treatments-xml/data/03/CD/7B/03CD7B0D1426FF91DBB553BAE906E7C3.xml

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<mods:title id="89C15B378C424DC6690832B9785E1095">MADACHIRONOMUS, A NEW GENUS OF TRIBE PSEUDOCHIRONOMINI (DIPTERA: CHIRONOMIDAE, CHIRONOMINAE) FROM MADAGASCAR Abstract</mods:title>
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<mods:date id="693E8DA4C160B2B50D2826B45E8CCBFC">2016</mods:date>
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<treatment id="03CD7B0D1426FF91DBB553BAE906E7C3" LSID="urn:lsid:plazi:treatment:03CD7B0D1426FF91DBB553BAE906E7C3" httpUri="http://treatment.plazi.org/id/03CD7B0D1426FF91DBB553BAE906E7C3" lastPageId="8" lastPageNumber="23" pageId="3" pageNumber="18">
<subSubSection id="C37E99901426FF9ADBB553BAEACFE490" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="8BDBCA1B1426FF9ADBB553BAE94CE4C3" blockId="3.[227,595,658,682]" box="[227,595,658,682]" pageId="3" pageNumber="18">
<heading id="D0937D771426FF9ADBB553BAE94CE4C3" bold="true" box="[227,595,658,682]" fontSize="10" level="2" pageId="3" pageNumber="18" reason="3">
<emphasis id="B91016091426FF9ADBB553BAE94CE4C3" bold="true" box="[227,595,658,682]" pageId="3" pageNumber="18">
<taxonomicName id="4C64B1981426FF9ADBB553BAE90FE4C3" ID-CoL="8CPRM" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[227,528,658,682]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="lakazana" status="sp. n.">
<emphasis id="B91016091426FF9ADBB553BAE90FE4C3" bold="true" box="[227,528,658,682]" italics="true" pageId="3" pageNumber="18">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721426FF9AD94153BAE94CE4C3" box="[535,595,658,682]" pageId="3" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<emphasis id="B91016091426FF9ADBB553E9EACFE490" italics="true" pageId="3" pageNumber="18">
http://zoobank.org/ 
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</paragraph>
</subSubSection>
<subSubSection id="C37E99901426FF9ADBB55239EA35E260" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BDBCA1B1426FF9ADBB55239EA35E260" blockId="3.[227,764,784,1033]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9ADBB55239EA94E541" bold="true" box="[227,395,784,809]" pageId="3" pageNumber="18">Type material.</emphasis>
<materialsCitation id="3B0CC0461426FF9ADAC25238E974E5A0" collectingDate="1996-11-20" collectionCode="LRSAE, ORSTOM, ZSM" country="Madagascar" county="Analamanga region" latitude="-18.083334" location="Anjafy" longLatPrecision="1276" longitude="47.766666" municipality="Anjozorobe district" pageId="3" pageNumber="18" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="54DF74B91426FF9ADAC25238EAE8E541" box="[404,503,784,808]" pageId="3" pageNumber="18">Holotype</typeStatus>
male, 
<collectingCountry id="F3738A8B1426FF9AD9125238E9D8E541" box="[580,711,784,808]" name="Madagascar" pageId="3" pageNumber="18">Madagascar</collectingCountry>
, 
<collectingRegion id="49A004F91426FF9AD9845239EAA4E521" country="Madagascar" name="Antananarivo" pageId="3" pageNumber="18">Antananarivo province</collectingRegion>
, 
<collectingCounty id="62BAB2971426FF9ADA935218E985E521" box="[453,666,816,840]" pageId="3" pageNumber="18">Analamanga region</collectingCounty>
, 
<collectingMunicipality id="6BBF50611426FF9AD9F25218EA78E501" pageId="3" pageNumber="18">Anjozorobe district</collectingMunicipality>
, 
<location id="8EBB9CC01426FF9ADA275278EAA4E501" LSID="urn:lsid:plazi:treatment:03CD7B0D1426FF91DBB553BAE906E7C3:8EBB9CC01426FF9ADA275278EAA4E501" box="[369,443,848,872]" country="Madagascar" county="Analamanga region" latitude="-18.083334" longLatPrecision="1276" longitude="47.766666" municipality="Anjozorobe district" name="Anjafy" pageId="3" pageNumber="18" stateProvince="Antananarivo">Anjafy</location>
high plains, 
<location id="8EBB9CC01426FF9AD91F5278E9ABE501" LSID="urn:lsid:plazi:treatment:03CD7B0D1426FF91DBB553BAE906E7C3:8EBB9CC01426FF9AD91F5278E9ABE501" box="[585,692,848,872]" country="Madagascar" county="Analamanga region" latitude="-18.083334" longLatPrecision="1276" longitude="47.766666" municipality="Anjozorobe district" name="Betsiboka" pageId="3" pageNumber="18" stateProvince="Antananarivo">Betsiboka</location>
drainage, 
<taxonomicName id="4C64B1981426FF9ADA4D5258EAD7E5E1" authority="River" authorityName="River" box="[283,456,880,904]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="lakazana">Lakazana River</taxonomicName>
at 
<location id="8EBB9CC01426FF9ADAA55258E985E5E1" LSID="urn:lsid:plazi:treatment:03CD7B0D1426FF91DBB553BAE906E7C3:8EBB9CC01426FF9ADAA55258E985E5E1" box="[499,666,880,904]" country="Madagascar" county="Analamanga region" latitude="-18.083334" longLatPrecision="1276" longitude="47.766666" municipality="Anjozorobe district" name="Ankondondona" pageId="3" pageNumber="18" stateProvince="Antananarivo">Ankondondona</location>
, approx. 
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<geoCoordinate id="EE50ACDC1426FF9ADA1D52B8EABAE5C1" box="[331,421,912,936]" degrees="18" direction="south" minutes="05" orientation="latitude" pageId="3" pageNumber="18" precision="925" value="-18.083334">18°05’S</geoCoordinate>
, evening of 
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<collectingDate id="EF9E15331426FF9AD91452B8E9A8E5C1" box="[578,695,912,936]" pageId="3" pageNumber="18" value="1996-11-20">20.xi.1996</collectingDate>
</date>
, 
<collectingMethod id="5225B20C1426FF9AD99D52B8EA10E5A1" pageId="3" pageNumber="18">light trap</collectingMethod>
, leg. 
<collectionCode id="ED7552DE1426FF9ADA1D5299EA80E5A0" box="[331,415,945,969]" pageId="3" pageNumber="18">LRSAE</collectionCode>
/ 
<collectionCode id="ED7552DE1426FF9ADAFE5299E909E5A0" box="[424,534,945,969]" country="Bolivia" httpUri="http://grbio.org/cool/mdr4-z094" name="Office de la Recherche scientifique et Technique Outre-mer" pageId="3" pageNumber="18">ORSTOM</collectionCode>
(
<collectionCode id="ED7552DE1426FF9AD9705299E97DE5A0" box="[550,610,945,969]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="18" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B0CC0461426FF9AD9205299EA39E260" collectingDate="1996-11-20" collectionCode="ZMBN, ZSM" country="Madagascar" county="Analamanga region" latitude="-18.083334" location="Anjafy" longLatPrecision="1276" longitude="47.766666" municipality="Anjozorobe district" pageId="3" pageNumber="18" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="54DF74B91426FF9AD9205299E9FDE5A0" box="[630,738,945,969]" pageId="3" pageNumber="18">Paratypes</typeStatus>
: 
<specimenCount id="9D6201921426FF9AD9B85299EA06E581" pageId="3" pageNumber="18" type="male">1 male</specimenCount>
, 
<specimenCount id="9D6201921426FF9ADA7152F9EA8BE581" box="[295,404,976,1001]" pageId="3" pageNumber="18" type="female">2 females</specimenCount>
, same data as holotype (
<collectionCode id="ED7552DE1426FF9AD9F052F9E9E8E580" box="[678,759,977,1001]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="3" pageNumber="18">ZMBN</collectionCode>
, 
<collectionCode id="ED7552DE1426FF9ADBB552D9EA01E260" box="[227,286,1009,1033]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="18" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E99901426FF9ADBB55508EAE2E3AE" pageId="3" pageNumber="18" type="etymology">
<paragraph id="8BDBCA1B1426FF9ADBB55508EAAEE2F1" blockId="3.[227,764,1056,1176]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9ADBB55508EA7CE251" bold="true" box="[227,355,1056,1080]" pageId="3" pageNumber="18">Etymology.</emphasis>
Named after 
<taxonomicName id="4C64B1981426FF9AD9555508E9AEE251" authority="River" authorityName="River" box="[515,689,1056,1080]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="lakazana">Lakazana River</taxonomicName>
, 
<collectingRegion id="49A004F91426FF9AD9E95508EAB8E231" country="Madagascar" name="Antananarivo" pageId="3" pageNumber="18">Antananarivo province</collectingRegion>
, 
<collectingCountry id="F3738A8B1426FF9ADAE45568E92AE231" box="[434,565,1088,1112]" name="Madagascar" pageId="3" pageNumber="18">Madagascar</collectingCountry>
, where the species was collected. The name is to be regarded as a noun in apposition.
</paragraph>
<paragraph id="8BDBCA1B1426FF9ADBB55586EAE2E3AE" blockId="3.[227,764,1198,1479]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9ADBB55586EAFDE2AE" bold="true" box="[227,482,1198,1223]" pageId="3" pageNumber="18">Diagnostic characters.</emphasis>
See diagnostic characters for the genus. The female can be separated from the female of 
<taxonomicName id="4C64B1981426FF9ADA2D55C7E902E36F" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[379,541,1263,1286]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="rongaronga" status="sp. n.">
<emphasis id="B91016091426FF9ADA2D55C7E902E36F" box="[379,541,1263,1286]" italics="true" pageId="3" pageNumber="18">M. rongaronga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721426FF9AD97055C7E97EE36F" box="[550,609,1263,1286]" pageId="3" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
as it is larger, with a wing length of 
<quantity id="4C9C67FE1426FF9ADA855427E971E34F" box="[467,622,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="4.17" metricValueMax="4.18" metricValueMin="4.16" pageId="3" pageNumber="18" unit="mm" value="4.17" valueMax="4.18" valueMin="4.16">4.16–4.18 mm</quantity>
compared to 
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in 
<emphasis id="B91016091426FF9ADAFB5407E945E32F" box="[429,602,1327,1350]" italics="true" pageId="3" pageNumber="18">
<taxonomicName id="4C64B1981426FF9ADAFB5407E949E32F" authorityName="Andersen" authorityYear="2016" box="[429,598,1327,1350]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="rongaronga">M. rongaronga</taxonomicName>
,
</emphasis>
has a slightly lower antennal ratio (AR = 0.58–0.66 compared to AR = 0.70–0.81) and has distinctly more setae on segment X to each side of vagina (189–231 setae compared to 53–78 setae).
</paragraph>
</subSubSection>
<subSubSection id="C37E99901426FF91DBB554F6E906E7C3" lastPageId="8" lastPageNumber="23" pageId="3" pageNumber="18" type="description">
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<emphasis id="B91016091426FF9ADBB554F6EA78E39F" bold="true" box="[227,359,1502,1526]" pageId="3" pageNumber="18">Description</emphasis>
</paragraph>
<paragraph id="8BDBCA1B1426FF9ADBB55725EA98E0EC" blockId="3.[227,764,1549,1669]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9ADBB55725EA7EE04C" bold="true" box="[227,353,1549,1573]" pageId="3" pageNumber="18">Adult male</emphasis>
(n = 2–3). Total length 
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. Wing length 
<quantity id="4C9C67FE1426FF9ADA235705E907E02D" box="[373,536,1581,1605]" metricMagnitude="-3" metricUnit="m" metricValue="4.14" metricValueMax="4.16" metricValueMin="4.12" pageId="3" pageNumber="18" unit="mm" value="4.14" valueMax="4.16" valueMin="4.12">4.12–4.16 mm</quantity>
. Total length / wing length 2.23–2.27. Wing length / length of profe- mur 2.32–2.35.
</paragraph>
<paragraph id="8BDBCA1B1426FF9ADBB557B4EF72E712" blockId="3.[227,764,1692,2010]" lastBlockId="3.[824,1361,227,379]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9ADBB557B4EA40E0DD" box="[227,351,1692,1716]" italics="true" pageId="3" pageNumber="18">Coloration.</emphasis>
Head, antennae and palpi brown. Thorax mostly brown with lateral mesonotal dark brown spot. Legs medium brown, foreleg with tibia and ta 
<subScript id="17E0C85E1426FF9ADA375624EA69E173" attach="left" box="[353,374,1804,1818]" fontSize="6" pageId="3" pageNumber="18">2-5</subScript>
brown, fore tibia with dark brown apex, fore ta 
<subScript id="17E0C85E1426FF9ADA315604EA70E153" attach="left" box="[359,367,1836,1850]" fontSize="6" pageId="3" pageNumber="18">1</subScript>
lighter brown with dark brown apex; mid- and hind legs with lighter brown tarsi. Wing membrane (
<figureCitation id="135FD69E1426FF9ADA375674EABFE11D" box="[353,416,1884,1908]" captionStart="Figures 8-9" captionStartId="6.[227,299,851,873]" captionTargetBox="[315,1273,227,834]" captionTargetId="figure-2@6.[315,1273,227,834]" captionTargetPageId="6" captionText="Figures 8-9. Madachironomus gen. n., wings. 8) M. lakazana sp. n., male; 9) M. rongaronga sp. n., female (photo Torbjørn Ekrem)." figureDoi="http://doi.org/10.5281/zenodo.7987251" httpUri="https://zenodo.org/record/7987251/files/figure.png" pageId="3" pageNumber="18">Fig. 8</figureCitation>
) translucent with brownish stain and some light shaded areas e.g. proximally and distally in cell c, along most of sc, proximal in r 
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and along Cu and proximal parts of M 
<subScript id="17E0C85E1426FF9AD9E256E4E9D3E1B3" attach="left" box="[692,716,1996,2010]" fontSize="6" pageId="3" pageNumber="18">3+4</subScript>
and Cu 
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; wing veins brownish, crossvein 
<collectingRegion id="49A004F91426FF9ADF8751CCEFE5E692" box="[1233,1274,228,251]" country="Peru" name="Lima" pageId="3" pageNumber="18">RM</collectingRegion>
and radial fork darker brown. Abdominal segment 1 pale brown, abdominal segments 2–5 light brown with narrow anterior transverse brown band; segments 6–8 and hypopygium brown.
</paragraph>
<paragraph id="8BDBCA1B1426FF9AD86E50BBEF07E7A3" blockId="3.[824,1360,402,459]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E50BBE887E7C3" box="[824,920,403,426]" italics="true" pageId="3" pageNumber="18">Antenna.</emphasis>
AR 2.37–2.49. Terminal flagellomere 1503–1560 µm long.
</paragraph>
<paragraph id="8BDBCA1B1426FF9AD86E50C9EE20E570" blockId="3.[824,1361,481,794]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E50C9E86DE790" box="[824,882,481,505]" italics="true" pageId="3" pageNumber="18">Head</emphasis>
(
<figureCitation id="135FD69E1426FF9AD8D650C9E8A0E790" box="[896,959,481,505]" captionStart="Figures 1-5" captionStartId="4.[227,299,1317,1339]" captionTargetBox="[336,1263,247,1297]" captionTargetId="figure-1@4.[313,1274,227,1380]" captionTargetPageId="4" captionText="Figures 1-5. Madachironomus lakazana gen. n., sp. n., male. 1) head; 2) tentorium, stipes and cibarial pump; 3) thorax; 4) comb of foreleg; 5) combs of hind leg." figureDoi="http://doi.org/10.5281/zenodo.7987245" httpUri="https://zenodo.org/record/7987245/files/figure.png" pageId="3" pageNumber="18">Fig. 1</figureCitation>
). Temporal setae 22–25, briefly bi- to tri-serial near transition of eye to its dorsomedial extension, consisting of inseparably intergrading verticals and postorbitals. Clypeus with 62–68 setae. Tentorium, stipes and cibarial pump as in 
<figureCitation id="135FD69E1426FF9AD86E53A9E88AE4F0" box="[824,917,641,665]" captionStart="Figures 1-5" captionStartId="4.[227,299,1317,1339]" captionTargetBox="[336,1263,247,1297]" captionTargetId="figure-1@4.[313,1274,227,1380]" captionTargetPageId="4" captionText="Figures 1-5. Madachironomus lakazana gen. n., sp. n., male. 1) head; 2) tentorium, stipes and cibarial pump; 3) thorax; 4) comb of foreleg; 5) combs of hind leg." figureDoi="http://doi.org/10.5281/zenodo.7987245" httpUri="https://zenodo.org/record/7987245/files/figure.png" pageId="3" pageNumber="18">Figure 2</figureCitation>
. Tentorium 277–312 µm long, 82–90 µm wide. Stipes 242–267 µm long, 21–25 µm wide. Palpomere lengths (in µm): 76–92, 112–128, 396– 412, 380–384, 640–651. Third palpomere with 2–3 sensilla clavata apically, longest 19–25 µm long.
</paragraph>
<paragraph id="8BDBCA1B1426FF9AD86E5218EF43E5A1" blockId="3.[824,1361,816,969]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E5218E89AE521" box="[824,901,816,840]" italics="true" pageId="3" pageNumber="18">Thorax</emphasis>
(
<figureCitation id="135FD69E1426FF9AD8CF5218E8C0E520" box="[921,991,816,841]" captionStart="Figures 1-5" captionStartId="4.[227,299,1317,1339]" captionTargetBox="[336,1263,247,1297]" captionTargetId="figure-1@4.[313,1274,227,1380]" captionTargetPageId="4" captionText="Figures 1-5. Madachironomus lakazana gen. n., sp. n., male. 1) head; 2) tentorium, stipes and cibarial pump; 3) thorax; 4) comb of foreleg; 5) combs of hind leg." figureDoi="http://doi.org/10.5281/zenodo.7987245" httpUri="https://zenodo.org/record/7987245/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
). Antepronotum with 4–6 dorsal and 15–17 ventrolateral setae. Acrostichals appar- ently about 40; dorsocentrals 42–45, weak; pre- alars 3–11; supraalars 0–1. Scutellum with 48–52 setae in 2–3 irregular rows.
</paragraph>
<paragraph id="8BDBCA1B1426FF9AD86E52C8EE22E211" blockId="3.[824,1361,992,1144]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E52C8E870E59E" box="[824,879,992,1015]" italics="true" pageId="3" pageNumber="18">Wing</emphasis>
(
<figureCitation id="135FD69E1426FF9AD82852C8E8DFE591" box="[894,960,992,1016]" captionStart="Figures 8-9" captionStartId="6.[227,299,851,873]" captionTargetBox="[315,1273,227,834]" captionTargetId="figure-2@6.[315,1273,227,834]" captionTargetPageId="6" captionText="Figures 8-9. Madachironomus gen. n., wings. 8) M. lakazana sp. n., male; 9) M. rongaronga sp. n., female (photo Torbjørn Ekrem)." figureDoi="http://doi.org/10.5281/zenodo.7987251" httpUri="https://zenodo.org/record/7987251/files/figure.png" pageId="3" pageNumber="18">Fig. 8</figureCitation>
). VR 0.98–0.99. C extension 72–109 µm long. Brachiolum with 2–3 setae; C extension with 4–6 non-marginal setae; R 
<subScript id="17E0C85E1426FF9ADFCE5518EFAFE257" attach="left" box="[1176,1200,1072,1086]" fontSize="6" pageId="3" pageNumber="18">4+5</subScript>
with 0–1 seta apically; other veins bare. Wing membrane bare. Squama with 28–36 setae, partly bi- to tri-serial.
</paragraph>
<paragraph id="8BDBCA1B1426FF9AD86E55A7EFDCE3CE" blockId="3.[824,1361,1167,1447]" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E55A7E875E2CF" box="[824,874,1167,1190]" italics="true" pageId="3" pageNumber="18">Legs</emphasis>
(
<figureCitation id="135FD69E1426FF9AD82E55A7E8C6E2CE" box="[888,985,1167,1191]" captionStart="Figures 1-5" captionStartId="4.[227,299,1317,1339]" captionTargetBox="[336,1263,247,1297]" captionTargetId="figure-1@4.[313,1274,227,1380]" captionTargetPageId="4" captionText="Figures 1-5. Madachironomus lakazana gen. n., sp. n., male. 1) head; 2) tentorium, stipes and cibarial pump; 3) thorax; 4) comb of foreleg; 5) combs of hind leg." figureDoi="http://doi.org/10.5281/zenodo.7987245" httpUri="https://zenodo.org/record/7987245/files/figure.png" pageId="3" pageNumber="18">Figs 4–5</figureCitation>
). Spur of fore tibia 80–89 µm long; spurs of mid tibia 100–115 µm and 103–121 µm long; spurs of hind tibia 105–121 µm and 113–127 µm long. Width at apex of fore tibia 111–113 µm; of mid tibia 127–131 µm; of hind tibia 135–139 µm. Mid ta 
<subScript id="17E0C85E1426FF9AD8E05417E8A1E324" attach="left" box="[950,958,1343,1357]" fontSize="6" pageId="3" pageNumber="18">1</subScript>
with about 15 sensilla chaetica in 3 rows in proximal 1/3, hind ta 
<subScript id="17E0C85E1426FF9ADF205477EF61E304" attach="left" box="[1142,1150,1375,1389]" fontSize="6" pageId="3" pageNumber="18">1</subScript>
with about 25 sensilla chaetica in 3 rows in proximal 1/3. Lengths and proportions of legs as in 
<tableCitation id="C6E6FFA01426FF9ADF3B54A7EFA2E3CE" box="[1133,1213,1423,1447]" captionStart="Table 1" captionStartId="4.[227,280,1408,1430]" captionText="Table 1. Lengths (in µm) and proportions of legs of Madachironomus lakazana gen. n., sp. n., male (n = 2–3)." pageId="3" pageNumber="18">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BDBCA1B1426FF9DD86E5497E97EE134" blockId="3.[824,1361,1470,2006]" lastBlockId="4.[227,764,1733,1885]" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
<emphasis id="B91016091426FF9AD86E5497E8DFE3BF" box="[824,960,1471,1494]" italics="true" pageId="3" pageNumber="18">Hypopygium</emphasis>
(
<figureCitation id="135FD69E1426FF9AD8985496EF30E3BF" box="[974,1071,1470,1494]" captionStart="Figures 6-7" captionStartId="5.[227,299,1221,1243]" captionTargetBox="[311,1275,314,1116]" captionTargetId="figure-312@5.[291,1296,227,1232]" captionTargetPageId="5" captionText="Figures 6-7. Madachironomus lakazana gen. n., sp. n., male. 6) hypopygium, dorsal view; 7) hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.7987249" httpUri="https://zenodo.org/record/7987249/files/figure.png" pageId="3" pageNumber="18">Figs 6–7</figureCitation>
). Tergite IX with 13–17 se- tae along posterior margin to each side of base of anal point; laterosternite IX with 15–23 setae. Anal point tapering to tongue-shaped apex, 115–123 µm long, 66–75 µm wide at base, 29–35 µm wide sub- apically. Phallapodeme 316–324 µm long, includ- ing 88–96 µm long, 8–12 µm wide, curved oral projection. Transverse sternapodeme 120–128 µm long. Gonocoxite 380–404 µm long. Median vol- sella with main stem split medially in two digitiform projections; longest, caudal branch 72–80 µm long, 16–20 µm wide medially; shortest, oral branch 66–84 µm long; densely covered with sub- ulate setae up to 66–85 µm in length. Superior vol- sella 92–108 µm long, including 34–38 µm long, hooked apical portion; with 1–2 strong, 28–30 µm long setae dorsolaterally, hooked apical portion with 3–6 medial to ventral setae. Inferior volsella broadly digitiform, 208–212 µm long, 80–100 µm wide at base, 48–56 µm wide medially, with 63–82 normal to strong setae. Gonostylus 328–344 µm long. HR 1.11–1.20. HV 2.74–2.80.
</paragraph>
<caption id="DF1B9A931421FF9DDBB5540DE96FE331" ID-DOI="http://doi.org/10.5281/zenodo.7987245" ID-Zenodo-Dep="7987245" httpUri="https://zenodo.org/record/7987245/files/figure.png" pageId="4" pageNumber="19" startId="4.[227,299,1317,1339]" targetBox="[336,1263,247,1297]" targetPageId="4" targetType="figure">
<paragraph id="8BDBCA1B1421FF9DDBB5540DE96FE331" blockId="4.[227,1361,1317,1368]" pageId="4" pageNumber="19">
Figures 1-5. 
<taxonomicName id="4C64B1981421FF9DDA0A540DE97EE353" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[348,609,1317,1338]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091421FF9DDA0A540DE97EE353" box="[348,609,1317,1338]" italics="true" pageId="4" pageNumber="19">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721421FF9DD930540EE9F9E352" box="[614,742,1318,1339]" pageId="4" pageNumber="19" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, male. 1) head; 2) tentorium, stipes and cibarial pump; 3) thorax; 4) comb of foreleg; 5) combs of hind leg.
</paragraph>
</caption>
<caption id="DF1B9A931421FF9DDBB554A8EE1DE3FF" box="[227,1282,1408,1430]" pageId="4" pageNumber="19" startId="4.[227,280,1408,1430]" targetType="table">
<paragraph id="8BDBCA1B1421FF9DDBB554A8EE1DE3FF" blockId="4.[227,1282,1408,1430]" box="[227,1282,1408,1430]" pageId="4" pageNumber="19">
Table 1. Lengths (in µm) and proportions of legs of 
<taxonomicName id="4C64B1981421FF9DD98054A8E8C3E3FC" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[726,988,1408,1429]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091421FF9DD98054A8E8C3E3FC" box="[726,988,1408,1429]" italics="true" pageId="4" pageNumber="19">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721421FF9DD8B754A9EF7CE3FF" box="[993,1123,1409,1430]" pageId="4" pageNumber="19" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, male (n = 2–3).
</paragraph>
</caption>
<paragraph id="8BDBCA1B1421FF9DDA4654E1EE31E0F4" pageId="4" pageNumber="19">
<table id="F96438BB14210066DA5A548EEE30E0CA" box="[268,1327,1446,1699]" gridcols="7" gridrows="2" pageId="4" pageNumber="19">
<tr id="3554C85914210066DA5A548EEE30E04A" box="[268,1327,1446,1571]" gridrow="0" pageId="4" pageNumber="19">
<th id="7685A12514210066DA5A548EEA38E04A" box="[268,295,1446,1571]" gridcol="0" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDA4654E1EA00E388" bold="true" box="[272,287,1481,1505]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA4954F1EA38E38E" bold="true" box="[287,295,1497,1511]" pageId="4" pageNumber="19">1</emphasis>
<emphasis id="B91016091421FF9DDA4654CFEA00E396" bold="true" box="[272,287,1511,1535]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA4954DFEA38E06C" bold="true" box="[287,295,1527,1541]" pageId="4" pageNumber="19">2</emphasis>
<emphasis id="B91016091421FF9DDA46572DEA00E074" bold="true" box="[272,287,1541,1565]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA49573DEA38E04A" bold="true" box="[287,295,1557,1571]" pageId="4" pageNumber="19">3</emphasis>
</th>
<td id="7685A12514210066DA3C548EEAFDE04A" box="[362,482,1446,1571]" gridcol="1" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDACD548EEAAFE3D7" bold="true" box="[411,432,1446,1470]" pageId="4" pageNumber="19">fe</emphasis>
1730–1812 2266–2307 2369–2410
</td>
<td id="7685A12514210066D946548EE997E04A" box="[528,648,1446,1571]" gridcol="2" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DD912548EE94BE3D7" bold="true" box="[580,596,1446,1470]" pageId="4" pageNumber="19">ti</emphasis>
2368–2410 2575–2657 2822–2843
</td>
<td id="7685A12514210066D9EC548EE82DE04A" box="[698,818,1446,1571]" gridcol="3" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DD9B1548EE9E2E3D7" bold="true" box="[743,765,1446,1470]" pageId="4" pageNumber="19">ta</emphasis>
<emphasis id="B91016091421FF9DD9AB549EE81AE3AD" bold="true" box="[765,773,1462,1476]" pageId="4" pageNumber="19">1</emphasis>
1854–1957 1009–1092 1215–1318
</td>
<td id="7685A12514210066D838548EE8D3E04A" box="[878,972,1446,1571]" gridcol="4" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DD8D8548EE8BBE3D7" bold="true" box="[910,932,1446,1470]" pageId="4" pageNumber="19">ta</emphasis>
<emphasis id="B91016091421FF9DD8F2549EE8B3E3AD" bold="true" box="[932,940,1462,1476]" pageId="4" pageNumber="19">2</emphasis>
741–824 556–577 638–701
</td>
<td id="7685A12514210066D8B1548EEF6DE04A" box="[999,1138,1446,1571]" gridcol="5" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDF62548EEF55E3D7" bold="true" box="[1076,1098,1446,1470]" pageId="4" pageNumber="19">ta</emphasis>
<emphasis id="B91016091421FF9DDF1D549EEF4CE3AD" bold="true" box="[1099,1107,1462,1476]" pageId="4" pageNumber="19">3</emphasis>
597–698 494–536 577–639
</td>
<td id="7685A12514210066DFFF548EEE30E04A" box="[1193,1327,1446,1571]" gridcol="6" gridrow="0" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDFB3548EEFE4E3D7" bold="true" box="[1253,1275,1446,1470]" pageId="4" pageNumber="19">ta</emphasis>
<emphasis id="B91016091421FF9DDFAD549EEE1CE3AD" bold="true" box="[1275,1283,1462,1476]" pageId="4" pageNumber="19">4</emphasis>
494–556 309–330 330–391
</td>
</tr>
<tr id="3554C85914210066DA5A570FEE30E0CA" box="[268,1327,1575,1699]" gridrow="1" pageId="4" pageNumber="19">
<th id="7685A12514210066DA5A570FEA38E0CA" box="[268,295,1575,1699]" gridcol="0" gridrow="1" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDA5A5762EA04E00B" bold="true" box="[268,283,1610,1634]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA4D5772EA3CE001" bold="true" box="[283,291,1626,1640]" pageId="4" pageNumber="19">1</emphasis>
<emphasis id="B91016091421FF9DDA5A5740EA04E0E9" bold="true" box="[268,283,1640,1664]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA4D575FEA3CE0EC" bold="true" box="[283,291,1655,1669]" pageId="4" pageNumber="19">2</emphasis>
<emphasis id="B91016091421FF9DDA5A57ADEA04E0F4" bold="true" box="[268,283,1669,1693]" pageId="4" pageNumber="19">p</emphasis>
<emphasis id="B91016091421FF9DDA4D57BDEA3CE0CA" bold="true" box="[283,291,1685,1699]" pageId="4" pageNumber="19">3</emphasis>
</th>
<td id="7685A12514210066DA3C570FEAFDE0CA" box="[362,482,1575,1699]" gridcol="1" gridrow="1" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDACE570FEAB1E056" bold="true" box="[408,430,1575,1599]" pageId="4" pageNumber="19">ta</emphasis>
<emphasis id="B91016091421FF9DDAF8571FEAA9E02C" bold="true" box="[430,438,1591,1605]" pageId="4" pageNumber="19">5</emphasis>
206–247 144–165 154–165
</td>
<td id="7685A12514210066D946570FE8D3E0CA" box="[528,972,1575,1699]" colspan="3" colspanRight="2" gridcol="2" gridrow="1" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DD930570FE86EE056" bold="true" box="[614,881,1575,1599]" pageId="4" pageNumber="19">LR BV</emphasis>
0.769–0.826 2.661–2.949 0.380–0.424 3.753–3.959 0.431–0.467 3.467–3.747
</td>
<td id="7685A12514210066D8B1570FEF6DE0CA" box="[999,1138,1575,1699]" gridcol="5" gridrow="1" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDF4F570FEF24E056" bold="true" box="[1049,1083,1575,1599]" pageId="4" pageNumber="19">SV</emphasis>
2.137–2.244 4.434–4.898 3.938–4.271
</td>
<td id="7685A12514210066DFFF570FEE30E0CA" box="[1193,1327,1575,1699]" gridcol="6" gridrow="1" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDF8C570FEFE0E056" bold="true" box="[1242,1279,1575,1599]" pageId="4" pageNumber="19">BR</emphasis>
1.813–1.866 2.125–2.166 2.211–2.500
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDBCA1B1421FF9DDBB5565CE9E3E1A5" blockId="4.[227,764,1908,1996]" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DDBB5565CEA68E1E5" bold="true" box="[227,375,1908,1932]" pageId="4" pageNumber="19">Adult female</emphasis>
(n = 2). Total length 
<quantity id="4C9C67FE1421FF9DD90C565CE9E7E1E5" box="[602,760,1908,1932]" metricMagnitude="-3" metricUnit="m" metricValue="9.879999999999999" metricValueMax="9.93" metricValueMin="9.83" pageId="4" pageNumber="19" unit="mm" value="9.879999999999999" valueMax="9.93" valueMin="9.83">9.83–9.93 mm</quantity>
. Wing length 
<quantity id="4C9C67FE1421FF9DDA2356BCE907E1C5" box="[373,536,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="4.17" metricValueMax="4.18" metricValueMin="4.16" pageId="4" pageNumber="19" unit="mm" value="4.17" valueMax="4.18" valueMin="4.16">4.16–4.18 mm</quantity>
. Total length / wing length 2.36–2.39. Wing length / length of profe-
</paragraph>
<paragraph id="8BDBCA1B1421FF9DD86E57EEE8C3E0B4" blockId="4.[824,988,1733,1757]" box="[824,988,1733,1757]" pageId="4" pageNumber="19">mur 2.39–2.43.</paragraph>
<paragraph id="8BDBCA1B1421FF9CD86E57DCEA31E004" blockId="4.[824,1361,1780,1996]" lastBlockId="5.[227,764,1301,1645]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<emphasis id="B91016091421FF9DD86E57DCE8ABE165" box="[824,948,1780,1804]" italics="true" pageId="4" pageNumber="19">Coloration.</emphasis>
Generally distinctly darker than male. Head, antennae and palpi brown. Thorax mostly brown, lateral mesonotal dark spot less contrasting than in male, indistinct in some specimens. Legs medium brown; foreleg with tibia and ta 
<subScript id="17E0C85E1421FF9DDFBB56ACEE1AE1FB" attach="left" box="[1261,1285,1924,1938]" fontSize="6" pageId="4" pageNumber="19">2–5</subScript>
darker brown, fore tibia with dark brown apex, fore ta 1 lighter brown with dark brown apex; mid- and hind legs with lighter brown femoral apices and tarsi. Wing membrane translucent with brownish stain and some more darkly shaded areas, e.g. proximally and distally in cell c, along most of sc, proximally in r 
<subScript id="17E0C85E1420FF9CDADF548DEA8EE3DA" attach="left" box="[393,401,1445,1459]" fontSize="6" pageId="5" pageNumber="20">1</subScript>
and r 
<subScript id="17E0C85E1420FF9CDA82548DEAF3E3DA" attach="left" box="[468,492,1445,1459]" fontSize="6" pageId="5" pageNumber="20">4+5</subScript>
, along Cu and proximal parts of M 
<subScript id="17E0C85E1420FF9CDA0554EDEA74E3BA" attach="left" box="[339,363,1477,1491]" fontSize="6" pageId="5" pageNumber="20">3+4</subScript>
and Cu 
<subScript id="17E0C85E1420FF9CDAEB54EDEADAE3BA" attach="left" box="[445,453,1477,1491]" fontSize="6" pageId="5" pageNumber="20">1</subScript>
; veins brown, crossvein 
<collectingRegion id="49A004F91420FF9CD984549DE9E4E3A5" box="[722,763,1461,1484]" country="Peru" name="Lima" pageId="5" pageNumber="20">RM</collectingRegion>
and radial fork darker brown. Abdominal tergites and posterior sternites brown, anterior sternites light brown, anterior transverse segment bands indicated in some specimens; hypopygium medium brown.
</paragraph>
<caption id="DF1B9A931420FF9CDBB555EDEF12E291" ID-DOI="http://doi.org/10.5281/zenodo.7987249" ID-Zenodo-Dep="7987249" httpUri="https://zenodo.org/record/7987249/files/figure.png" pageId="5" pageNumber="20" startId="5.[227,299,1221,1243]" targetBox="[311,1275,314,1116]" targetPageId="5" targetType="figure">
<paragraph id="8BDBCA1B1420FF9CDBB555EDEF12E291" blockId="5.[227,1361,1221,1272]" pageId="5" pageNumber="20">
Figures 6-7. 
<taxonomicName id="4C64B1981420FF9CDA0955EDE979E2B3" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[351,614,1221,1242]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091420FF9CDA0955EDE979E2B3" box="[351,614,1221,1242]" italics="true" pageId="5" pageNumber="20">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721420FF9CD93B55EEE9ECE2B2" box="[621,755,1222,1243]" pageId="5" pageNumber="20" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, male. 6) hypopygium, dorsal view; 7) hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right.
</paragraph>
</caption>
<paragraph id="8BDBCA1B1420FF9CDBB557ACE9AAE1F5" blockId="5.[227,764,1668,1948]" pageId="5" pageNumber="20">
<emphasis id="B91016091420FF9CDBB557ACEA23E0F2" box="[227,316,1668,1691]" italics="true" pageId="5" pageNumber="20">Antenna</emphasis>
(
<figureCitation id="135FD69E1420FF9CDA0357ACEAB0E0F5" box="[341,431,1668,1692]" captionStart="Figures 10-13" captionStartId="6.[227,299,1999,2021]" captionTargetBox="[327,1279,951,1974]" captionTargetId="figure-1@6.[294,1296,894,2030]" captionTargetPageId="6" captionText="Figures 10-13. Madachironomus lakazana gen. n., sp. n., female. 10) head; 11) tentorium, stipes and cibarial pump; 12) antenna; 13) thorax." figureDoi="http://doi.org/10.5281/zenodo.7987255" httpUri="https://zenodo.org/record/7987255/files/figure.png" pageId="5" pageNumber="20">Fig. 12</figureCitation>
). With 6 flagellomeres; AR = 0.58–0.66. Length and width of pedicel and flagellomeres 1–6 (in µm) as: 80–92 / 128–132, 100–104 / 56–60, 84–88 / 40–44, 92–104 / 40–44, 96–104 / 38–42, 88–92 / 36–40, 272–300 / 28–32. Flagellomeres 1–5 with ring of sensilla chaetica subapically, flagellomere 6 with sensilla chaetica in apical 2/3. Flagellomere 6 with 1–2 strong setae subapically, longest 118–148, 133 µm long.
</paragraph>
<paragraph id="8BDBCA1B1420FF9CDBB5569BE886E024" blockId="5.[227,764,1971,1995]" lastBlockId="5.[824,1361,1301,1613]" pageId="5" pageNumber="20">
<emphasis id="B91016091420FF9CDBB5569BEA02E1A2" box="[227,285,1971,1995]" italics="true" pageId="5" pageNumber="20">Head</emphasis>
(
<figureCitation id="135FD69E1420FF9CDA7B569BEA61E1A2" box="[301,382,1971,1995]" captionStart="Figures 10-13" captionStartId="6.[227,299,1999,2021]" captionTargetBox="[327,1279,951,1974]" captionTargetId="figure-1@6.[294,1296,894,2030]" captionTargetPageId="6" captionText="Figures 10-13. Madachironomus lakazana gen. n., sp. n., female. 10) head; 11) tentorium, stipes and cibarial pump; 12) antenna; 13) thorax." figureDoi="http://doi.org/10.5281/zenodo.7987255" httpUri="https://zenodo.org/record/7987255/files/figure.png" pageId="5" pageNumber="20">Fig. 10</figureCitation>
). Temporal setae 22–24, briefly bito tri-serial near transition of eye to its dorsomedial extension, consisting of inseparably intergrading verticals and postorbitals. Clypeus with 105–112 setae. Tentorium, stipes and cibarial pump as in 
<figureCitation id="135FD69E1420FF9CD86E54BDE880E3C4" box="[824,927,1429,1453]" captionStart="Figures 10-13" captionStartId="6.[227,299,1999,2021]" captionTargetBox="[327,1279,951,1974]" captionTargetId="figure-1@6.[294,1296,894,2030]" captionTargetPageId="6" captionText="Figures 10-13. Madachironomus lakazana gen. n., sp. n., female. 10) head; 11) tentorium, stipes and cibarial pump; 12) antenna; 13) thorax." figureDoi="http://doi.org/10.5281/zenodo.7987255" httpUri="https://zenodo.org/record/7987255/files/figure.png" pageId="5" pageNumber="20">Figure 11</figureCitation>
. Tentorium 304–316 µm long, 76–80 µm wide. Stipes 272–312 µm long, 24–28 µm wide. Palp segment lengths (in µm): 87–98, 108–114, 376–380, 380–388, 556–588. Third palpomere with 2–3 sensilla clavata apically, longest 21–25 µm long.
</paragraph>
<paragraph id="8BDBCA1B1420FF9CD86E574CE8E6E092" blockId="5.[824,1361,1636,1788]" pageId="5" pageNumber="20">
<emphasis id="B91016091420FF9CD86E574CE89AE015" box="[824,901,1636,1660]" italics="true" pageId="5" pageNumber="20">Thorax</emphasis>
(
<figureCitation id="135FD69E1420FF9CD8C1574CE8F6E015" box="[919,1001,1636,1660]" captionStart="Figures 10-13" captionStartId="6.[227,299,1999,2021]" captionTargetBox="[327,1279,951,1974]" captionTargetId="figure-1@6.[294,1296,894,2030]" captionTargetPageId="6" captionText="Figures 10-13. Madachironomus lakazana gen. n., sp. n., female. 10) head; 11) tentorium, stipes and cibarial pump; 12) antenna; 13) thorax." figureDoi="http://doi.org/10.5281/zenodo.7987255" httpUri="https://zenodo.org/record/7987255/files/figure.png" pageId="5" pageNumber="20">Fig. 13</figureCitation>
). Antepronotum with 6–9 dorsal and 14–15 ventrolateral setae. Acrostichals appar- ently about 40; dorsocentrals 36–41 weak, in 1–3 irregular rows; prealars 7–8. Scutellum with 44–48 setae in 2–3 rows.
</paragraph>
<paragraph id="8BDBCA1B1420FF9CD86E563CEF47E1A2" blockId="5.[824,1361,1811,1995]" pageId="5" pageNumber="20">
<emphasis id="B91016091420FF9CD86E563CE870E142" box="[824,879,1812,1835]" italics="true" pageId="5" pageNumber="20">Wing</emphasis>
. VR 1.01–1.07. C extension about 60 µm long. Brachiolum with 2 setae, C extension with 2–5 non-marginal setae, R 
<subScript id="17E0C85E1420FF9CDF06564BEF47E118" attach="left" box="[1104,1112,1891,1905]" fontSize="6" pageId="5" pageNumber="20">1</subScript>
with 8–12 setae in api- cal 1/3, R 
<subScript id="17E0C85E1420FF9CD8FF56ABE8DEE1F8" attach="left" box="[937,961,1923,1937]" fontSize="6" pageId="5" pageNumber="20">4+5</subScript>
with 2–5 seta apically, other veins bare. Wing membrane bare. Squama with 31–35 setae, partly bi- to triserial.
</paragraph>
<caption id="DF1B9A931423FF9FDBB5527BEA93E5EC" ID-DOI="http://doi.org/10.5281/zenodo.7987251" ID-Zenodo-Dep="7987251" httpUri="https://zenodo.org/record/7987251/files/figure.png" pageId="6" pageNumber="21" startId="6.[227,299,851,873]" targetBox="[315,1273,227,834]" targetPageId="6" targetType="figure">
<paragraph id="8BDBCA1B1423FF9FDBB5527BEA93E5EC" blockId="6.[227,1360,851,901]" pageId="6" pageNumber="21">
Figures 8-9. 
<taxonomicName id="4C64B1981423FF9FDA34527BE915E501" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[354,522,851,872]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis id="B91016091423FF9FDA34527BE915E501" box="[354,522,851,872]" italics="true" pageId="6" pageNumber="21">Madachironomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721423FF9FD945527BE948E501" box="[531,599,851,872]" pageId="6" pageNumber="21" rank="genus">gen. n.</taxonomicNameLabel>
, wings. 8) 
<taxonomicName id="4C64B1981423FF9FD99E527BE85DE501" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[712,834,851,872]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="lakazana" status="sp. n.">
<emphasis id="B91016091423FF9FD99E527BE85DE501" box="[712,834,851,872]" italics="true" pageId="6" pageNumber="21">M. lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721423FF9FD81C527BE89EE501" box="[842,897,851,872]" pageId="6" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
, male; 9) 
<taxonomicName id="4C64B1981423FF9FD8BE527BEF63E501" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[1000,1148,851,872]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="rongaronga" status="sp. n.">
<emphasis id="B91016091423FF9FD8BE527BEF63E501" box="[1000,1148,851,872]" italics="true" pageId="6" pageNumber="21">M. rongaronga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721423FF9FDFD2527BEFA4E501" box="[1156,1211,851,872]" pageId="6" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
, female (photo Torbjørn Ekrem).
</paragraph>
</caption>
<caption id="DF1B9A931423FF9FDBB556E7EAD4EE6B" ID-DOI="http://doi.org/10.5281/zenodo.7987255" ID-Zenodo-Dep="7987255" httpUri="https://zenodo.org/record/7987255/files/figure.png" pageId="6" pageNumber="21" startId="6.[227,299,1999,2021]" targetBox="[327,1279,951,1974]" targetPageId="6" targetType="figure">
<paragraph id="8BDBCA1B1423FF9FDBB556E7EAD4EE6B" blockId="6.[227,1361,1999,2050]" pageId="6" pageNumber="21">
Figures 10-13. 
<taxonomicName id="4C64B1981423FF9FDA2E56E7E960E18D" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[376,639,1999,2020]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091423FF9FDA2E56E7E960E18D" box="[376,639,1999,2020]" italics="true" pageId="6" pageNumber="21">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721423FF9FD9D056E7E813E18D" box="[646,780,1999,2020]" pageId="6" pageNumber="21" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, female. 10) head; 11) tentorium, stipes and cibarial pump; 12) antenna; 13) thorax.
</paragraph>
</caption>
<caption id="DF1B9A931422FF9EDBB75443E871E3F7" ID-DOI="http://doi.org/10.5281/zenodo.7987257" ID-Zenodo-Dep="7987257" httpUri="https://zenodo.org/record/7987257/files/figure.png" pageId="7" pageNumber="22" startId="7.[225,297,1387,1409]" targetBox="[225,1361,251,1510]" targetPageId="7" targetType="figure">
<paragraph id="8BDBCA1B1422FF9EDBB75443E871E3F7" blockId="7.[225,1358,1387,1438]" pageId="7" pageNumber="22">
Figures 14-19. 
<taxonomicName id="4C64B1981422FF9EDA235443E964E3E9" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[373,635,1387,1408]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091422FF9EDA235443E964E3E9" box="[373,635,1387,1408]" italics="true" pageId="7" pageNumber="22">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721422FF9ED9D45444E818E3E8" box="[642,775,1388,1409]" pageId="7" pageNumber="22" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, female. 14) genitalia, ventral view; 15) tergite IX; 16) dor- somesal lobe; 17) ventrolateral lobe; 18) apodeme lobe; 19) labium.
</paragraph>
</caption>
<caption id="DF1B9A931422FF9EDBB354E0EE5FE08D" pageId="7" pageNumber="22" startId="7.[229,282,1480,1502]" targetType="table">
<paragraph id="8BDBCA1B1422FF9EDBB354E0EE0FE065" blockId="7.[229,1361,1480,1770]" pageId="7" pageNumber="22">
Table 2. Lengths (in µm) and proportions of legs of 
<taxonomicName id="4C64B1981422FF9ED85254E0EF12E3B4" authority="Andersen, 2016" authorityName="Andersen" authorityYear="2016" box="[772,1037,1480,1501]" class="Insecta" family="Chironomidae" genus="Madachironomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="lakazana" status="gen. n., sp. n.">
<emphasis id="B91016091422FF9ED85254E0EF12E3B4" box="[772,1037,1480,1501]" italics="true" pageId="7" pageNumber="22">Madachironomus lakazana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223AB721422FF9EDF4154E0EFB9E3B4" box="[1047,1190,1480,1501]" pageId="7" pageNumber="22" rank="species">gen. n., sp. n.</taxonomicNameLabel>
, female (n = 2). 
<emphasis id="B91016091422FF9EDA9754C6EE0FE065" bold="true" box="[449,1296,1518,1548]" pageId="7" pageNumber="22">fe ti ta1 ta2 ta3 ta4</emphasis>
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDA775739EE2FE040" blockId="7.[229,1361,1480,1770]" box="[289,1328,1553,1583]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDA775739EA28E046" bold="true" box="[289,311,1553,1583]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA795709EA28E046" attach="left" box="[303,311,1569,1583]" fontSize="6" pageId="7" pageNumber="22">1</subScript>
</emphasis>
1710–1751 2348–2410 1710–1812 700–721 618–639 494–515
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDA775706EE2FE02F" blockId="7.[229,1361,1480,1770]" box="[289,1328,1582,1612]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDA775706EA28E025" bold="true" box="[289,311,1582,1612]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA795716EA28E025" attach="left" box="[303,311,1598,1612]" fontSize="6" pageId="7" pageNumber="22">2</subScript>
</emphasis>
2245–2266 2554–2698 948–1009 515–556 453–474 288–330
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDA775764EE30E00D" blockId="7.[229,1361,1480,1770]" box="[289,1327,1612,1642]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDA775764EA28E003" bold="true" box="[289,311,1612,1642]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA795774EA28E003" attach="left" box="[303,311,1628,1642]" fontSize="6" pageId="7" pageNumber="22">3</subScript>
</emphasis>
2369–2390 2760–2946 1195–1257 597–700 556–618 350–371
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDAEB5746EACCE0EF" blockId="7.[229,1361,1480,1770]" box="[445,467,1646,1670]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDAEB5746EACCE0EF" bold="true" box="[445,467,1646,1670]" pageId="7" pageNumber="22">ta</emphasis>
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDA855757EAC4E0E4" blockId="7.[229,1361,1480,1770]" box="[467,475,1663,1677]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDA855757EAC4E0E4" bold="true" box="[467,475,1663,1677]" pageId="7" pageNumber="22">5</emphasis>
</paragraph>
<paragraph id="8BDBCA1B1422FF9ED9C75746EE0FE0EF" blockId="7.[229,1361,1480,1770]" box="[657,1296,1646,1670]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9ED9C75746EE0FE0EF" bold="true" box="[657,1296,1646,1670]" pageId="7" pageNumber="22">LR BV SV BR</emphasis>
</paragraph>
<paragraph id="8BDBCA1B1422FF9EDA4B57B9EE5FE08D" blockId="7.[229,1361,1480,1770]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDA4B57B9EA2CE0C6" bold="true" box="[285,307,1681,1711]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA7D5789EA2CE0C6" attach="left" box="[299,307,1697,1711]" fontSize="6" pageId="7" pageNumber="22">1</subScript>
</emphasis>
226–247 0.728–0.752 2.800–2.843 2.295–2.373 1.733–1.750 
<emphasis id="B91016091422FF9EDA4B5787EA2CE0A4" bold="true" box="[285,307,1711,1741]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA7D5797EA2CE0A4" attach="left" box="[299,307,1727,1741]" fontSize="6" pageId="7" pageNumber="22">2</subScript>
</emphasis>
144–165 0.371–0.374 3.905–4.118 4.898–5.087 1.750–1.888 
<emphasis id="B91016091422FF9EDA4A57E4EA2CE083" bold="true" box="[284,307,1740,1770]" pageId="7" pageNumber="22">
p 
<subScript id="17E0C85E1422FF9EDA7D57F4EA2CE083" attach="left" box="[299,307,1756,1770]" fontSize="6" pageId="7" pageNumber="22">3</subScript>
</emphasis>
144–165 0.427–0.433 3.556–3.837 4.293–4.246 1.600–1.700
</paragraph>
</caption>
<paragraph id="8BDBCA1B1422FF9EDBB55631EF6DE119" blockId="7.[227,764,1816,2000]" lastBlockId="7.[824,1361,1816,1904]" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9EDBB55631EA03E159" box="[227,284,1817,1840]" italics="true" pageId="7" pageNumber="22">Legs.</emphasis>
Spur of fore tibia 76–80 µm long, spurs of mid tibia 100–104 µm and 112–116 µm long, of hind tibia 100–108 µm and 116–120 µm long. Width at apex of fore tibia 116–124 µm, of mid tibia 128–136 µm, of hind tibia 136–144 µm. Mid ta 
<subScript id="17E0C85E1422FF9EDBA056E0EBE1E1BF" attach="left" box="[246,254,1992,2006]" fontSize="6" pageId="7" pageNumber="22">1</subScript>
with about 50 sensilla chaetica in 2–3 rows in proximal 1/3, hind ta 
<subScript id="17E0C85E1422FF9EDF435600EF02E15F" attach="left" box="[1045,1053,1832,1846]" fontSize="6" pageId="7" pageNumber="22">1</subScript>
with about 35 sensilla chaetica in 2–3 rows in proximal 1/3. Lengths and pro- portions of legs as in 
<tableCitation id="C6E6FFA01422FF9EDF4A5670EF71E119" box="[1052,1134,1880,1904]" captionStart="Table 2" captionStartId="7.[229,282,1480,1502]" captionText="Table 2. Lengths (in µm) and proportions of legs of Madachironomus lakazana gen. n., sp. n., female (n = 2). fe ti ta1 ta2 ta3 ta4 p1 1710–1751 2348–2410 1710–1812 700–721 618–639 494–515 p2 2245–2266 2554–2698 948–1009 515–556 453–474 288–330 p3 2369–2390 2760–2946 1195–1257 597–700 556–618 350–371 ta 5 LR BV SV BR p1 226–247 0.728–0.752 2.800–2.843 2.295–2.373 1.733–1.750 p2 144–165 0.371–0.374 3.905–4.118 4.898–5.087 1.750–1.888 p3 144–165 0.427–0.433 3.556–3.837 4.293–4.246 1.600–1.700" pageId="7" pageNumber="22">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BDBCA1B1422FF91D86E56AFEA6DE712" blockId="7.[824,1361,1927,2015]" lastBlockId="8.[227,764,227,380]" lastPageId="8" lastPageNumber="23" pageId="7" pageNumber="22">
<emphasis id="B91016091422FF9ED86E56AFE882E1F6" box="[824,925,1927,1951]" italics="true" pageId="7" pageNumber="22">Genitalia</emphasis>
(
<figureCitation id="135FD69E1422FF9ED8FA56AFEF36E1F6" box="[940,1065,1927,1951]" captionStart="Figures 14-19" captionStartId="7.[225,297,1387,1409]" captionTargetBox="[225,1361,251,1510]" captionTargetId="figure-1@7.[293,1294,174,1508]" captionTargetPageId="7" captionText="Figures 14-19. Madachironomus lakazana gen. n., sp. n., female. 14) genitalia, ventral view; 15) tergite IX; 16) dor- somesal lobe; 17) ventrolateral lobe; 18) apodeme lobe; 19) labium." figureDoi="http://doi.org/10.5281/zenodo.7987257" httpUri="https://zenodo.org/record/7987257/files/figure.png" pageId="7" pageNumber="22">Figs 14–19</figureCitation>
). Segment X with 189–231 setae to each side of vagina. Seminal capsule ovoid, 176–184 μm long, not including 18–20 μm long neck, 120–132 μm wide. Notum 204–228 μm long. Dorsomesal lobe 218–224 μm long from base of vagina to apex. Gonocoxite IX with 28–30 setae. Tergite IX with 31–37 setae. Cercus 284– 292 μm long.
</paragraph>
<paragraph id="8BDBCA1B142DFF91DBB550BAE906E7C3" blockId="8.[227,537,402,426]" box="[227,537,402,426]" pageId="8" pageNumber="23">
<emphasis id="B9101609142DFF91DBB550BAEABEE7C3" bold="true" box="[227,417,402,426]" pageId="8" pageNumber="23">Larva and pupa.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>