280 lines
26 KiB
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280 lines
26 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.529.6139" ID-GBIF-Dataset="db8513fd-e2ce-4f63-a857-3530566adade" ID-PMC="PMC4668899" ID-Pensoft-Pub="1313-2970-529-1" ID-PubMed="26692792" ID-ZBK="AD4323B4913C447A88A7CE05EC8862A3" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-529-1" ModsDocOrigin="ZooKeys 529" ModsDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" checkinTime="1451243868489" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Budha, Prem B., Naggs, Fred, Backeljau, Thierry & Asami, Takahiro" docDate="2015" docId="4B2E75C0ED954B5635182B0FD969AC6D" docLanguage="en" docName="ZooKeys 529: 1-70" docOrigin="ZooKeys 529" docSource="http://dx.doi.org/10.3897/zookeys.529.6139" docTitle="Endothyrella nepalica Budha & Pall-Gergely, sp. n." docType="treatment" docUuid="1ED614EA-455F-4507-B1EF-F5129052F4E0" docUuidSource="ZooBank" docVersion="4" lastPageNumber="34" masterDocId="E206FFD8FFF5FFFCFFF16833B258FB6B" masterDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="70" masterPageNumber="1" pageNumber="34" updateTime="1668160861707" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Budha, Prem B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Naggs, Fred</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Backeljau, Thierry</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Asami, Takahiro</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>529</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.529.6139</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.529.6139</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-529-1</mods:identifier>
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<mods:identifier type="ZBK">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
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<mods:identifier type="ZooBank">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152062819" LSID="urn:lsid:zoobank.org:act:1ED614EA-455F-4507-B1EF-F5129052F4E0" httpUri="http://treatment.plazi.org/id/4B2E75C0ED954B5635182B0FD969AC6D" lastPageNumber="34" pageId="33" pageNumber="34">
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<subSubSection pageId="33" pageNumber="34" type="multiple">
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<paragraph pageId="33" pageNumber="34">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="nomenclature">
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<paragraph pageId="33" pageNumber="34">
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<taxonomicName LSID="http://zoobank.org/1ED614EA-455F-4507-B1EF-F5129052F4E0" authority="Budha & Pall-Gergely" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">
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Endothyrella nepalica Budha &
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="33" pageNumber="34">sp. n.</taxonomicNameLabel>
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Figures 6E, 8
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 9
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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, 24
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 25
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="reference_group">
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<paragraph pageId="33" pageNumber="34">
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<taxonomicName LSID="http://zoobank.org/1ED614EA-455F-4507-B1EF-F5129052F4E0" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
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2015
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella affinis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="affinis">Endothyrella affinis</taxonomicName>
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, - Budha et al., ZooKeys, 492: 18.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="type material">
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<paragraph pageId="33" pageNumber="34">Type material.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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Champadevi, Kirtipur, Kathmandu District, 1326-1500 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.654867">27.654868°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.24409">85.244084°E</geoCoordinate>
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, leg. Budha, P., 02.10.2010., holotype (CDZMTU005.1), paratypes CDZMTU005.2-16 (15 shells), CDZMTU005P (2 paratypes = specimens dissected and preserved, 3 dry shells = paratypes, 2 juvenile shells = not paratype); W-Nepal, Dhaulagiri Zone, Myagdi District, Annapurna Conservation Area, right side of Kali Gandaki valley, 300 m NNW of Suke Bagar village along
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<normalizedToken originalValue="“Tatopani-Dana”">"Tatopani-Dana"</normalizedToken>
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track, 1430 m alt., 14.05.1996., leg. A. Kuznetsov, WM/10 paratypes; Nepal, Kathmandu Valley, NW end of Kathmandu, middle part of S slope of Swoyambhunath Hill, in dry oak forest, 1500 m, 25.04.1995, leg A. Kuznetsov, WM/4 sinistral and 1 dextral paratypes; W Nepal, Daulagiri zone, Hyagdi distr., Annapurna NP., right side of Kali Gandaki v., NNW od Suke Bagar, Tatop, leg. A. Kuznetsov, 14.05.1996., ex coll. W. Maassen, HNHM 95867/1 paratype (labelled as paratype of "
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis nepalensis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalensis">Plectopylis nepalensis</taxonomicName>
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Schileyko and Kuznetsov"); Nepal, Swoyambhunath, Kathmandu District, 1366 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.71697">27.716971N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.28938">85.289386 E</geoCoordinate>
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, leg. Budha, P., 05.09.2008, CDZMTU006 (24 paratypes = shells); Siddha Cave, Tanahun District, 600 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.94718">27.94718°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="84.42134">84.421338°E</geoCoordinate>
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, leg. Budha, P., 24.10.2008, CDZMTU004, CDZMTU007 (11 paratypes = shells, and one juvenile shell, which is not paratype) (Figs 6E, 24A); Dhunche, Rasuwa, 1985 m,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="28.1092">28.1092°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="85.2916">85.2916°E</geoCoordinate>
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, leg. Budha, P., 31.05.2007., CDZMTU008 (2 shell = paratypes, and one damaged shell which is not paratype); Balaju, Kathmandu District, 1356 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.741173">27.741173°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.29376">85.293763°E</geoCoordinate>
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, leg. Budha, P., 04.01.2009., CDZMTU009 (8 paratypes = shells), CDZMTU009P (2 paratypes = specimens preserved, 4 dry shells = paratypes); Mahadevsthan, Thankot, Kathmandu District 1500 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.683367">27.683366°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.21384">85.213834°E</geoCoordinate>
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, leg. Budha, P., 06.02.2007., CDZMTU010 (25 paratypes = shells), CDZMTU010P 2 paratypes = specimens preserved, 4 dry shells = paratype, 5 juvenile shells = not paratypes); Arjewa, Baglung, 900 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.154392">28.154393°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.63071">83.630703°E</geoCoordinate>
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, leg. Budha, P., 13.09.2006., CDZMTU011 (14 paratypes = shells, one juvenile shells = not paratype); Majhbeni, Parbat, 700 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.205708">28.205708°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.67461">83.674605°E</geoCoordinate>
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, leg. Budha, P., 13.09.2006., CDZMTU012 (9 paratypes = shells, 6 juvenile/damaged shells = not paratypes); Sirsuwa, Parbat District, 780 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.136478">28.136478°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.642136">83.642135°E</geoCoordinate>
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, leg. Budha, P., 13.09.2006., CDZMTU013 (6 paratypes = shells); Foksing, Parbat District, 790 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.093252">28.093252°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.60429">83.604283°E</geoCoordinate>
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, leg. Budha, P., 11.06.2006., CDZMTU014 (11 paratypes = shells, 2 juvenile shells 7 not paratypes); Godawari, Lalitpur, 1868 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.94718">27.94718°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="84.42134">84.421338°E</geoCoordinate>
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, leg. Budha, P., 01.10.2008., CDZMTU015a (1 paratype); Annapurna Conservation Area, Tatopani, 1282 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.495173">28.495172°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.62888">83.628883°E</geoCoordinate>
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, leg. Budha, P., 01.10.2008., CDZMTU016 2 (2 paratypes = shells); Godawari, Lalitpur, 1575 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.596458">27.596459°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.389435">85.389432°E</geoCoordinate>
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, leg. Budha, P., 30.06.2007., CDZMTU015b (1 paratype = shell); Ridi, Gulmi, 832 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.945621">27.945621°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="83.43215">83.43215°E</geoCoordinate>
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, leg. Budha, P., 30.06.2007., CDZMTU017 (5 paratypes = shells); Godawari Botanical Garden, Lalitpur, 1453 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.59667">27.596671°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.38176">85.381758°E</geoCoordinate>
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, leg. Budha, P., 03.09.2008., CDZMTU015c (50 paratypes = shells); Nepal, Pokhara, Khare, 1520 m alt.,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="28.286">28.2860°N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="83.8472">83.8472°E</geoCoordinate>
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, leg. C. Huber, 18.03.1991, NMBE 527538/1 paratype (Figure 24C).
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</paragraph>
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<caption pageId="33" pageNumber="34">
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<paragraph pageId="33" pageNumber="34">
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Figure 24. Shells of
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">Endothyrella</taxonomicName>
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species. A
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
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Budha &
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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, sp. n., paratype, same data as on Fig. 6E B
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
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Budha &
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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, sp. n., holotype C
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
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Budha &
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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, sp. n., paratype, NMBE 527538 D
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella pinacis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pinacis">Endothyrella pinacis</taxonomicName>
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(Benson, 1859), (holotype of
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<taxonomicName class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea pinacis" order="Cycloneritida" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pinacis">pinacis</taxonomicName>
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) E
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella pinacis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pinacis">Endothyrella pinacis</taxonomicName>
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(holotype of
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix pettos" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pettos">Helix pettos</taxonomicName>
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) F
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella pinacis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pinacis">Endothyrella pinacis</taxonomicName>
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, NHMUK 1906.2.2.143. Photos: B.
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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(A), E. Bochud (B, C), J. Gundry (D), Ch. Zorn (E), H. Taylor (F). Scale represent 5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="diagnosis">
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<paragraph pageId="33" pageNumber="34">Diagnosis.</paragraph>
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<paragraph pageId="33" pageNumber="34">A small to middle-sized, hairless species with domed dorsal surface and rounded body whorl; parietal lamella simple with one or two denticles posteriorly and sometimes a plica below the lamella, middle palatal plicae divided or almost divided.</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="description">
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<paragraph pageId="33" pageNumber="34">Description.</paragraph>
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<paragraph pageId="33" pageNumber="34">Shell very small to small, sinistral, with somewhat elevated spire and domed dorsal surface; protoconch slightly elevates from the dorsal surface; usually brownish but sometimes turns into yellowish; protoconch consists of 1.5-1.75 whorls, very finely, regularly ribbed; teleoconch with very weak, irregular growth lines on the ventral surface and fine reticulated sculpture on the dorsal surface; in high magnification the surface is covered by flat periostracal folds; no spirally arranged large deciduous folds found; whorls 5.5-6.25, moderately bulging, separated by relatively deep suture; umbilicus wide and deep, whorls almost flat inside, resulting in an funnel-like shape, apertural lip whitish, rather thin, slightly reflexed; callus inconspicuous, but present, slightly S-shaped; no fold in the aperture.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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Ten specimens were opened from different populations. Parietal wall with one slightly curved lamella with arms pointing in the direction of the aperture; lower end on the lamella more conspicuously curved than the upper end; two small denticles above and below posteriorly of the lamella (exceptionally, the lower one is missing); in some populations (e.g. Majhbeni - Parbat District, Champadevi - Kathmandu District and Siddha Cave - Tanahu District) with short plica under the lamella; palatal wall with six plicae; first slim and short, parallel with the suture; second plica is the longest, it shows a tendency towards dividing in the middle, but the two parts always fused; third, fourth and fifth plicae usually divided (third one sometimes not); last plica short, slightly curved with arms pointing in the direction of the lower suture (Figures 9
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="measurements">
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<paragraph pageId="33" pageNumber="34">Measurements</paragraph>
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<paragraph pageId="33" pageNumber="34">(in mm): D: 8.2-14.9, H: 4.0-6.0, Wh: 5.5-7.5 (n = 35, different populations).</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="differential diagnosis">
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<paragraph pageId="33" pageNumber="34">Differential diagnosis.</paragraph>
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<paragraph pageId="33" pageNumber="34">
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
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sp. n. is usually larger than
|
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|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella angulata" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="angulata">Endothyrella angulata</taxonomicName>
|
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sp. n., it has a domed dorsal surface, rounded body whorl and lacks hairs standing in spiral rows, whereas
|
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella angulata" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="angulata">Endothyrella angulata</taxonomicName>
|
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sp. n. has a flat dorsal surface, shouldered body whorl and has hairs which are arranged in spiral rows.
|
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<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella dolakhaensis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="dolakhaensis">Endothyrella dolakhaensis</taxonomicName>
|
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|
sp. n. differs from
|
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|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
|
|||
|
sp. n. by the usually smaller size, fewer whorls, stronger sculpture, comparatively larger protoconch, conical dorsal surface, slightly angulated body whorl and the presence of hairs standing in five spiral lines. For comparison with
|
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|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella oakesi" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="oakesi">Endothyrella oakesi</taxonomicName>
|
|||
|
and
|
|||
|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella pinacis" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pinacis">Endothyrella pinacis</taxonomicName>
|
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|
, see under those species. See also Table 5.
|
|||
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</paragraph>
|
|||
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</subSubSection>
|
|||
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<subSubSection pageId="33" pageNumber="34" type="description of the genitalia">
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<paragraph pageId="33" pageNumber="34">Description of the genitalia</paragraph>
|
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<paragraph pageId="33" pageNumber="34">
|
|||
|
(Figures 25
|
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
|||
|
): Three specimens from three populations were anatomically examined (Champadevi, Balaju of Kathmandu District and Godawari Botanical Garden, Lalitpur District). Penis short, narrow distally and slowly tapers toward the proximal end; internal surface with several tubercles including minute calcareous hooks; epiphallus slender, cylindrical, longer than the penis, it enters penis laterally; penial caecum very short, blunt, cylindrical, with a short retractor muscle attached at its proximal end; vas deferens thin and nearly 1.5 times longer than epiphallus, convoluted before connection to prostate; vagina shorter than the penis with well-developed vaginal bulb; gametolytic sac very thin throughout and ends into a small rounded sac; there is a slender diverticulum running parallel with the gametolytic sac; it is as long as the gametolytic sac.
|
|||
|
</paragraph>
|
|||
|
<caption pageId="33" pageNumber="34">
|
|||
|
<paragraph pageId="33" pageNumber="34">
|
|||
|
Figure 25. Genital anatomy of
|
|||
|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
|
|||
|
sp. n. A Specimen from Godawari B, C penis of a specimen form Balaju. Diagrammatic. Abbreviations: AG albumen glandD diverticulumE epiphallusGS gametolytic sacP penisPC penial caecumRM retractor muscleSO spermoviductV vaginaVD vas deferens.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection pageId="33" pageNumber="34" type="etymology">
|
|||
|
<paragraph pageId="33" pageNumber="34">Etymology.</paragraph>
|
|||
|
<paragraph pageId="33" pageNumber="34">
|
|||
|
The name
|
|||
|
<taxonomicName class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea nepalica" order="Cycloneritida" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">nepalica</taxonomicName>
|
|||
|
refers to the country (Nepal) where the new species lives.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection pageId="33" pageNumber="34" type="type locality">
|
|||
|
<paragraph pageId="33" pageNumber="34">Type locality.</paragraph>
|
|||
|
<paragraph pageId="33" pageNumber="34">
|
|||
|
Champadevi, Kirtipur, Kathmandu District, Nepal, 1326-1500 m,
|
|||
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.654867">27.654868°N</geoCoordinate>
|
|||
|
,
|
|||
|
<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.24409">85.244084°E</geoCoordinate>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection pageId="33" pageNumber="34" type="distribution">
|
|||
|
<paragraph pageId="33" pageNumber="34">Distribution.</paragraph>
|
|||
|
<paragraph pageId="33" pageNumber="34">
|
|||
|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
|
|||
|
sp. n. inhabits a relatively large area in western and central Nepal (Figure 15).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection pageId="33" pageNumber="34" type="remarks">
|
|||
|
<paragraph pageId="33" pageNumber="34">Remarks.</paragraph>
|
|||
|
<paragraph pageId="33" pageNumber="34">
|
|||
|
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="50" pageNumber="51" pagination="435 - 564" title="Treatise on Recent Terrestrial Pulmonate Molluscs, Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimulidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae." volume="2" year="1999">Schileyko (1999)</bibRefCitation>
|
|||
|
figured a shell from the "SW slope of Swayambhunat (= Swoyambhunath) hill, Kathmandu valley, Nepal" (Fig. 594.). The figured specimen is probably
|
|||
|
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
|
|||
|
sp. n., but the drawing is not sufficient for identification.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|