">
<taxonomicName id="4C0E4D17FFB8E46780F44A2FFE96FA67" ID-CoL="86NM2" box="[151,336,1401,1427]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="219" phylum="Bryozoa" rank="species" species="alma" status="sp. nov.">
<emphasis id="B97AEA86FFB8E46780F44A2FFE96FA67" bold="true" box="[151,336,1401,1427]" italics="true" pageId="18" pageNumber="219">Olisthella alma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957FDFFB8E46781344A2FFE5FFA67" box="[343,409,1401,1427]" pageId="18" pageNumber="219" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB13694FFB8E46780F44ACAFF3EFA40" blockId="18.[151,409,1401,1461]" box="[151,248,1436,1461]" pageId="18" pageNumber="219">
(
<figureCitation id="13352A11FFB8E46780FC4ACAFF29FA40" box="[159,239,1436,1461]" captionStart="FIGURE 11" captionStartId="19.[151,250,1900,1922]" captionTargetBox="[258,1323,338,1870]" captionTargetId="figure@19.[257,1323,337,1870]" captionTargetPageId="19" captionText="FIGURE 11. Olisthella alma n. gen., n. sp., A C, holotype, NIWA 98229; B, E, F, paratype, NIWA 98230; D, unregistered, on same substratum as paratype: A, zooids, one with an ooecium; B, zooids with well-preserved cryptocystal shelves; C, ooecium of zooid illustrated in A; D, zooid in profile; E, reparative kenozooid functioning as an opesial closure; F, complete autozooid with opesia occluded by reparative kenozooid. Scalebars: A, 500 µm; B, 200 µm; C, D, F, 100 µm; E, 50 µm." httpUri="https://zenodo.org/record/376367/files/figure.png" pageId="18" pageNumber="219">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C314651FFFB8E46780F44AB2FE7CF99C" pageId="18" pageNumber="219" type="materials_examined">
<paragraph id="8BB13694FFB8E46780F44AB2FE7CF99C" blockId="18.[151,1437,1508,2037]" pageId="18" pageNumber="219">
<materialsCitation id="3B663CC9FFB8E46780F44AB2FBBDF9B0" ID-GBIF-Occurrence="1438637077" collectionCode="NIWA" country="New Zealand" county="Motel" elevation="50" latitude="-45.1154" location="Waiareka Volcanic Formation" longLatPrecision="6" longitude="170.9074" municipality="Late Eocene" pageId="18" pageNumber="219" specimenCode="NIWA 98231" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
<emphasis id="B97AEA86FFB8E46780F44AB2FE47FA09" bold="true" box="[151,385,1508,1533]" pageId="18" pageNumber="219">Material examined.</emphasis>
<typeStatus id="54B58836FFB8E46781EB4AB2FE28FA08" box="[392,494,1508,1532]" pageId="18" pageNumber="219" type="holotype">
<emphasis id="B97AEA86FFB8E46781EB4AB2FE28FA08" box="[392,494,1508,1532]" italics="true" pageId="18" pageNumber="219">Holotype</emphasis>
</typeStatus>
:
<specimenCode id="DBA89EEFFFB8E467819D4AB2FD5EFA08" box="[510,664,1508,1532]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="18" pageNumber="219">
<collectionCode id="ED1FAE51FFB8E467819D4AB2FD81FA08" box="[510,583,1508,1532]" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="18" pageNumber="219">NIWA</collectionCode>
98231
</specimenCode>
,
<taxonomicName id="4C0E4D17FFB8E46782C74AB2FD23FA09" box="[676,741,1508,1533]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="221" phylum="Bryozoa" rank="species" species="alma" status="sp. nov.">Alma</taxonomicName>
,
<collectingRegion id="49CAF876FFB8E46782934AB2FCF1FA08" box="[752,823,1508,1532]" country="New Zealand" name="Otago" pageId="18" pageNumber="219">Otago</collectingRegion>
,
<collectingCountry id="F3197604FFB8E46783204AB3FC1CFA09" box="[835,986,1508,1533]" name="New Zealand" pageId="18" pageNumber="219">New Zealand</collectingCountry>
Fossil Record Number for outcrop J41/ f0088,
<location id="8ED1604FFFB8E46780844951FDFDF9D4" LSID="urn:lsid:plazi:treatment:03A78782FFB8E46180F44A2FFE41FE24:8ED1604FFFB8E46780844951FDFDF9D4" box="[231,571,1543,1568]" country="New Zealand" county="Motel" latitude="-45.1154" longLatPrecision="6" longitude="170.9074" municipality="Late Eocene" name="Waiareka Volcanic Formation" pageId="18" pageNumber="219" stateProvince="Otago">Waiareka Volcanic Formation</location>
,
<location id="8ED1604FFFB8E4678224495EFD7FF9D4" LSID="urn:lsid:plazi:treatment:03A78782FFB8E46180F44A2FFE41FE24:8ED1604FFFB8E4678224495EFD7FF9D4" box="[583,697,1544,1568]" country="New Zealand" county="Motel" latitude="-45.1154" longLatPrecision="6" longitude="170.9074" municipality="Late Eocene" name="Runangan" pageId="18" pageNumber="219" stateProvince="Otago">Runangan</location>
(Priabonian),
<collectingMunicipality id="6BD5ACEEFFB8E467833F495EFC2AF9EB" box="[860,1004,1544,1567]" pageId="18" pageNumber="219">Late Eocene</collectingMunicipality>
,
<geoCoordinate id="EE3A5053FFB8E467839B4951FBB0F9D4" box="[1016,1142,1543,1568]" direction="south" orientation="latitude" pageId="18" pageNumber="219" precision="5" value="-45.1154">45.1154° S</geoCoordinate>
,
<geoCoordinate id="EE3A5053FFB8E46784E7495EFAD2F9EB" box="[1156,1300,1544,1568]" direction="east" orientation="longitude" pageId="18" pageNumber="219" precision="5" value="170.9074">170.9074° E</geoCoordinate>
, from a tuff in a low road-cutting,
<elevation id="0023D1A7FFB8E46781F0497AFE0BF9B0" box="[403,461,1580,1605]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="219" unit="m" value="50.0">
<quantity id="4CF69B71FFB8E46781F0497AFE0BF9B0" box="[403,461,1580,1605]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="219" unit="m" value="50.0">50 m</quantity>
</elevation>
south of
<taxonomicName id="4C0E4D17FFB8E467825A497AFDB1F9B1" box="[569,631,1580,1605]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="221" phylum="Bryozoa" rank="species" species="alma" status="sp. nov.">Alma</taxonomicName>
<collectingCounty id="62D04E18FFB8E467821C497AFD04F9B1" box="[639,706,1580,1605]" pageId="18" pageNumber="219">Motel</collectingCounty>
on east side of main south road (SH1)
</materialsCitation>
.
<materialsCitation id="3B663CC9FFB8E46784E4497BFE71F99C" ID-GBIF-Occurrence="1438637092" collectionCode="NIWA" country="New Zealand" county="Motel" elevation="50" latitude="-45.1154" location="Waiareka Volcanic Formation" longLatPrecision="6" longitude="170.9074" municipality="Late Eocene" pageId="18" pageNumber="219" specimenCode="NIWA 98232" specimenCount="1" stateProvince="Otago" typeStatus="paratype">
<typeStatus id="54B58836FFB8E46784E4497BFB28F9B0" box="[1159,1262,1581,1604]" pageId="18" pageNumber="219" type="paratype">
<emphasis id="B97AEA86FFB8E46784E4497BFB28F9B0" box="[1159,1262,1581,1604]" italics="true" pageId="18" pageNumber="219">Paratype</emphasis>
</typeStatus>
:
<specimenCode id="DBA89EEFFFB8E467849D497AFA5EF9B0" box="[1278,1432,1580,1604]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="18" pageNumber="219">
<collectionCode id="ED1FAE51FFB8E467849D497AFA81F9B0" box="[1278,1351,1580,1604]" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="18" pageNumber="219">NIWA</collectionCode>
98232
</specimenCode>
, same locality as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C314651FFFB8E46780A44922FB88F979" box="[199,1102,1652,1677]" pageId="18" pageNumber="219" type="etymology">
<paragraph id="8BB13694FFB8E46780A44922FB88F979" blockId="18.[151,1437,1508,2037]" box="[199,1102,1652,1677]" pageId="18" pageNumber="219">
<emphasis id="B97AEA86FFB8E46780A44922FE88F979" bold="true" box="[199,334,1652,1677]" pageId="18" pageNumber="219">Etymology.</emphasis>
Alluding to the
<typeStatus id="54B58836FFB8E46782664923FDF3F979" box="[517,565,1653,1677]" pageId="18" pageNumber="219">type</typeStatus>
locality, the name used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C314651FFFB8E46680A449C1FBE7FEE9" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219" type="description">
<paragraph id="8BB13694FFB8E46680A449C1FBE7FEE9" blockId="18.[151,1437,1508,2037]" lastBlockId="19.[151,1436,151,285]" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<emphasis id="B97AEA86FFB8E46780A449C1FE9FF944" bold="true" box="[199,345,1687,1712]" pageId="18" pageNumber="219">Description.</emphasis>
Colonies spot-like, comprising up to 17 non-caudate contiguous autozooids; maximum spread of fertile colonies
<emphasis id="B97AEA86FFB8E467812849EBFE91F920" box="[331,343,1725,1748]" italics="true" pageId="18" pageNumber="219">c</emphasis>
.
<quantity id="4CF69B71FFB8E467810A49EAFE05F920" box="[361,451,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="219" unit="mm" value="1.7">1.7 mm</quantity>
, hence small and spot-like. Individual zooids oval [ZL 332456 (384); ZW 266367 (318)], arranged in quincunx with at least two pairs of lateral septula as well as one distal and one proximal. Gymnocyst encircling entire autozooid at colony margin, narrow from frontal view, especially laterally and distally, but zooid relatively high, steeply sloping to substratum. Cryptocyst comprising a very broad, flat shelf surrounded by a raised rim that is circular to oval; radial length of shelf midproximally 99152 (116) µm (i.e. same length as opesia), surface even, with indications of lineations trending more or less radially from rim towards opesia; texture in life uncertain owing to diagenesis. Opesia longer than wide, typically slightly wider in distal third than proximal third, with slight narrowing of middle third [OpL 99157 (116); OpW 76111 (90)]; distal margin evenly rounded, proximal margin weakly rounded to not quite straight. Eight articulated spines, extending around the distal half of the opesial margin only, the bases slightly broader proximad; only one additional spine on the gymnocyst laterally. No avicularia. Ooecium subglobular, more or less as wide as long or slightly wider, texture in life uncertain owing to irregularities in preserved surface, the inferred ooecial heterozooid completely concealed beneath it [OoL 178200 (191); OoW 176211 (196)]. Ancestrula not identified.
</paragraph>
</subSubSection>
<caption id="DF71661CFFB9E46680F4483AFB2FF82A" httpUri="https://zenodo.org/record/376367/files/figure.png" pageId="19" pageNumber="220" targetBox="[258,1323,338,1870]" targetPageId="19">
<paragraph id="8BB13694FFB9E46680F4483AFB2FF82A" blockId="19.[151,1436,1900,2014]" pageId="19" pageNumber="220">
<emphasis id="B97AEA86FFB9E46680F4483AFED8F875" bold="true" box="[151,286,1900,1922]" pageId="19" pageNumber="220">FIGURE 11.</emphasis>
<taxonomicName id="4C0E4D17FFB9E4668147483AFE07F876" box="[292,449,1900,1922]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="220" phylum="Bryozoa" rank="species" species="alma" status="gen. nov.">
<emphasis id="B97AEA86FFB9E4668147483AFE07F876" box="[292,449,1900,1922]" italics="true" pageId="19" pageNumber="220">Olisthella alma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957FDFFB9E46681A4483BFD95F876" box="[455,595,1901,1922]" pageId="19" pageNumber="220" rank="species">
<emphasis id="B97AEA86FFB9E46681A4483BFD95F876" bold="true" box="[455,595,1901,1922]" pageId="19" pageNumber="220">n. gen., n. sp.</emphasis>
</taxonomicNameLabel>
, A C, holotype, NIWA 98229; B, E, F, paratype, NIWA 98230; D, unregistered, on same substratum as paratype:
<emphasis id="B97AEA86FFB9E46681A448DDFE19F854" bold="true" box="[455,479,1931,1952]" pageId="19" pageNumber="220">A,</emphasis>
zooids, one with an ooecium;
<emphasis id="B97AEA86FFB9E466837648DDFCEAF854" bold="true" box="[789,812,1931,1952]" pageId="19" pageNumber="220">B,</emphasis>
zooids with well-preserved cryptocystal shelves;
<emphasis id="B97AEA86FFB9E466854548DDFAF9F855" bold="true" box="[1318,1343,1931,1953]" pageId="19" pageNumber="220">C,</emphasis>
ooecium of zooid illustrated in A;
<emphasis id="B97AEA86FFB9E46681FA48FCFE77F84B" bold="true" box="[409,433,1962,1983]" pageId="19" pageNumber="220">D,</emphasis>
zooid in profile;
<emphasis id="B97AEA86FFB9E466820748FCFDB3F84B" bold="true" box="[612,629,1962,1983]" pageId="19" pageNumber="220">E</emphasis>
, reparative kenozooid functioning as an opesial closure;
<emphasis id="B97AEA86FFB9E46684D948FCFB0DF84B" bold="true" box="[1210,1227,1962,1983]" pageId="19" pageNumber="220">F,</emphasis>
complete autozooid with opesia occluded by reparative kenozooid. Scalebars: A, 500 µm; B, 200 µm; C, D, F, 100 µm; E, 50 µm.
</paragraph>
</caption>
<subSubSection id="C314651FFFBEE46180A44FC1FE06FE7C" pageId="20" pageNumber="221" type="discussion">
<paragraph id="8BB13694FFBEE46180A44FC1FB90FEE9" blockId="20.[151,1437,151,464]" pageId="20" pageNumber="221">
<emphasis id="B97AEA86FFBEE46180A44FC1FEFDFF44" bold="true" box="[199,315,151,176]" pageId="20" pageNumber="221">Remarks.</emphasis>
This species is represented by three colonies, one with a complete ooecium, a second with to frontal broken ooecia, and a third non-ovicellate.
<taxonomicName id="4C0E4D17FFBEE461820E4FEAFCDCFF20" box="[621,794,188,212]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="221" phylum="Bryozoa" rank="species" species="alma" status="sp. nov.">
<emphasis id="B97AEA86FFBEE461820E4FEAFCDCFF20" box="[621,794,188,212]" italics="true" pageId="20" pageNumber="221">Olisthella alma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957FDFFBEE46183424FEAFC99FF21" box="[801,863,188,213]" pageId="20" pageNumber="221" rank="species">
<emphasis id="B97AEA86FFBEE46183424FEAFC99FF21" bold="true" box="[801,863,188,213]" pageId="20" pageNumber="221">n. sp.</emphasis>
</taxonomicNameLabel>
most resembles
<taxonomicName id="4C0E4D17FFBEE461847A4FEAFB5BFF20" box="[1049,1181,188,212]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="221" phylum="Bryozoa" rank="species" species="contigua" status="sp. nov.">
<emphasis id="B97AEA86FFBEE461847A4FEAFB5BFF20" box="[1049,1181,188,212]" italics="true" pageId="20" pageNumber="221">O. contigua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957FDFFBEE46184C74FEAFB24FF21" box="[1188,1250,188,213]" pageId="20" pageNumber="221" rank="species">
<emphasis id="B97AEA86FFBEE46184C74FEAFB24FF21" bold="true" box="[1188,1250,188,213]" pageId="20" pageNumber="221">n. sp.</emphasis>
</taxonomicNameLabel>
—the cryptocyst is broad and extensive and opesial shape is not very dissimilar, but zooids of
<taxonomicName id="4C0E4D17FFBEE46184774FB6FBB7FF0C" box="[1044,1137,224,248]" class="Gymnolaemata" family="Calloporidae" genus="Olisthella" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="221" phylum="Bryozoa" rank="species" species="alma" status="sp. nov.">
<emphasis id="B97AEA86FFBEE46184774FB6FBB7FF0C" box="[1044,1137,224,248]" italics="true" pageId="20" pageNumber="221">O. alma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957FDFFBEE46184194F89FB7DFF0C" box="[1146,1211,223,248]" pageId="20" pageNumber="221" rank="species">
<emphasis id="B97AEA86FFBEE46184194F89FB7DFF0C" bold="true" box="[1146,1211,223,248]" pageId="20" pageNumber="221">n. sp.</emphasis>
</taxonomicNameLabel>
are far smaller, the spine bases are not completely pericryptocystal and there is only one accessory spine.
</paragraph>
<paragraph id="8BB13694FFBEE46180A44E7EFE06FE7C" blockId="20.[151,1437,151,464]" pageId="20" pageNumber="221">
<bibRefCitation id="EF9F4B65FFBEE46180A44E7EFE4FFEB4" box="[199,393,296,320]" pageId="20" pageNumber="221" refString="Lee, D. E., Scholz, J. &amp; Gordon, D. P. (1997) Paleoecology of a Late Eocene mobile rockground biota from North Otago, New Zealand. Palaios, 12, 568 - 581." type="journal article">
Lee
<emphasis id="B97AEA86FFBEE46180984E7FFEE8FEB4" box="[251,302,296,320]" italics="true" pageId="20" pageNumber="221">et al</emphasis>
. (1997)
</bibRefCitation>
described the paleoecological setting as a warm-water (subtropical), mobile volcaniclastic (basaltic) pebble and cobble rockground in a current-swept channel about
<quantity id="4CF69B71FFBEE46183BC4E1AFB84FE90" box="[991,1090,332,357]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="20" pageNumber="221" unit="m" value="37.5" valueMax="50.0" valueMin="25.0">2550 m</quantity>
depth several kilometres from the Late Eocene shoreline.
</paragraph>
</subSubSection>
<subSubSection id="C314651FFFBEE46180A44EC2FE41FE24" pageId="20" pageNumber="221" type="distribution">
<paragraph id="8BB13694FFBEE46180A44EC2FE41FE24" blockId="20.[151,1437,151,464]" pageId="20" pageNumber="221">
<emphasis id="B97AEA86FFBEE46180A44EC2FEA6FE59" bold="true" box="[199,352,404,429]" pageId="20" pageNumber="221">Distribution.</emphasis>
Known only from the Runangan (Priabonian) stage at the
<typeStatus id="54B58836FFBEE46184694EC3FBFCFE59" box="[1034,1082,405,429]" pageId="20" pageNumber="221">type</typeStatus>
locality in north
<collectingRegion id="49CAF876FFBEE46185654EC2FA8BFE58" box="[1286,1357,404,428]" country="New Zealand" name="Otago" pageId="20" pageNumber="221">Otago</collectingRegion>
, South
<collectingCountry id="F3197604FFBEE46180F44EE1FF18FE24" box="[151,222,439,464]" name="Iceland" pageId="20" pageNumber="221">Island</collectingCountry>
,
<collectingCountry id="F3197604FFBEE461808A4EEEFE45FE24" box="[233,387,439,464]" name="New Zealand" pageId="20" pageNumber="221">New Zealand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>