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<document id="56A359DA66884BFDEA8FDCF6C64D8F20" ID-DOI="10.11646/zootaxa.5023.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5226026" ID-ZooBank="CABE11FD-4FAE-4716-80F8-390376AF1B73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629427582629" checkinUser="plazi" docAuthor="Johansson, Niklas" docDate="2021" docId="03EE4833FFD10828FF6FFF1E83863BE2" docLanguage="en" docName="zootaxa.5023.3.1.pdf" docOrigin="Zootaxa 5023 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophion albanicus Johansson 2021, sp. nov." docType="treatment" docUuid="3633E7BD-BFFF-4D9A-A189-26F2614CDE44" docUuidSource="ZooBank" docVersion="3" lastPageNumber="313" masterDocId="FFD7304BFFDD0824FFF8FF8981453F22" masterDocTitle="Contribution to the subfamily Ophioninae (Hymenoptera: Ichneumonidae) of Southern Europe with the description of one new genus and 15 new species" masterLastPageNumber="334" masterPageNumber="301" pageNumber="313" updateTime="1699072774709" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E013C92F3618CDEF92078E683FEA1058">Contribution to the subfamily Ophioninae (Hymenoptera: Ichneumonidae) of Southern Europe with the description of one new genus and 15 new species</mods:title>
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<mods:namePart id="0A08C25D42E3EDF84ADF2FF031771A56">Johansson, Niklas</mods:namePart>
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<mods:title id="97A732D454D7214700F5F77871D236C4">Zootaxa</mods:title>
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<mods:part id="C6943A4FCEBE0F447793677A8F8481A9">
<mods:date id="EC7811B334DCF3FC1E1BA052434CCCCA">2021</mods:date>
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<mods:number id="D3830E1DBB00522F372A7E178167BB44">2021-08-19</mods:number>
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<mods:number id="60AE6C8CBCE97D20A386E5EC76695211">5023</mods:number>
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<mods:number id="BBDA1BA2B7E367197D6C38150B078412">3</mods:number>
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<mods:classification id="4024266623176D1BBCD79ED5C522F5BA">journal article</mods:classification>
<mods:identifier id="25B0E1F8F1675A0CCBF06734D5598E4C" type="DOI">10.11646/zootaxa.5023.3.1</mods:identifier>
<mods:identifier id="CDF1F1281C341305E463C89455453F77" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="89BF27DCEF89C76E6CC9EA7AF48CC00F" type="Zenodo-Dep">5226026</mods:identifier>
<mods:identifier id="0B9835D0DFE07385E0E6F3B57804BFCB" type="ZooBank">CABE11FD-4FAE-4716-80F8-390376AF1B73</mods:identifier>
</mods:mods>
<treatment id="03EE4833FFD10828FF6FFF1E83863BE2" LSID="urn:lsid:zoobank.org:act:3633E7BD-BFFF-4D9A-A189-26F2614CDE44" httpUri="http://treatment.plazi.org/id/03EE4833FFD10828FF6FFF1E83863BE2" lastPageNumber="313" pageId="12" pageNumber="313">
<subSubSection id="C35DAAAEFFD10828FF6FFF1E80913F93" box="[151,468,151,178]" pageId="12" pageNumber="313" type="nomenclature">
<paragraph id="8BF8F925FFD10828FF6FFF1E80913F93" blockId="12.[151,468,151,214]" box="[151,468,151,178]" pageId="12" pageNumber="313">
<heading id="D0B04E49FFD10828FF6FFF1E80913F93" bold="true" box="[151,468,151,178]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
<emphasis id="B9332537FFD10828FF6FFF1E80913F93" bold="true" box="[151,468,151,178]" pageId="12" pageNumber="313">
<taxonomicName id="4C4782A6FFD10828FF6FFF1E802A3F90" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[151,367,151,178]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">
<emphasis id="B9332537FFD10828FF6FFF1E802A3F90" bold="true" box="[151,367,151,178]" italics="true" pageId="12" pageNumber="313">Ophion albanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD10828FE8FFF1E80913F93" box="[375,468,151,177]" pageId="12" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD10828FF6FFF3282C53E35" pageId="12" pageNumber="313" type="description">
<paragraph id="8BF8F925FFD10828FF6FFF3282C53E35" blockId="12.[151,468,151,214]" lastBlockId="12.[151,896,255,279]" pageId="12" pageNumber="313">
<figureCitation id="137CE5A0FFD10828FF6FFF3280733FF4" box="[151,310,187,214]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Figs 8 A–C, E</figureCitation>
<uri id="FFD6F527FFD10828FF6FFF7682C53E35" box="[151,896,255,279]" pageId="12" pageNumber="313">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD10828FF6FFECF83D43EA6" pageId="12" pageNumber="313" type="materials_examined">
<paragraph id="8BF8F925FFD10828FF6FFECF83D43EA6" blockId="12.[151,1437,325,1216]" pageId="12" pageNumber="313">
<materialsCitation id="3B2FF378FFD10828FF6FFECF83D43EA6" collectingDate="2019-06-05" collectionCode="NMS" collectorName="Gropa Mt. Above Shen Meri &amp; S. Beshkov &amp; A. Nahirnic" country="Albania" county="Mt. Above Shen Meri" elevation="1405" location="Mali" municipality="Gropa Mt." pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B9332537FFD10828FF6FFECF80413E42" bold="true" box="[151,260,326,352]" pageId="12" pageNumber="313">
<typeStatus id="54FC4787FFD10828FF6FFECF80413E42" box="[151,260,326,352]" pageId="12" pageNumber="313" type="holotype">Holotype</typeStatus>
</emphasis>
:
<emphasis id="B9332537FFD10828FEEBFECC80D13E7D" bold="true" box="[275,404,325,351]" pageId="12" pageNumber="313">
<collectingCountry id="F350B9B5FFD10828FEEBFECC80D13E7D" box="[275,404,325,351]" name="Albania" pageId="12" pageNumber="313">ALBANIA</collectingCountry>
</emphasis>
:
<specimenCount id="9D4132ACFFD10828FE5CFECF80FC3E42" box="[420,441,326,352]" pageId="12" pageNumber="313" type="female">♀</specimenCount>
,
<location id="8E98AFFEFFD10828FE31FECF80BB3E42" LSID="urn:lsid:plazi:treatment:03EE4833FFD10828FF6FFF1E83863BE2:8E98AFFEFFD10828FE31FECF80BB3E42" box="[457,510,326,352]" country="Albania" county="Mt. Above Shen Meri" municipality="Gropa Mt." name="Mali" pageId="12" pageNumber="313">Mali</location>
me
<collectorName id="26B29CF3FFD10828FDC9FECF823F3E42" box="[561,890,326,352]" pageId="12" pageNumber="313">
<collectingMunicipality id="6B9C635FFFD10828FDC9FECF83E23E42" box="[561,679,326,352]" pageId="12" pageNumber="313">
Gropa
<collectingCounty id="629981A9FFD10828FD87FECF83E23E42" box="[639,679,326,352]" pageId="12" pageNumber="313">Mt.</collectingCounty>
</collectingMunicipality>
Above Shen Meri
</collectorName>
, 41˚3524’N, 020˚0465’E,
<quantity id="4CBF54C0FFD10828FB5FFECF84453E42" box="[1191,1280,326,353]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="12" pageNumber="313" unit="m" value="1405.0">
<elevation id="006A1E16FFD10828FB5FFECF84453E42" box="[1191,1280,326,353]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="12" pageNumber="313" unit="m" value="1405.0">1405 m</elevation>
</quantity>
. a. s.,
<date id="FFF9DFE5FFD10828FAB2FECF81973EA6" pageId="12" pageNumber="313" value="2019-06-05">
<collectingDate id="EFBD260DFFD10828FAB2FECF81973EA6" pageId="12" pageNumber="313" value="2019-06-05">05 Jun. 2019</collectingDate>
</date>
,
<collectorName id="26B29CF3FFD10828FF25FEE380183EA6" box="[221,349,362,388]" pageId="12" pageNumber="313">S. Beshkov</collectorName>
&amp;
<collectorName id="26B29CF3FFD10828FE87FEE383463EA6" box="[383,515,362,388]" pageId="12" pageNumber="313">A. Nahirnic</collectorName>
leg. (
<collectionCode id="ED5661E0FFD10828FDBBFEE383C03EA6" box="[579,645,362,388]" country="United Kingdom" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="12" pageNumber="313" type="Museum">NMS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD10828FF3FFE0785B23E8A" box="[199,1271,398,424]" pageId="12" pageNumber="313" type="etymology">
<paragraph id="8BF8F925FFD10828FF3FFE0785B23E8A" blockId="12.[151,1437,325,1216]" box="[199,1271,398,424]" pageId="12" pageNumber="313">
<emphasis id="B9332537FFD10828FF3FFE07800D3E8A" bold="true" box="[199,328,398,424]" pageId="12" pageNumber="313">Etymology</emphasis>
: The name
<taxonomicName id="4C4782A6FFD10828FE35FE07837F3E8A" authorityName="Johansson" authorityYear="2021" box="[461,570,398,424]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="albanicus">
<emphasis id="B9332537FFD10828FE35FE07837F3E8A" box="[461,570,398,424]" italics="true" pageId="12" pageNumber="313">albanicus</emphasis>
</taxonomicName>
is derived from
<collectingCountry id="F350B9B5FFD10828FD0AFE07820B3E8A" box="[754,846,398,424]" name="Albania" pageId="12" pageNumber="313">Albania</collectingCountry>
, where the
<typeStatus id="54FC4787FFD10828FC35FE0782B83E8A" box="[973,1021,398,424]" pageId="12" pageNumber="313">type</typeStatus>
female was collected.
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD10828FF3FFE3885283D1A" pageId="12" pageNumber="313" type="diagnosis">
<paragraph id="8BF8F925FFD10828FF3FFE3885283D1A" blockId="12.[151,1437,325,1216]" pageId="12" pageNumber="313">
<emphasis id="B9332537FFD10828FF3FFE38807F3EE9" bold="true" box="[199,314,433,459]" pageId="12" pageNumber="313">Diagnosis</emphasis>
: Most similar to
<taxonomicName id="4C4782A6FFD10828FDFDFE3B821C3EEE" authority="Aubert, 1980" authorityName="Aubert" authorityYear="1980" box="[517,857,434,460]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="bipictor">
<emphasis id="B9332537FFD10828FDFDFE3B83FE3EEE" box="[517,699,434,460]" italics="true" pageId="12" pageNumber="313">Ophion bipictor</emphasis>
Aubert, 1980
</taxonomicName>
(
<figureCitation id="137CE5A0FFD10828FC92FE3B82AF3EEE" box="[874,1002,434,460]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Figs 8D, F</figureCitation>
), but with clypeus narrower, flagellomeres distinctly longer and fewer (
<quantity id="4CBF54C0FFD10828FDDBFE5F83CC3ED2" box="[547,649,470,497]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" metricValueMax="1.6002000000000003" metricValueMin="1.4986000000000002" pageId="12" pageNumber="313" unit="in" value="61.0" valueMax="63.0" valueMin="59.0">59‒63 in</quantity>
<taxonomicName id="4C4782A6FFD10828FD69FE5F82483ED2" authorityName="Aubert" authorityYear="1980" box="[657,781,470,496]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="bipictor">
<emphasis id="B9332537FFD10828FD69FE5F82483ED2" box="[657,781,470,496]" italics="true" pageId="12" pageNumber="313">O. bipictor</emphasis>
</taxonomicName>
). Also potentially similar to
<taxonomicName id="4C4782A6FFD10828FBADFE5F81973D36" authority="Johansson, 2019" authorityName="Johansson" authorityYear="2019" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="sylvestris">
<emphasis id="B9332537FFD10828FBADFE5F845F3ED2" box="[1109,1306,470,496]" italics="true" pageId="12" pageNumber="313">Ophion sylvestris</emphasis>
<bibRefCitation id="EFD684D4FFD10828FADAFE5F81973D36" author="Johansson, N. &amp; Cederberg, B." pageId="12" pageNumber="313" pagination="1 - 136" refId="ref17269" refString="Johansson, N. &amp; Cederberg, B. (2019) Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species. European Journal of Taxonomy, 550, 1 - 136. https: // doi. org / 10.5852 / ejt. 2019.560" type="journal article" year="2019">Johansson, 2019</bibRefCitation>
</taxonomicName>
, but with the carination of propodeum more complete and the apical flagellomeres more elongate. The known female also has the sclerotised part of first sternite ending distinctly anterior to spiracle.
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD10828FF3FFDC883863BE2" pageId="12" pageNumber="313" type="description">
<paragraph id="8BF8F925FFD10828FF3FFDC885743B5A" blockId="12.[151,1437,325,1216]" pageId="12" pageNumber="313">
<emphasis id="B9332537FFD10828FF3FFDC880143D79" bold="true" box="[199,337,577,603]" pageId="12" pageNumber="313">Description</emphasis>
: Male unknown. Fore wing length
<quantity id="4CBF54C0FFD10828FD19FDCB82713D7E" box="[737,820,578,604]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="313" unit="mm" value="12.0">12 mm</quantity>
. Body length
<quantity id="4CBF54C0FFD10828FC29FDCB85663D7E" box="[977,1059,578,604]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="313" unit="mm" value="14.0">14 mm</quantity>
. Antennae with 52 flagellomeres. Temple narrowed behind compound eyes, in lateral view about 0.3 × as long as compound eye (
<figureCitation id="137CE5A0FFD10828FB10FDEF84063DA2" box="[1256,1347,614,640]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Fig. 8A</figureCitation>
). Malar space narrow, 0.1 × the width of the mandibular base. Space between compound eye and lateral ocellus narrow, about 0.1 × the diameter of ocellus (
<figureCitation id="137CE5A0FFD10828FDB8FD2783DE3DEA" box="[576,667,686,712]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Fig. 8C</figureCitation>
). Face relatively narrow (
<figureCitation id="137CE5A0FFD10828FC3DFD27855A3DEA" box="[965,1055,686,712]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Fig. 8C</figureCitation>
), in anterior view, about as wide as compound eye. Occipital carina dorsally slightly angulate. Face below antennal sockets densely punctate on a polished background, interstices between punctures about equal to their diameter. Clypeus with large, slightly scarcer punctures than the face, polished. Mandibular gape acute angled, with quite indistinct internal angles. First flagellomere 3.5 × as long as wide. Central flagellomeres about 2.0 × as long as wide. Mesoscutum polished with very dense and distinct punctures, space between punctures at most equal to their diameter. Mesopleuron polished with very dense and distinct punctures. Epicnemial carina, in antero-ventral view, with pleurosternal angles obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum weakly rugulose with all carinae strong (
<figureCitation id="137CE5A0FFD10828FF15FC4780013CCA" box="[237,324,974,1000]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Fig. 8B</figureCitation>
). Sclerotised part of first sternite ending distinctly anterior to spiracle. Hind trochantellus distinctly shorter than wide in dorsal view. Hind femur slender, about 6.0 × as long as wide. Inner spur of hind tibia about 0.3 × as long as hind metatarsus. Wing membrane clear. Ramellus in fore wing long, reaching 0.4 × the width of the discosubmarginal cell. Outer part of radius weakly sinuous, its inner part straight and not widened before junction with pterostigma. Nervellus broken distinctly below the middle by the discoidella.
</paragraph>
<paragraph id="8BF8F925FFD10828FF3FFB0B83863BE2" blockId="12.[151,1437,325,1216]" pageId="12" pageNumber="313">
<emphasis id="B9332537FFD10828FF3FFB0B80153BBE" box="[199,336,1154,1180]" italics="true" pageId="12" pageNumber="313">Colouration</emphasis>
: Body quite dark, reddish. Inner and outer orbits widely whitish (
<figureCitation id="137CE5A0FFD10828FBD2FB0B85EF3BBE" box="[1066,1194,1154,1180]" captionStart="FIGURE 8" captionStartId="11.[151,250,1783,1807]" captionTargetBox="[151,1436,525,1760]" captionTargetId="figure-158@11.[151,1436,525,1760]" captionTargetPageId="11" captionText="FIGURE 8. A) Ophion albanicus sp. nov. holotype female (NMS), habitus lateral view; B) Ophion albanicus sp. nov. holotype female, propodeum, dorsal view; C) Ophion albanicus sp. nov. holotype female, head, anterior view; D) Ophion bipictor Aubert, 1980, head anterior view; E) Ophion albanicus sp. nov. holotype female, apical part of antenna; F) Ophion bipictor Aubert, 1980, apical part of antenna." figureDoi="http://doi.org/10.5281/zenodo.5226042" httpUri="https://zenodo.org/record/5226042/files/figure.png" pageId="12" pageNumber="313">Figs 8A–C</figureCitation>
). Subalar prominence with white elongate spot. Mandibular teeth black.
</paragraph>
</subSubSection>
</treatment>
</document>
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