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<document id="912A68F87F084272ACA44852258D8179" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB2D5751FF6EDB00FB20F845" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) foliiseta Duda (Duda 1923" docType="treatment" docVersion="10" lastPageNumber="12" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="12" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AF0190B8B0D0836BB46ADE8896763962">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="C83E1804B816B31108D3C7959C0CC4C8">Toda, Masanori J.</mods:namePart>
<mods:affiliation id="D550C25DF96822C4070A3470DCAAD234">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="35C782E302FE63472AC60D3F1C12BF49">Bänziger, Hans</mods:namePart>
<mods:affiliation id="75B144E65BA40470C9BE3FE88FC8F8AE">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="AEC1815721C53C535C2FAC4243A94796">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="49CA88CE836C7AAA90FF78F6176DDA0B">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="FDBD23716C13D2FDA401E42D0CAF73A8">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="06ECE9386DDAD22285511F010774CAFA">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="017F76E1BE76CB29A4349E4C2D08B88A">Suwito, Awit</mods:namePart>
<mods:affiliation id="552EE5B99553EB7AADECD49A49DBC6AB">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="678DD109320710B886AA0B2FD2769959">Katoh, Toru</mods:namePart>
<mods:affiliation id="78C57C6F7443D89715EF7EE3D59E3E72">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<treatment id="03C86C16EB2D5751FF6EDB00FB20F845" ID-DOI="http://doi.org/10.5281/zenodo.5221276" ID-GBIF-Taxon="164612798" ID-Zenodo-Dep="5221276" LSID="urn:lsid:plazi:treatment:03C86C16EB2D5751FF6EDB00FB20F845" httpUri="http://treatment.plazi.org/id/03C86C16EB2D5751FF6EDB00FB20F845" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="C37B8E8BEB2D5751FF6EDB00FD06F988" box="[151,749,1543,1570]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BDEDD00EB2D5751FF6EDB00FD06F988" blockId="11.[151,749,1543,1570]" box="[151,749,1543,1570]" pageId="11" pageNumber="12">
<heading id="D0966A6CEB2D5751FF6EDB00FD06F988" bold="true" box="[151,749,1543,1570]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B9150112EB2D5751FF6EDB00FD06F988" bold="true" box="[151,749,1543,1570]" pageId="11" pageNumber="12">
2.
<taxonomicName id="4C61A683EB2D5751FF4CDB0FFE17F988" ID-CoL="4GS4N" authorityName="Duda (Duda" authorityYear="1923" box="[181,508,1543,1570]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
<emphasis id="B9150112EB2D5751FF4CDB0FFEF6F988" bold="true" box="[181,285,1543,1570]" italics="true" pageId="11" pageNumber="12">Phortica</emphasis>
(
<emphasis id="B9150112EB2D5751FED7DB0FFE60F988" bold="true" box="[302,395,1543,1570]" italics="true" pageId="11" pageNumber="12">Ashima</emphasis>
)
<emphasis id="B9150112EB2D5751FE65DB0FFE17F988" bold="true" box="[412,508,1543,1570]" italics="true" pageId="11" pageNumber="12">foliiseta</emphasis>
</taxonomicName>
species group, new
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB2D5751FF6EDB40FB20F845" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BDEDD00EB2D5751FF6EDB40FE66F92D" blockId="11.[151,1437,1608,2031]" pageId="11" pageNumber="12">
<emphasis id="B9150112EB2D5751FF6EDB40FEF9F9C9" bold="true" box="[151,274,1608,1635]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Pregonital, apical knob with fine serrations (ch.45-1;
<figureCitation id="135AC185EB2D5751FC84DB41FC32F9C9" box="[893,985,1609,1635]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4C</figureCitation>
,
<figureCitation id="135AC185EB2D5751FC1EDB40FBE1F9C9" box="[999,1034,1608,1635]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5D</figureCitation>
) and round barbule (ch.46-0, 47-1;
<figureCitation id="135AC185EB2D5751FF6EDB65FF19F92D" box="[151,242,1645,1671]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4C</figureCitation>
,
<figureCitation id="135AC185EB2D5751FF04DB64FEC9F92D" box="[253,290,1644,1671]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FED4DB65FEA4F92D" box="[301,335,1645,1671]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FEA3DB65FE6BF92D" box="[346,384,1645,1671]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7G</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB2D5751FF3EDB98FD91F861" blockId="11.[151,1437,1608,2031]" pageId="11" pageNumber="12">
<emphasis id="B9150112EB2D5751FF3EDB98FE41F901" bold="true" box="[199,426,1680,1707]" pageId="11" pageNumber="12">Shared characters.</emphasis>
Palpus with hollow-shaped sense organ (ch.9-2). Some anterior, cibarial, medial sensilla short, apically blunt, peg-like (ch.11-1). Pseudotracheae in labellum 6 per side (ch.14-2). Mid and hind femora and hind tibia entirely yellow (ch.16,19-0). Epandrium widely round on ventral margin (ch.31-1;
<figureCitation id="135AC185EB2D5751FB51DBD1FAE9F959" box="[1192,1282,1753,1779]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4B</figureCitation>
,
<figureCitation id="135AC185EB2D5751FAF4DBD0FADBF959" box="[1293,1328,1752,1779]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5C</figureCitation>
,
<figureCitation id="135AC185EB2D5751FAC5DBD1FA8BF959" box="[1340,1376,1753,1779]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FA95DBD1FA64F959" box="[1388,1423,1753,1779]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7E</figureCitation>
); setae on and/or near ventral margin of epandrium not denser than those on other portions (ch.32-0;
<figureCitation id="135AC185EB2D5751FAF2DBF5FA8CF8BD" box="[1291,1383,1789,1815]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4B</figureCitation>
,
<figureCitation id="135AC185EB2D5751FA8CDBF4FA73F8BD" box="[1397,1432,1788,1815]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5C</figureCitation>
,
<figureCitation id="135AC185EB2D5751FF6EDA29FF57F891" box="[151,188,1825,1851]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FF31DA29FF00F891" box="[200,235,1825,1851]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7E</figureCitation>
). Hypandrium symmetric (ch.40-0;
<figureCitation id="135AC185EB2D5751FD78DA29FD35F891" box="[641,734,1825,1851]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FD13DA28FCE7F891" box="[746,780,1824,1851]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FCE1DA29FCD6F891" box="[792,829,1825,1851]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6G</figureCitation>
,
<figureCitation id="135AC185EB2D5751FCB3DA29FC84F891" box="[842,879,1825,1851]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7H</figureCitation>
). Pregonite (
<figureCitation id="135AC185EB2D5751FBF9DA29FBB3F891" box="[1024,1112,1825,1851]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Fig. 4C</figureCitation>
,
<figureCitation id="135AC185EB2D5751FB9DDA28FB62F891" box="[1124,1161,1824,1851]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FB6FDA29FB5CF891" box="[1174,1207,1825,1851]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FB3DDA29FB0CF891" box="[1220,1255,1825,1851]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7G</figureCitation>
) partly strongly sclerotized (ch.43-0), rod-like, with apical knob (ch.44-1) and sensilla only on basal portion (ch.53-1) but no serration on ventrodistal portion (ch.50-0). Aedeagal, outer membrane without apical, long, narrow tube (ch.57-0;
<figureCitation id="135AC185EB2D5751FA95DA61FF57F80D" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">Figs 5D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FF30DA85FF01F80D" box="[201,234,1933,1959]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FF0EDA85FEF6F80D" box="[247,285,1933,1959]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7G</figureCitation>
); aedeagal free sclerite absent (ch.59-0;
<figureCitation id="135AC185EB2D5751FD10DA85FCB4F80D" box="[745,863,1932,1959]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">Figs 5D,E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FC95DA85FC43F80D" box="[876,936,1933,1959]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6E,G</figureCitation>
,
<figureCitation id="135AC185EB2D5751FC4FDA85FC1CF80D" box="[950,1015,1933,1959]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7G,H</figureCitation>
). Aedeagal sheath and/or postgonites asymmetric (ch.63-1;
<figureCitation id="135AC185EB2D5751FE77DAB9FE01F861" box="[398,490,1969,1995]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="11" pageNumber="12">Figs 4D</figureCitation>
,
<figureCitation id="135AC185EB2D5751FE0CDAB8FDFCF861" box="[501,535,1968,1995]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="11" pageNumber="12">5E</figureCitation>
,
<figureCitation id="135AC185EB2D5751FDDBDAB9FDAAF861" box="[546,577,1969,1995]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen &amp; Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="11" pageNumber="12">6F</figureCitation>
,
<figureCitation id="135AC185EB2D5751FDB5DAB9FD87F861" box="[588,620,1969,1995]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="11" pageNumber="12">7F</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB2D5751FF3EDADDFB20F845" blockId="11.[151,1437,1608,2031]" box="[199,1227,2005,2031]" pageId="11" pageNumber="12">
<emphasis id="B9150112EB2D5751FF3EDADDFE99F845" bold="true" box="[199,370,2005,2031]" pageId="11" pageNumber="12">Included taxa.</emphasis>
<taxonomicName id="4C61A683EB2D5751FE80DADDFD41F845" authorityName="Duda (Duda" authorityYear="1923" box="[377,682,2005,2031]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
<emphasis id="B9150112EB2D5751FE80DADDFE30F845" box="[377,475,2005,2031]" italics="true" pageId="11" pageNumber="12">Phortica</emphasis>
(
<emphasis id="B9150112EB2D5751FE15DADDFDABF845" box="[492,576,2005,2031]" italics="true" pageId="11" pageNumber="12">Ashima</emphasis>
)
<emphasis id="B9150112EB2D5751FDA9DADDFD41F845" box="[592,682,2005,2031]" italics="true" pageId="11" pageNumber="12">foliiseta</emphasis>
</taxonomicName>
species subgroup; and
<emphasis id="B9150112EB2D5751FC56DADEFC2BF845" box="[943,960,2006,2031]" italics="true" pageId="11" pageNumber="12">P</emphasis>
.
<taxonomicName id="4C61A683EB2D5751FC36DADDFBBFF845" box="[975,1108,2005,2031]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tanabei" subGenus="Ashima">
(
<emphasis id="B9150112EB2D5751FC21DADDFC02F845" box="[984,1001,2005,2031]" italics="true" pageId="11" pageNumber="12">A</emphasis>
.)
<emphasis id="B9150112EB2D5751FBF9DADDFBBFF845" box="[1024,1108,2005,2031]" italics="true" pageId="11" pageNumber="12">tanabei</emphasis>
</taxonomicName>
subgroup.
</paragraph>
</subSubSection>
</treatment>
</document>
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