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<document id="BB22BFDF8E268C7ECAAD68CEFFEB333A" ID-DOI="10.5281/zenodo.178191" ID-GBIF-Dataset="016b9301-24ee-46a2-b3ec-aa764e1e01bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="178191" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459885279670" checkinUser="plazi" docAuthor="Kerr, Peter H." docDate="2007" docId="03F8A5124D073259FF15F9D8FC7B69BF" docLanguage="en" docName="zt01554p040.pdf" docOrigin="Zootaxa 1554" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Novakia lisae Kerr, 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFC1DD6A4D013250FF82FF8BFFFD6E4D" masterDocTitle="Revision of the Holarctic genus Novakia Strobl (Diptera: Mycetophilidae)" masterLastPageNumber="40" masterPageNumber="27" pageNumber="33" updateTime="1698227745365" updateUser="plazi">
<mods:mods id="587AC539C0BD509E40988C7D5AD9E449" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="76C52F0D6DE72254AF7F1E7AB427E625">
<mods:title id="62110724343C7526C3476B37F20F5D6C">Revision of the Holarctic genus Novakia Strobl (Diptera: Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="9897F49E30F8494BDE4F2F8DA2362BF6" type="personal">
<mods:role id="F94797A9889460895F54506FBAEC7416">
<mods:roleTerm id="426626D03E53967B2678F9B71D2FB394">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4314CA690DA7408EDBB627A53F17FE78">Kerr, Peter H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2006986A4E6E9D63C2A810946C4BF950">text</mods:typeOfResource>
<mods:relatedItem id="FD0F4B436025ED5EACEC608B712FA903" type="host">
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<mods:title id="1563846E4D10E8D7A56B105E7D94FA22">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9714B7AC86E3C72AACE47BA8F8E04BA0">
<mods:date id="699904DEEA4021CDE62953BCAB6CF40E">2007</mods:date>
<mods:detail id="C1C7A7BD35F275B2668D0E50575F00A8" type="volume">
<mods:number id="09347FE208CD3C253C331A04C2DB25D9">1554</mods:number>
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<mods:classification id="B4FB1F7B4312C8E58E00396345AA6569">journal article</mods:classification>
<mods:identifier id="295C4B108CBE17F44105382257FA419C" type="DOI">10.5281/zenodo.178191</mods:identifier>
<mods:identifier id="6B2A730B3EA3AE0205D76E7C8AFD1268" type="GBIF-Dataset">016b9301-24ee-46a2-b3ec-aa764e1e01bb</mods:identifier>
<mods:identifier id="4136B6E3467CB2F2697F7CAD03120B76" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="30C85F4CA5D51C6691E367DCAEA8F1B7" type="Zenodo-Dep">178191</mods:identifier>
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<treatment id="03F8A5124D073259FF15F9D8FC7B69BF" ID-DOI="http://doi.org/10.5281/zenodo.6249881" ID-GBIF-Taxon="119371424" ID-Zenodo-Dep="6249881" LSID="urn:lsid:plazi:treatment:03F8A5124D073259FF15F9D8FC7B69BF" httpUri="http://treatment.plazi.org/id/03F8A5124D073259FF15F9D8FC7B69BF" lastPageId="9" lastPageNumber="36" pageId="6" pageNumber="33">
<subSubSection id="C34B478F4D073256FF15F9D8FEAE68DD" pageId="6" pageNumber="33" type="nomenclature">
<paragraph id="8BEE14044D073256FF15F9D8FE626821" blockId="6.[151,415,1618,1680]" box="[151,415,1618,1645]" pageId="6" pageNumber="33">
<heading id="D0A6A3684D073256FF15F9D8FE626821" bold="true" box="[151,415,1618,1645]" fontSize="11" level="1" pageId="6" pageNumber="33" reason="1">
<emphasis id="B925C8164D073256FF15F9D8FE626821" bold="true" box="[151,415,1618,1645]" pageId="6" pageNumber="33">
<taxonomicName id="4C516F874D073256FF15F9D8FEC66820" ID-CoL="47ZXL" box="[151,315,1619,1645]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="lisae" status="sp. nov.">
<emphasis id="B925C8164D073256FF15F9D8FEC66820" bold="true" box="[151,315,1619,1645]" italics="true" pageId="6" pageNumber="33">Novakia lisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216756D4D073256FEC0F9D9FE626821" box="[322,415,1618,1644]" pageId="6" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE14044D073256FF15F9FDFEAE68DD" blockId="6.[151,415,1618,1680]" box="[151,339,1654,1680]" pageId="6" pageNumber="33">
(
<figureCitation id="136A08814D073256FF1DF9FDFEB768DD" box="[159,330,1654,1680]" captionStart-0="FIGURES 10 – 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStart-3="FIGURE 18" captionStartId-0="7.[151,269,1923,1947]" captionStartId-1="9.[151,255,904,928]" captionStartId-2="9.[151,255,1348,1372]" captionStartId-3="10.[151,255,915,939]" captionTargetBox-0="[214,1379,155,1905]" captionTargetBox-2="[434,1153,975,1333]" captionTargetBox-3="[240,1346,194,891]" captionTargetId-0="figure@7.[214,1379,155,1906]" captionTargetId-2="figure@9.[434,1154,975,1333]" captionTargetId-3="figure@10.[240,1346,194,891]" captionTargetPageId-0="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURES 10 – 15. Novakia lisae, male. 10, head, anteroventral view; 11, head, oblique lateral view; 12, first tarsus, ventral view; 13, genitalia, lateral view; 14, genitalia, dorsal view; 15, genitalia, terminal view. In figures 11 – 12 and 14 – 16, scale bar = 0.1 mm; in figure 13, scale bar = 0.01 mm." captionText-1="FIGURE 16. Novakia lisae, female, wing, dorsal view." captionText-2="FIGURE 17. Novakia lisae, male genitalia. A, hypandrium, dorsal view; B, epandrium, dorsal view. Scale bar = 0.1 mm." captionText-3="FIGURE 18. Novakia lisae, female genitalia. A, terminalia, lateral view; B, sternite 8, ventral view. Scale bar = 0.1 mm. Abbreviations: tg = tergite; s = seta." httpUri-0="https://zenodo.org/record/178195/files/figure.png" httpUri-2="https://zenodo.org/record/178196/files/figure.png" httpUri-3="https://zenodo.org/record/178197/files/figure.png" pageId="6" pageNumber="33">Figures 10–18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34B478F4D073258FF15F94DFD406D99" lastPageId="8" lastPageNumber="35" pageId="6" pageNumber="33" type="materials_examined">
<paragraph id="8BEE14044D073256FF15F94DFEC668AD" blockId="6.[151,1436,1734,1920]" box="[151,315,1734,1760]" pageId="6" pageNumber="33">
<typeStatus id="54EAAAA64D073256FF15F94DFF2C68AD" box="[151,209,1734,1760]" pageId="6" pageNumber="33">Type</typeStatus>
Material
</paragraph>
<paragraph id="8BEE14044D073256FF44F965FAD369CD" blockId="6.[151,1436,1734,1920]" pageId="6" pageNumber="33">
<typeStatus id="54EAAAA64D073256FF44F965FEC56945" box="[198,312,1774,1800]" pageId="6" pageNumber="33" type="holotype">Holotype</typeStatus>
: ♂,
<collectingCountry id="F34654944D073256FEE9F965FE576945" box="[363,426,1774,1800]" name="United States of America" pageId="6" pageNumber="33">USA</collectingCountry>
: CA: San Bernardino Co.,
<quantity id="4CA9B9E14D073256FD7AF965FCCA6945" box="[760,823,1774,1800]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="33" unit="km" value="9.0">9 km</quantity>
SE Wrightwood. Lone Pine Canyon,
<geoCoordinate id="EE6572C34D073256FA88F965FA6A6945" box="[1290,1431,1774,1800]" direction="north" orientation="latitude" pageId="6" pageNumber="33" precision="9" value="34.302834">34°18.17’N</geoCoordinate>
,
<geoCoordinate id="EE6572C34D073256FF15F89DFEC8697D" box="[151,309,1814,1840]" direction="west" orientation="longitude" pageId="6" pageNumber="33" precision="9" value="-117.53017">117°31.81’W</geoCoordinate>
, elev. 4127’,
<date id="FFEF32C44D073256FE5CF89DFD82697D" box="[478,639,1814,1840]" pageId="6" pageNumber="33" value="2005-05-21" valueMax="2005-05-28" valueMin="2005-05-21">21–28.v.2005</date>
, S.L. Winterton &amp; A.R. Cline, Malaise
<collectionCode id="ED408CC14D073256FBEFF89DFB41697D" box="[1133,1212,1814,1840]" pageId="6" pageNumber="33">CDFA</collectionCode>
2005-010 [
<collectionCode id="ED408CC14D073256FABEF89DFA6F697D" LSID="urn:lsid:biocol.org:col:33156" box="[1340,1426,1814,1840]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="6" pageNumber="33">CSCA</collectionCode>
]. Specimen on point mount, missing all legs on left side, otherwise in excellent condition.
<typeStatus id="54EAAAA64D073256FF44F8EDFECC69CD" box="[198,305,1894,1920]" pageId="6" pageNumber="33" type="paratype">Paratype</typeStatus>
: 1 Ψ, same data as
<typeStatus id="54EAAAA64D073256FD8CF8EDFD8969CD" box="[526,628,1894,1920]" pageId="6" pageNumber="33" type="holotype">holotype</typeStatus>
[
<collectionCode id="ED408CC14D073256FD01F8EDFD2769CD" box="[643,730,1894,1920]" pageId="6" pageNumber="33">PHKC</collectionCode>
]. Specimen on point mount, in excellent condition.
</paragraph>
<caption id="DF2E448C4D063257FF15F808FD2669AD" httpUri="https://zenodo.org/record/178195/files/figure.png" pageId="7" pageNumber="34" targetBox="[214,1379,155,1905]" targetPageId="7">
<paragraph id="8BEE14044D063257FF15F808FD2669AD" blockId="7.[151,1436,1923,2016]" pageId="7" pageNumber="34">
<emphasis id="B925C8164D063257FF15F808FEA069D6" bold="true" box="[151,349,1923,1947]" pageId="7" pageNumber="34">FIGURES 10–15.</emphasis>
<taxonomicName id="4C516F874D063257FEE6F80FFE0869D6" box="[356,501,1924,1947]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B925C8164D063257FEE6F80FFE0869D6" box="[356,501,1924,1947]" italics="true" pageId="7" pageNumber="34">Novakia lisae</emphasis>
</taxonomicName>
, male. 10, head, anteroventral view; 11, head, oblique lateral view; 12, first tarsus, ventral view; 13, genitalia, lateral view; 14, genitalia, dorsal view; 15, genitalia, terminal view. In figures 11–12 and 14–16, scale bar = 0.1 mm; in figure 13, scale bar = 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BEE14044D093258FF44FF13FD566F67" blockId="8.[151,1436,152,298]" pageId="8" pageNumber="35">
Additional
<typeStatus id="54EAAAA64D093258FECDFF13FE426EFF" box="[335,447,152,178]" pageId="8" pageNumber="35" type="paratype">paratypes</typeStatus>
in alcohol:
<specimenCount id="9D57DF8D4D093258FDD7FF13FD7C6EE6" box="[597,641,152,178]" pageId="8" pageNumber="35" type="generic">1 ♂</specimenCount>
, 1 Ψ, same data as
<typeStatus id="54EAAAA64D093258FCF5FF13FC206EFF" box="[887,989,152,178]" pageId="8" pageNumber="35" type="holotype">holotype</typeStatus>
[CSCA-FTC, database det. numbers
<date id="FFEF32C44D093258FF15FF4BFF366E97" box="[151,203,192,218]" pageId="8" pageNumber="35" value="2006-10">06X</date>
608,
<date id="FFEF32C44D093258FE84FF4BFEC06E97" box="[262,317,192,218]" pageId="8" pageNumber="35" value="2006-10">06X</date>
609]; 1 Ψ,
<collectingCountry id="F34654944D093258FE3EFF4BFE066E97" box="[444,507,192,218]" name="United States of America" pageId="8" pageNumber="35">USA</collectingCountry>
: CA: Los Angeles Co.,
<quantity id="4CA9B9E14D093258FC9CFF4BFCA16E97" box="[798,860,192,218]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="35" unit="km" value="1.0">1 km</quantity>
NW Wrightwood, Big Pines Hwy.,
<geoCoordinate id="EE6572C34D093258FA90FF4BFA616E97" box="[1298,1436,192,218]" direction="north" orientation="latitude" pageId="8" pageNumber="35" precision="9" value="34.3755">34°22.53’N</geoCoordinate>
<geoCoordinate id="EE6572C34D093258FF15FF63FEC56F4F" box="[151,312,232,258]" direction="west" orientation="longitude" pageId="8" pageNumber="35" precision="9" value="-117.66967">117°40.18’W</geoCoordinate>
; elev. 6476’,
<date id="FFEF32C44D093258FE60FF63FD796F4F" box="[482,644,232,258]" pageId="8" pageNumber="35" value="2005-05-21" valueMax="2005-05-28" valueMin="2005-05-21">21–28.v.2005</date>
, S.L. Winterton &amp; A.R. Cline, Malaise in wash
<collectionCode id="ED408CC14D093258FB63FF63FACC6F4F" box="[1249,1329,232,258]" pageId="8" pageNumber="35">CDFA</collectionCode>
2005-011 [CSCA-FTC, database det. number
<date id="FFEF32C44D093258FDB9FE9BFD8D6F67" box="[571,624,272,298]" pageId="8" pageNumber="35" value="2006-10">06X</date>
706].
</paragraph>
<paragraph id="8BEE14044D093258FF15FEEBFEF16F37" blockId="8.[151,1436,352,498]" box="[151,268,352,378]" pageId="8" pageNumber="35">Diagnosis</paragraph>
<paragraph id="8BEE14044D093258FF44FE03FAB46FBF" blockId="8.[151,1436,352,498]" pageId="8" pageNumber="35">
This species is the most distinctive of the genus. The most prominent diagnostic features that separate it from all other
<taxonomicName id="4C516F874D093258FEC2FE3BFE5F6F84" box="[320,418,432,457]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B925C8164D093258FEC2FE3BFE5F6F84" box="[320,418,432,457]" italics="true" pageId="8" pageNumber="35">Novakia</emphasis>
</taxonomicName>
species include pale yellowish or white colored mid and hind legs, antennal bases that touch medially, and lateral ocellus positioned approximately its own diameter or less from eye margin.
</paragraph>
<paragraph id="8BEE14044D093258FF15FDA3FEDD6C0F" blockId="8.[151,1437,552,2020]" box="[151,288,552,578]" pageId="8" pageNumber="35">Description</paragraph>
<paragraph id="8BEE14044D093258FF44FDDBFA896C27" blockId="8.[151,1437,552,2020]" box="[198,1396,592,618]" pageId="8" pageNumber="35">
Body length:
<quantity id="4CA9B9E14D093258FEE4FDDBFE0A6C27" box="[358,503,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="8" pageNumber="35" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
. Wing length:
<quantity id="4CA9B9E14D093258FD23FDDBFCCE6C27" box="[673,819,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="8" pageNumber="35" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
. Males and females approximately the same size.
</paragraph>
<paragraph id="8BEE14044D093258FF44FDF3FD406D99" blockId="8.[151,1437,552,2020]" pageId="8" pageNumber="35">
<emphasis id="B925C8164D093258FF44FDF3FEBA6CDC" box="[198,327,632,657]" italics="true" pageId="8" pageNumber="35">Coloration</emphasis>
. Head dark brown to black. Eye color brown when preserved fresh in alcohol, eyes gray when dried. Antenna brown to dark brown. Clypeus dark brown to black, basal palpomeres pale brown, segments becoming increasingly white distally; palpomere 5 entirely white. Scutum and scutellum brown to black. Thoracic sclerites brown. Coxae whitish or pale yellow, femora and tibiae whitish or pale yellow to pale yellowish brown; tibial spurs yellowish brown to brown; tarsi yellowish brown. Wing with brownish tint over most of membrane, narrow hyaline area posterior of wing vein R5, cubital veins slightly darker than medial veins (wing veins not noticeably darker in female), haltere stem white and knob pale to dark brown. Abdominal segments brown, with faint black band at posterior margin. Terminalia brown to dark brown in male; in female, cercus pale yellowish to pale yellowish brown.
</paragraph>
</subSubSection>
<subSubSection id="C34B478F4D093259FF44FC68FB116897" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="description">
<paragraph id="8BEE14044D093258FF44FC68FCCE6831" blockId="8.[151,1437,552,2020]" pageId="8" pageNumber="35">
<emphasis id="B925C8164D093258FF44FC68FEFB6DB1" box="[198,262,995,1020]" italics="true" pageId="8" pageNumber="35">Head</emphasis>
(
<figureCitation id="136A08814D093258FE95FC69FE596DB1" box="[279,420,994,1020]" captionStart="FIGURES 10 – 15" captionStartId="7.[151,269,1923,1947]" captionTargetBox="[214,1379,155,1905]" captionTargetId="figure@7.[214,1379,155,1906]" captionTargetPageId="7" captionText="FIGURES 10 – 15. Novakia lisae, male. 10, head, anteroventral view; 11, head, oblique lateral view; 12, first tarsus, ventral view; 13, genitalia, lateral view; 14, genitalia, dorsal view; 15, genitalia, terminal view. In figures 11 – 12 and 14 – 16, scale bar = 0.1 mm; in figure 13, scale bar = 0.01 mm." httpUri="https://zenodo.org/record/178195/files/figure.png" pageId="8" pageNumber="35">Figs. 10–11</figureCitation>
). Median ocellus less than half diameter of lateral ocellus; lateral ocellus less than its own diameter from eye margin (in female, lateral ocellus closer to eye margin). Postocular region and top of head with evenly distributed, short brown setae; frons bare. Frontal tubercle above antennal bases; transverse frontoclypeal membrane oblique, incomplete. Face and clypeus not separated by frontoclypeal membrane medially; face and clypeus swollen and projecting forward, beyond rest of head; area between base of antennae and clypeolabral articulation (= face + clypeus) longer than wide. Palp about as long as head or slightly longer; segments l–2 short, lacking setae; palpomere 3 longer, with short dark setae scattered throughout, also with small and inconspicuous shallow pit on basal half of inner surface containing sensilla cochleariformis (
<quantity id="4CA9B9E14D093258FF22FAA9FF166B71" box="[160,235,1314,1340]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="8" pageNumber="35" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5–8 in</quantity>
male); palpomere 4 longer than palpomere 3, with short pale brown setae; palpomere 5 cylindrical and elongate, longer than combined length of other palpomeres, more than twice length of palpomere 4, with short pale colored setae. Antennal length in male approximately 3.0 times the length of head; in female, antennal length approximately 2.2 times the length of head; antennal bases separated; scape with dorsal extension covering base of pedicel, with ring of dark setae, setae shorter than length of scape, slightly longer ventrally; pedicel in lateral view as wide as scape, with ring of dark setae distally, shorter than length of pedicel; all flagellomeres densely setulose, in male, flagellomeres 1–13 longer than wide and terminal flagellomere approximately twice as long as wide; in female, flagellomeres 1–13 approximately as long as wide and terminal flagellomere slightly less than twice as long as wide.
</paragraph>
<paragraph id="8BEE14044D093259FF44F900FDEE6F98" blockId="8.[151,1437,552,2020]" lastBlockId="9.[151,1436,152,469]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
<emphasis id="B925C8164D093258FF44F900FEE668E9" box="[198,283,1675,1700]" italics="true" pageId="8" pageNumber="35">Thorax</emphasis>
. Scutellum bearing two pairs of long black setae on posterior margin, several shorter setae also present on posterior margin. Setae of legs black, short. Fore coxa setose over entire anterior surface and with a few setae on the posterior surface at apex only, mid coxa setose anteriorly, hind coxa with single row of 3–5 setae at distal 2/3 of dorsal surface and with setae at apical margin. Approximate femur length to tibia length ratio 1.2 (foreleg), 1.0 (midleg), and 0.9 (hindleg). Trichia of tibia and tarsus brown, prominent; all tibial setae shorter than width of tibia; fore tibia with ovate anteroapical depressed area bearing several rows of small yellowish setae; mid tibia with 6 anterior setae, 4–5 dorsal setae, and 3–4 posterior setae on apical third; hind tibia with a row of 5–9 anterior setae, a row of dorsal setae, and a 0–3 posterior setae on apical third, and a cluster of approximately 25 short setae posteroapically. In male, tarsal claw finely dissected terminally, modified into a 6-tined comb (
<figureCitation id="136A08814D083259FE40FF13FDE46EFF" box="[450,537,152,178]" captionStart="FIGURES 10 – 15" captionStartId="7.[151,269,1923,1947]" captionTargetBox="[214,1379,155,1905]" captionTargetId="figure@7.[214,1379,155,1906]" captionTargetPageId="7" captionText="FIGURES 10 – 15. Novakia lisae, male. 10, head, anteroventral view; 11, head, oblique lateral view; 12, first tarsus, ventral view; 13, genitalia, lateral view; 14, genitalia, dorsal view; 15, genitalia, terminal view. In figures 11 – 12 and 14 – 16, scale bar = 0.1 mm; in figure 13, scale bar = 0.01 mm." httpUri="https://zenodo.org/record/178195/files/figure.png" pageId="9" pageNumber="36">Fig. 12</figureCitation>
); in female, tarsal claw simple; empodium pulvilliform. Wing (
<figureCitation id="136A08814D083259FA80FF13FAAA6EFF" box="[1282,1367,152,178]" captionStart="FIGURE 16" captionStartId="9.[151,255,904,928]" captionText="FIGURE 16. Novakia lisae, female, wing, dorsal view." pageId="9" pageNumber="36">Fig. 16</figureCitation>
) with small, irregularly arranged trichia over cell membrane surface; costal vein extends approximately 0.4 of distance between R5 and M1; subcostal vein short, thick, bare, and ending free; radial veins with setulae on upper surface; vein R1 longer than half length of r-m; crossvein r-m close to R along length; Rs absent but position marked where radial veins touch before R1 turns to costal vein; r-m and R5 join seamlessly (as R and R1); stem of medial fork readily apparent for its entire length, shorter than length of fork; M1 and M2 with reduced, inconspicuous setulae on apical half of upper surface only; basal transversal vein readily apparent; cubital veins bare; anal vein short, bare.
</paragraph>
<caption id="DF2E448C4D083259FF15FC03FD1B6DED" box="[151,742,904,928]" pageId="9" pageNumber="36">
<paragraph id="8BEE14044D083259FF15FC03FD1B6DED" blockId="9.[151,742,904,928]" box="[151,742,904,928]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF15FC03FEDB6DED" bold="true" box="[151,294,904,928]" pageId="9" pageNumber="36">FIGURE 16.</emphasis>
<taxonomicName id="4C516F874D083259FEAFFC02FE436DED" box="[301,446,905,928]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B925C8164D083259FEAFFC02FE436DED" box="[301,446,905,928]" italics="true" pageId="9" pageNumber="36">Novakia lisae</emphasis>
</taxonomicName>
, female, wing, dorsal view.
</paragraph>
</caption>
<caption id="DF2E448C4D083259FF15FACFFA616B11" box="[151,1436,1348,1372]" httpUri="https://zenodo.org/record/178196/files/figure.png" pageId="9" pageNumber="36" targetBox="[434,1153,975,1333]" targetPageId="9">
<paragraph id="8BEE14044D083259FF15FACFFA616B11" blockId="9.[151,1436,1348,1372]" box="[151,1436,1348,1372]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF15FACFFEDB6B11" bold="true" box="[151,294,1348,1372]" pageId="9" pageNumber="36">FIGURE 17.</emphasis>
<taxonomicName id="4C516F874D083259FEAEFACEFE406B11" box="[300,445,1349,1372]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B925C8164D083259FEAEFACEFE406B11" box="[300,445,1349,1372]" italics="true" pageId="9" pageNumber="36">Novakia lisae</emphasis>
</taxonomicName>
, male genitalia. A, hypandrium, dorsal view; B, epandrium, dorsal view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEE14044D083259FF44FA23FF33682F" blockId="9.[151,1437,1448,1754]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF44FA23FE776B8C" box="[198,394,1448,1473]" italics="true" pageId="9" pageNumber="36">Male Terminalia</emphasis>
(
<figureCitation id="136A08814D083259FE18FA23FDD56B8F" box="[410,552,1448,1474]" captionStart="FIGURES 10 – 15" captionStartId="7.[151,269,1923,1947]" captionTargetBox="[214,1379,155,1905]" captionTargetId="figure@7.[214,1379,155,1906]" captionTargetPageId="7" captionText="FIGURES 10 – 15. Novakia lisae, male. 10, head, anteroventral view; 11, head, oblique lateral view; 12, first tarsus, ventral view; 13, genitalia, lateral view; 14, genitalia, dorsal view; 15, genitalia, terminal view. In figures 11 – 12 and 14 – 16, scale bar = 0.1 mm; in figure 13, scale bar = 0.01 mm." httpUri="https://zenodo.org/record/178195/files/figure.png" pageId="9" pageNumber="36">Figs. 13–15</figureCitation>
,
<figureCitation id="136A08814D083259FDB4FA23FDAB6B8F" box="[566,598,1448,1474]" captionStart="FIGURE 17" captionStartId="9.[151,255,1348,1372]" captionTargetBox="[434,1153,975,1333]" captionTargetId="figure@9.[434,1154,975,1333]" captionTargetPageId="9" captionText="FIGURE 17. Novakia lisae, male genitalia. A, hypandrium, dorsal view; B, epandrium, dorsal view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/178196/files/figure.png" pageId="9" pageNumber="36">17</figureCitation>
). Epandrium (= tergite 9) posterior margin not emarginate. Gonocoxal lobes (
<bibRefCitation id="EFC069F54D083259FF6AFA5BFE9B6BA7" author="Soli" box="[232,358,1488,1514]" pageId="9" pageNumber="40" refString="Soli, G. E. E. (1997) The adult morphology of Mycetophilidae (s. str.), with a tentative phylogeny of the family (Diptera, Sciaroidea). Entomologica Scandinavica Supplement, 50, 5 - 55." type="journal article" year="1997">Søli, 1997</bibRefCitation>
) separated medially, exposing aedeagal complex in ventral view; gonostylus armed with thick setae that extend past midline, with tapered apical lobe; internal lobe of gonostylus absent; aedeagus dissected medially, with apical fringe; parameres long, extending prominently beyond confines of genitalic capsule.
</paragraph>
<paragraph id="8BEE14044D083259FF44F9FBFB116897" blockId="9.[151,1437,1448,1754]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF44F9FBFE5568C4" box="[198,424,1648,1673]" italics="true" pageId="9" pageNumber="36">Female Terminalia</emphasis>
(
<figureCitation id="136A08814D083259FE38F9FBFDEE68C7" box="[442,531,1648,1674]" captionStart="FIGURE 18" captionStartId="10.[151,255,915,939]" captionTargetBox="[240,1346,194,891]" captionTargetId="figure@10.[240,1346,194,891]" captionTargetPageId="10" captionText="FIGURE 18. Novakia lisae, female genitalia. A, terminalia, lateral view; B, sternite 8, ventral view. Scale bar = 0.1 mm. Abbreviations: tg = tergite; s = seta." httpUri="https://zenodo.org/record/178197/files/figure.png" pageId="9" pageNumber="36">Fig. 18</figureCitation>
). First cercus segment broad, second cercus segment ovoid, often flattened with transversely flattened apical beak-like process; spermathecae connected directly to genital chamber via short spermathecal ducts, spermathecal duct length slightly longer than two-segmented cercus.
</paragraph>
</subSubSection>
<subSubSection id="C34B478F4D083259FF15F89BFCD26937" pageId="9" pageNumber="36" type="discussion">
<paragraph id="8BEE14044D083259FF15F89BFEEE6964" blockId="9.[151,1436,1808,1914]" box="[151,275,1808,1833]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF15F89BFEEE6964" box="[151,275,1808,1833]" italics="true" pageId="9" pageNumber="36">Comments</emphasis>
</paragraph>
<paragraph id="8BEE14044D083259FF44F8B3FCD26937" blockId="9.[151,1436,1808,1914]" pageId="9" pageNumber="36">
This species is only known from the San Gabriel Mountains (near Wrightwood, CA; Los Angeles and San Bernadino counties), where it is sympatric with
<taxonomicName id="4C516F874D083259FD48F8EBFCD56934" box="[714,808,1888,1913]" class="Insecta" family="Mycetophilidae" genus="Novakia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="miloi">
<emphasis id="B925C8164D083259FD48F8EBFCD56934" box="[714,808,1888,1913]" italics="true" pageId="9" pageNumber="36">N. miloi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B478F4D083259FF15F83BFC7B69BF" box="[151,902,1968,2034]" pageId="9" pageNumber="36" type="etymology">
<paragraph id="8BEE14044D083259FF15F83BFEE96984" blockId="9.[151,902,1968,2034]" box="[151,276,1968,1993]" pageId="9" pageNumber="36">
<emphasis id="B925C8164D083259FF15F83BFEE96984" box="[151,276,1968,1993]" italics="true" pageId="9" pageNumber="36">Etymology</emphasis>
</paragraph>
<paragraph id="8BEE14044D083259FF44F853FC7B69BF" blockId="9.[151,902,1968,2034]" box="[198,902,2008,2034]" pageId="9" pageNumber="36">
Named in honor of Lisa Sofia Kerr, born
<date id="FFEF32C44D083259FD29F853FC7F69BF" box="[683,898,2008,2034]" pageId="9" pageNumber="36" value="2006-11-03">3 November 2006</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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