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<document id="1F858399B02D814C64203648494B9944" ID-DOI="10.11646/zootaxa.3741.1.2" ID-GBIF-Dataset="6a6ba78e-a9c5-4d11-997a-3d06e5e31520" ID-ISSN="1175-5326" ID-Zenodo-Dep="223773" ID-ZooBank="BF5EA9B8-C6F4-448A-BEF9-1976AB4EC308" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460702326177" checkinUser="plazi" docAuthor="Lazányi, Eszter &amp; Vagalinski, Boyan" docDate="2013" docId="03C887D3FFFFFFB0FF3499B1FE27ADC4" docLanguage="en" docName="zt03741p100.pdf" docOrigin="Zootaxa 3741 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum metsovoni Strasser 1976" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FFF1FFABFFF9FFB7FFA39926FFF2AD11" masterDocTitle="Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)" masterLastPageNumber="100" masterPageNumber="55" pageNumber="61" updateTime="1698356536038" updateUser="plazi">
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<mods:title id="81C4A76C50CB6FA37724FF6E786357B1">Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="E5DE32033A640015D0653A3A6EE006F8">Lazányi, Eszter</mods:namePart>
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<mods:namePart id="D65CB5F14FF358AB5E8CA79BBC78D3B6">Vagalinski, Boyan</mods:namePart>
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<mods:date id="2E1AF49E3C123A839B5A2112FFDC7524">2013</mods:date>
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<treatment id="03C887D3FFFFFFB0FF3499B1FE27ADC4" ID-DOI="http://doi.org/10.5281/zenodo.6151705" ID-GBIF-Taxon="119583503" ID-Zenodo-Dep="6151705" LSID="urn:lsid:plazi:treatment:03C887D3FFFFFFB0FF3499B1FE27ADC4" httpUri="http://treatment.plazi.org/id/03C887D3FFFFFFB0FF3499B1FE27ADC4" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
<subSubSection id="C37B654EFFFFFFB1FF3499B1FCE8AC09" pageId="6" pageNumber="61" type="nomenclature">
<paragraph id="8BDE36C5FFFFFFB1FF3499B1FD74ADA3" blockId="6.[151,646,151,211]" box="[151,646,151,178]" pageId="6" pageNumber="61">
<heading id="D09681A9FFFFFFB1FF3499B1FD74ADA3" bold="true" box="[151,646,151,178]" fontSize="11" level="1" pageId="6" pageNumber="61" reason="1">
<taxonomicName id="4C614D46FFFFFFB1FF3499B1FD74ADA3" ID-CoL="8KR6R" authority="Strasser, 1976" authorityName="Strasser" authorityYear="1976" box="[151,646,151,178]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="metsovoni">
<emphasis id="B915EAD7FFFFFFB1FF3499B1FD74ADA3" bold="true" box="[151,646,151,178]" pageId="6" pageNumber="61">
<emphasis id="B915EAD7FFFFFFB1FF3499B1FE32ADA0" bold="true" box="[151,448,151,177]" italics="true" pageId="6" pageNumber="61">Megaphyllum metsovoni</emphasis>
(Strasser, 1976)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDE36C5FFFFFFB1FF34999CFEF2ADC2" blockId="6.[151,646,151,211]" box="[151,256,186,211]" pageId="6" pageNumber="61">
<figureCitation id="135A2A40FFFFFFB1FF34999CFEF2ADC2" box="[151,256,186,211]" captionStart="FIGURES 3 a f" captionStartId="6.[151,264,1559,1581]" captionTargetBox="[197,1391,396,1535]" captionTargetId="figure@6.[191,1396,394,1537]" captionTargetPageId="6" captionText="FIGURES 3 a f. Megaphyllum metsovoni male from Eptolofos, Parnassos Mts, Greece. (3 a) Gonopods, posterior view; (3 b) right gonopods, mesal view; (3 c) left promere, meso-posterior view; (3 d) left opisthomere, lateral view; penis, (3 e) posterior and (3 f) right lateral views. Abbreviations: ap: anterior process, asp: anterior process of the solenomere, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223776/files/figure.png" pageId="6" pageNumber="61">Figs 3af</figureCitation>
</paragraph>
<paragraph id="8BDE36C5FFFFFFB1FF349824FCE8AC09" blockId="6.[151,1096,258,342]" box="[151,794,258,280]" pageId="6" pageNumber="61">
<taxonomicName id="4C614D46FFFFFFB1FF349824FD91AC09" authority="Strasser, 1976: 614" authorityName="Strasser" authorityPageNumber="614" authorityYear="1976" box="[151,611,258,280]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="metsovoni">
<emphasis id="B915EAD7FFFFFFB1FF349824FE68AC09" box="[151,410,258,280]" italics="true" pageId="6" pageNumber="61">Chromatoiulus metsovoni</emphasis>
Strasser, 1976: 614
</taxonomicName>
616, Figs 5256.
</paragraph>
</subSubSection>
<subSubSection id="C37B654EFFFFFFB1FF349807FBBAAC47" pageId="6" pageNumber="61" type="description">
<paragraph id="8BDE36C5FFFFFFB1FF349807FDA8AC26" blockId="6.[151,1096,258,342]" box="[151,602,289,311]" pageId="6" pageNumber="61">
<taxonomicName id="4C614D46FFFFFFB1FF349807FDA5AC26" authority="Ceuca 1992: 425" authorityName="Ceuca" authorityPageNumber="425" authorityYear="1992" box="[151,599,289,311]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="metsovoni">
<emphasis id="B915EAD7FFFFFFB1FF349807FE68AC26" box="[151,410,289,311]" italics="true" pageId="6" pageNumber="61">Chromatoiulus metsovoni</emphasis>
: Ceuca 1992: 425
</taxonomicName>
.
</paragraph>
<paragraph id="8BDE36C5FFFFFFB1FF349866FBBAAC47" blockId="6.[151,1096,258,342]" box="[151,1096,320,342]" pageId="6" pageNumber="61">
<taxonomicName id="4C614D46FFFFFFB1FF349866FD71AC47" authority="Enghoff &amp; Kime 2009" authorityName="Enghoff &amp; Kime" authorityYear="2009" box="[151,643,320,342]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="metsovoni">
<emphasis id="B915EAD7FFFFFFB1FF349866FE7CAC47" box="[151,398,320,342]" italics="true" pageId="6" pageNumber="61">Megaphyllum metsovoni</emphasis>
: Enghoff &amp; Kime 2009
</taxonomicName>
; Lazányi
<emphasis id="B915EAD7FFFFFFB1FD469867FCEBAC47" box="[741,793,320,342]" italics="true" pageId="6" pageNumber="61">et al.</emphasis>
2012: 17, 19, 41,
<figureCitation id="135A2A40FFFFFFB1FC739866FBB7AC47" box="[976,1093,320,342]" captionStart-0="FIGURES 10 a g" captionStart-1="FIGURES 11 a g" captionStart-2="FIGURES 12 a g" captionStart-3="FIGURES 13 a f" captionStart-4="FIGURES 14 a g" captionStartId-0="19.[151,264,1833,1855]" captionStartId-1="20.[151,264,1683,1705]" captionStartId-2="22.[151,264,1861,1883]" captionStartId-3="24.[151,264,1853,1875]" captionStartId-4="26.[151,264,1570,1592]" captionTargetBox-0="[178,1409,801,1810]" captionTargetBox-1="[257,1326,640,1661]" captionTargetBox-2="[166,1422,633,1831]" captionTargetBox-3="[176,1412,609,1819]" captionTargetBox-4="[163,1419,377,1544]" captionTargetId-0="figure@19.[174,1413,797,1811]" captionTargetId-1="figure@20.[252,1334,637,1662]" captionTargetId-2="figure@22.[162,1425,625,1840]" captionTargetId-3="figure@24.[168,1419,605,1832]" captionTargetId-4="figure@26.[163,1423,373,1548]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="22" captionTargetPageId-3="24" captionTargetPageId-4="26" captionText-0="FIGURES 10 a g. Megaphyllum glossulifer males and a female from Rhodopi Mts, Bulgaria: (10 a) gonopods, posterior view; (10 b) left gonopods, mesal view; (10 c) left promere, meso-posterior view; (10 d) left opisthomere, lateral view; penis, (10 e) posterior and (10 f) left lateral views; (10 g) left vulva, meso-posterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-1="FIGURES 11 a g. Megaphyllum monticola. Males from Korakas Mts, Greece, ZMB 1300: (11 a) gonopods, posterior view and (11 b) right gonopods, mesal view and (11 c) left promere, meso-posterior view. Males from Ostrovo, Greece, Inv. No. 8019: (11 d) Right opisthomere, lateral view, penis, (11 e) posterior and (11 f) right lateral views, (11 f *) another, bent penis, left lateral view. (11 g) Left vulva, posterior view, female from Korakas Mts, Greece (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-2="FIGURES 12 a g. Megaphyllum bosniense. Syntype male from Bosnia, ZMB 2625: (12 a) gonopods, posterior view; (12 b) left gonopods, mesal view; (12 c) right promere, meso-posterior view; (12 d) left opisthomere, meso-posterior view. Male from Treskavac Mts, Montenegro: penis, (12 e) posterior and (12 f) right lateral views. Female from Maljen Mts, Serbia: (12 g) left vulva, meso-posterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-3="FIGURES 13 a f. Megaphyllum rosenauense. Syntype male from Transylvania, Romania, ZMB 2622: (13 a) gonopods, posterior view; (13 b) right gonopods, mesal view; (13 c) right promere, posterior view; (13 d) right opisthomere, lateral view. Male from Miercurea Ciuc, Romania: penis, (13 e) posterior and (13 f) right lateral views. Abbreviations: asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-4="FIGURES 14 a g. Megaphyllum transsylvanicum and Megaphyllum kievenese. M. transsylvanicum syntype male from Transylvania, Romania, ZMB 4380: (14 a) gonopods, posterior view; (14 b) left gonopods, mesal view; (14 c) left promere, meso-posterior view. M. transsylvanicum male from Dobrostan, Bulgaria: (14 d) left opisthomere, antero-lateral view. M. kievense male, after Lohmander (1928): (14 d *) opisthomere. M. transsylvanicum. male from Fruska Gora, Serbia: penis, (14 e) posterior and (14 f) rig
.
</paragraph>
</subSubSection>
<caption id="DF1E664DFFFFFFB1FF349F31FC8FAB98" httpUri="https://zenodo.org/record/223776/files/figure.png" pageId="6" pageNumber="61" targetBox="[197,1391,396,1535]" targetPageId="6">
<paragraph id="8BDE36C5FFFFFFB1FF349F31FC8FAB98" blockId="6.[151,1436,1559,1673]" pageId="6" pageNumber="61">
<emphasis id="B915EAD7FFFFFFB1FF349F31FEB6AB3C" bold="true" box="[151,324,1559,1581]" pageId="6" pageNumber="61">FIGURES 3af.</emphasis>
<taxonomicName id="4C614D46FFFFFFB1FEEF9F31FDB7AB3C" box="[332,581,1559,1581]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="metsovoni">
<emphasis id="B915EAD7FFFFFFB1FEEF9F31FDB7AB3C" box="[332,581,1559,1581]" italics="true" pageId="6" pageNumber="61">Megaphyllum metsovoni</emphasis>
</taxonomicName>
male from Eptolofos, Parnassos Mts, Greece.
<emphasis id="B915EAD7FFFFFFB1FB859F31FBA3AB3C" bold="true" box="[1062,1105,1559,1581]" pageId="6" pageNumber="61">(3a)</emphasis>
Gonopods, posterior view;
<emphasis id="B915EAD7FFFFFFB1FAD39F31FA6EAB3C" bold="true" box="[1392,1436,1559,1581]" pageId="6" pageNumber="61">(3b)</emphasis>
right gonopods, mesal view;
<emphasis id="B915EAD7FFFFFFB1FE619F10FE19AB5D" bold="true" box="[450,491,1590,1612]" pageId="6" pageNumber="61">(3c)</emphasis>
left promere, meso-posterior view;
<emphasis id="B915EAD7FFFFFFB1FCFD9F10FC78AB5D" bold="true" box="[862,906,1590,1612]" pageId="6" pageNumber="61">(3d)</emphasis>
left opisthomere, lateral view; penis,
<emphasis id="B915EAD7FFFFFFB1FAB29F10FAC8AB5D" bold="true" box="[1297,1338,1590,1612]" pageId="6" pageNumber="61">(3e)</emphasis>
posterior and
<emphasis id="B915EAD7FFFFFFB1FF619F72FF1AAB7B" bold="true" box="[194,232,1620,1642]" pageId="6" pageNumber="61">(3f)</emphasis>
right lateral views. Abbreviations:
<emphasis id="B915EAD7FFFFFFB1FDF39F73FD9BAB7B" bold="true" box="[592,617,1621,1642]" italics="true" pageId="6" pageNumber="61">ap</emphasis>
: anterior process,
<emphasis id="B915EAD7FFFFFFB1FC859F73FCBBAB7B" bold="true" box="[806,841,1621,1642]" italics="true" pageId="6" pageNumber="61">asp</emphasis>
: anterior process of the solenomere,
<emphasis id="B915EAD7FFFFFFB1FB619F73FB29AB7B" bold="true" box="[1218,1243,1621,1642]" italics="true" pageId="6" pageNumber="61">pp</emphasis>
: posterior process,
<emphasis id="B915EAD7FFFFFFB1FF349F55FF48AB99" bold="true" box="[151,186,1651,1672]" italics="true" pageId="6" pageNumber="61">psp</emphasis>
: posterior process of the solenomere,
<emphasis id="B915EAD7FFFFFFB1FD9F9F55FDB9AB99" bold="true" box="[572,587,1651,1672]" italics="true" pageId="6" pageNumber="61">P</emphasis>
: promere. Scale bars: 0.2mm.
</paragraph>
</caption>
<subSubSection id="C37B654EFFFFFFB1FF349F96FDD7ABFC" pageId="6" pageNumber="61" type="materials_examined">
<paragraph id="8BDE36C5FFFFFFB1FF349F96FDD7ABFC" blockId="6.[151,1437,1712,2025]" pageId="6" pageNumber="61">
<emphasis id="B915EAD7FFFFFFB1FF349F96FE2DABD8" bold="true" box="[151,479,1712,1737]" pageId="6" pageNumber="61">
Material examined.
<collectingCountry id="F3767655FFFFFFB1FE2F9F96FE2DABD8" box="[396,479,1712,1737]" name="Greece" pageId="6" pageNumber="61">Greece</collectingCountry>
</emphasis>
:
<typeStatus id="54DA8867FFFFFFB1FE519F96FDA6ABD8" box="[498,596,1712,1737]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
male,
<collectingCountry id="F3767655FFFFFFB1FD069F97FD06ABD8" box="[677,756,1713,1737]" name="Greece" pageId="6" pageNumber="61">Greece</collectingCountry>
(Epire), coll. de Metsovon, pâturages avec
<taxonomicName id="4C614D46FFFFFFB1FB4C9F97FA66ABD8" box="[1263,1428,1713,1737]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="B915EAD7FFFFFFB1FB4C9F97FA66ABD8" box="[1263,1428,1713,1737]" italics="true" pageId="6" pageNumber="61">Rhododendron</emphasis>
</taxonomicName>
,
<date id="FFDF1005FFFFFFB1FF349FF3FEF8ABFC" box="[151,266,1749,1773]" pageId="6" pageNumber="61" value="1973-04-28">28.4.1973</date>
, B. Hauser leg. (NHMG)
</paragraph>
</subSubSection>
<subSubSection id="C37B654EFFFFFFB1FF649FDEFCA8AA00" box="[199,858,1784,1809]" pageId="6" pageNumber="61" type="distribution">
<paragraph id="8BDE36C5FFFFFFB1FF649FDEFCA8AA00" blockId="6.[151,1437,1712,2025]" box="[199,858,1784,1809]" pageId="6" pageNumber="61">
<emphasis id="B915EAD7FFFFFFB1FF649FDEFE48AA00" bold="true" box="[199,442,1784,1809]" pageId="6" pageNumber="61">
Distribution.
<collectingCountry id="F3767655FFFFFFB1FEC49FDEFE48AA00" box="[359,442,1784,1809]" name="Greece" pageId="6" pageNumber="61">Greece</collectingCountry>
</emphasis>
(Strasser 1976; Lazányi
<emphasis id="B915EAD7FFFFFFB1FD719FDFFCF8AA00" box="[722,778,1785,1809]" italics="true" pageId="6" pageNumber="61">et al.</emphasis>
2012).
</paragraph>
</subSubSection>
<subSubSection id="C37B654EFFFFFFB1FF649E3AFCEFAAF8" pageId="6" pageNumber="61" type="diagnosis">
<paragraph id="8BDE36C5FFFFFFB1FF649E3AFDEBAA48" blockId="6.[151,1437,1712,2025]" pageId="6" pageNumber="61">
<emphasis id="B915EAD7FFFFFFB1FF649E3AFEB3AA24" bold="true" box="[199,321,1820,1845]" pageId="6" pageNumber="61">Diagnosis.</emphasis>
Differs from all consubgeners by the bent, plate-like anterior process of the opisthomere (
<emphasis id="B915EAD7FFFFFFB1FA919E3BFABCAA25" bold="true" box="[1330,1358,1821,1844]" italics="true" pageId="6" pageNumber="61">ap</emphasis>
), most visible from lateral view (
<figureCitation id="135A2A40FFFFFFB1FE1A9E66FDFEAA48" box="[441,524,1856,1881]" captionStart="FIGURES 3 a f" captionStartId="6.[151,264,1559,1581]" captionTargetBox="[197,1391,396,1535]" captionTargetId="figure@6.[191,1396,394,1537]" captionTargetPageId="6" captionText="FIGURES 3 a f. Megaphyllum metsovoni male from Eptolofos, Parnassos Mts, Greece. (3 a) Gonopods, posterior view; (3 b) right gonopods, mesal view; (3 c) left promere, meso-posterior view; (3 d) left opisthomere, lateral view; penis, (3 e) posterior and (3 f) right lateral views. Abbreviations: ap: anterior process, asp: anterior process of the solenomere, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223776/files/figure.png" pageId="6" pageNumber="61">Fig. 3d</figureCitation>
).
</paragraph>
<paragraph id="8BDE36C5FFFFFFB1FF649E42FD8EAAB0" blockId="6.[151,1437,1712,2025]" pageId="6" pageNumber="61">
Opisthomeres (
<figureCitation id="135A2A40FFFFFFB1FEDB9E42FDE0AA6C" box="[376,530,1892,1917]" captionStart="FIGURES 3 a f" captionStartId="6.[151,264,1559,1581]" captionTargetBox="[197,1391,396,1535]" captionTargetId="figure@6.[191,1396,394,1537]" captionTargetPageId="6" captionText="FIGURES 3 a f. Megaphyllum metsovoni male from Eptolofos, Parnassos Mts, Greece. (3 a) Gonopods, posterior view; (3 b) right gonopods, mesal view; (3 c) left promere, meso-posterior view; (3 d) left opisthomere, lateral view; penis, (3 e) posterior and (3 f) right lateral views. Abbreviations: ap: anterior process, asp: anterior process of the solenomere, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223776/files/figure.png" pageId="6" pageNumber="61">Figs 3ab, 3d</figureCitation>
) posterior process (
<emphasis id="B915EAD7FFFFFFB1FD4D9E43FCF8AA6D" bold="true" box="[750,778,1893,1916]" italics="true" pageId="6" pageNumber="61">pp</emphasis>
) thin and reaching the height of solenomere. Solenomeres processes (
<emphasis id="B915EAD7FFFFFFB1FEB19EAFFECBAAB1" bold="true" box="[274,313,1929,1952]" italics="true" pageId="6" pageNumber="61">asp</emphasis>
and
<emphasis id="B915EAD7FFFFFFB1FED39EAFFE65AAB1" bold="true" box="[368,407,1929,1952]" italics="true" pageId="6" pageNumber="61">psp</emphasis>
) subequal in length.
</paragraph>
<paragraph id="8BDE36C5FFFFFFB1FF649E8AFCEFAAF8" blockId="6.[151,1437,1712,2025]" pageId="6" pageNumber="61">
Both sexes are reddish-brown with two bright longitudinal dorsal bands. Body length and height: males:
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,
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; female (Strasser 1976):
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,
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.
</paragraph>
</subSubSection>
<subSubSection id="C37B654EFFFEFFB0FF6499B1FE27ADC4" pageId="7" pageNumber="62" type="discussion">
<paragraph id="8BDE36C5FFFEFFB0FF6499B1FE27ADC4" blockId="7.[199,632,151,213]" pageId="7" pageNumber="62">
<emphasis id="B915EAD7FFFEFFB0FF6499B1FEC9ADA1" bold="true" box="[199,315,151,176]" pageId="7" pageNumber="62">Remarks.</emphasis>
Endemic species to
<collectingCountry id="F3767655FFFEFFB0FD8299BEFD86ADA1" box="[545,628,152,176]" name="Greece" pageId="7" pageNumber="62">Greece</collectingCountry>
. Female vulva unknown.
</paragraph>
</subSubSection>
</treatment>
</document>