187 lines
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187 lines
18 KiB
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<mods:title id="544F7918ED9BDE3F0FC4F8BDCC9B0B6E">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="AE6157D70491CFE39403702AFC8CFEF5">Toda, Masanori J.</mods:namePart>
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<mods:affiliation id="8BA098DD7D9E935874B168914449D65E">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="17CAB97C974040A83FAC1093F174B21C">Bänziger, Hans</mods:namePart>
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<mods:affiliation id="19D86CEF2C49B724DF703C0AC3C4DBDD">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="0EBB888C6601B569A8CA6E919626A0E1">Sati, Pradeep C.</mods:namePart>
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<mods:affiliation id="36A0C91FBA4174C244231124FAF0792B">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="6C0E88B0DA1B126DBCCBB4CBE04D56F8">Fartyal, Rajendra S.</mods:namePart>
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<mods:affiliation id="A778284B813CC070C1E06F5C7387E781">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="A7C707E3EECE95C49A37A0F551CBF18E">Suwito, Awit</mods:namePart>
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<mods:affiliation id="E1078968C89022A95B0790C53FB5809A">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="0B8D668375343A7C54FED5641902587E">Katoh, Toru</mods:namePart>
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<mods:affiliation id="66CE3D39D01E64F1A74CD639E05120AC">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:title id="A27B08DE2B95579B39CF130630688787">Zootaxa</mods:title>
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<mods:part id="A3098861D61E7E87534429D91E6E7CAF">
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<mods:date id="FB22FC1EE2561B4CC20913809B801504">2020</mods:date>
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<mods:number id="5395F94A59FC88EB688E1E509045F6E5">2020-06-08</mods:number>
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<mods:number id="4FC08B986D9BF761D4B7C553792B5FF7">4789</mods:number>
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<mods:identifier id="141D1B573304FC82A60B15688768787D" type="DOI">10.11646/zootaxa.4789.1.1</mods:identifier>
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<treatment id="03C86C16EB39577AFF6EDBB6FCF7FEEE" ID-DOI="http://doi.org/10.5281/zenodo.5919016" ID-GBIF-Taxon="164612780" ID-Zenodo-Dep="5919016" LSID="urn:lsid:plazi:treatment:03C86C16EB39577AFF6EDBB6FCF7FEEE" httpUri="http://treatment.plazi.org/id/03C86C16EB39577AFF6EDBB6FCF7FEEE" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<subSubSection id="C37B8E8BEB395745FF6EDBB6FD1FF973" box="[151,756,1726,1753]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BDEDD00EB395745FF6EDBB6FD1FF973" blockId="31.[151,756,1726,1753]" box="[151,756,1726,1753]" pageId="31" pageNumber="32">
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<heading id="D0966A6CEB395745FF6EDBB6FD1FF973" bold="true" box="[151,756,1726,1753]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B9150112EB395745FF6EDBB6FD1FF973" bold="true" box="[151,756,1726,1753]" pageId="31" pageNumber="32">
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32)
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<taxonomicName id="4C61A683EB395745FF3FDBB6FD1FF973" ID-CoL="4GS4S" authority="Chen & Toda, 2005" authorityName="Chen & Toda" authorityYear="2005" box="[198,756,1726,1753]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="glabra" subGenus="Ashima">
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<emphasis id="B9150112EB395745FF3FDBB6FEC5F973" bold="true" box="[198,302,1726,1753]" italics="true" pageId="31" pageNumber="32">Phortica</emphasis>
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(
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<emphasis id="B9150112EB395745FEC6DBB6FE77F973" bold="true" box="[319,412,1726,1753]" italics="true" pageId="31" pageNumber="32">Ashima</emphasis>
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)
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<emphasis id="B9150112EB395745FE54DBB6FE10F973" bold="true" box="[429,507,1726,1753]" italics="true" pageId="31" pageNumber="32">glabra</emphasis>
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<bibRefCitation id="EFF0A0F1EB395745FDFADBB7FD1FF973" author="Chen & Toda" box="[515,756,1727,1753]" firstAuthor="Chen" pageId="31" pageNumber="32" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. & Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005">Chen & Toda, 2005</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B8E8BEB395745FF6EDA0BFD59F890" pageId="31" pageNumber="32" type="reference_group">
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<paragraph id="8BDEDD00EB395745FF6EDA0BFD36F8B1" blockId="31.[151,733,1794,1850]" box="[151,733,1794,1819]" pageId="31" pageNumber="32">
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<treatmentCitationGroup id="AB71FA2EEB395745FF6EDA0BFD36F8B1" box="[151,733,1794,1819]" pageId="31" pageNumber="32">
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<treatmentCitation id="0AC0FB11EB395745FF6EDA0BFD32F8B1" author="Chen, H. W. & Toda, M. J. & Gao, J. J." box="[151,729,1794,1819]" page="424" pageId="31" pageNumber="32" year="2005">
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<taxonomicName id="4C61A683EB395745FF6EDA0BFD32F8B1" ID-CoL="4GS4S" authority="Chen & Toda in Chen et al., 2005 b: 424" authorityName="Chen & Toda in Chen" authorityPageNumber="424" authorityYear="2005" box="[151,729,1794,1819]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="glabra">
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<emphasis id="B9150112EB395745FF6EDA0BFED1F8B1" box="[151,314,1795,1819]" italics="true" pageId="31" pageNumber="32">Phortica glabra</emphasis>
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Chen & Toda in
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<bibRefCitation id="EFF0A0F1EB395745FE11DA0AFD32F8B1" author="Chen" box="[488,729,1794,1819]" etAl="et al." firstAuthor="Chen" pageId="31" pageNumber="32" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. & Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005" yearSuffix="b">
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Chen
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<emphasis id="B9150112EB395745FDDDDA0CFDBAF8B1" box="[548,593,1795,1819]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
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., 2005b: 424
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</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BDEDD00EB395745FF6EDA2AFD59F890" blockId="31.[151,733,1794,1850]" box="[151,690,1825,1850]" pageId="31" pageNumber="32">
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<treatmentCitationGroup id="AB71FA2EEB395745FF6EDA2AFD59F890" box="[151,690,1825,1850]" pageId="31" pageNumber="32">
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<treatmentCitation id="0AC0FB11EB395745FF6EDA2AFD45F890" author="Chen, H. W. & Maca, J." box="[151,686,1825,1850]" page="507" pageId="31" pageNumber="32" year="2012">
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<taxonomicName id="4C61A683EB395745FF6EDA2AFD45F890" ID-CoL="4GS4S" authority=": Chen & Maca, 2012: 507" authorityName=": Chen & Maca" authorityPageNumber="507" authorityYear="2012" box="[151,686,1825,1850]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="glabra" subGenus="Ashima">
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<emphasis id="B9150112EB395745FF6EDA2AFF1BF890" box="[151,240,1826,1850]" italics="true" pageId="31" pageNumber="32">Phortica</emphasis>
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(
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<emphasis id="B9150112EB395745FF06DA2AFEA0F890" box="[255,331,1826,1850]" italics="true" pageId="31" pageNumber="32">Ashima</emphasis>
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)
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<emphasis id="B9150112EB395745FEA0DA2AFE76F890" box="[345,413,1826,1850]" italics="true" pageId="31" pageNumber="32">glabra</emphasis>
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:
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<bibRefCitation id="EFF0A0F1EB395745FE53DA29FD45F890" author="Chen & Maca" box="[426,686,1825,1850]" firstAuthor="Chen" pageId="31" pageNumber="32" pagination="493 - 509" refId="ref33941" refString="Chen, H. W. & Maca, J. (2012) Ten new species of the genus Phortica from the Afrotropical and Oriental regions (Diptera: Drosophilidae). Zootaxa, 3478 (1), 493 - 509. https: // doi. org / 10.11646 / zootaxa. 3478.1.42" type="journal article" year="2012">Chen & Máca, 2012: 507</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B8E8BEB39577AFF6EDA6FFF11FEB5" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8BDEDD00EB395745FF6EDA6FFCBEF860" blockId="31.[151,1437,1895,2031]" pageId="31" pageNumber="32">
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<emphasis id="B9150112EB395745FF6EDA6FFEF9F828" bold="true" box="[151,274,1895,1922]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
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Arista with 1 or 2 ventral branches (ch.6-1); posterior postgonite present only on right side of aedeagal sheath, short, apically blunt, and left lateral lobe long, broad, somewhat sinuated, apically with very large, trifurcated, anterior postgonite (“Figs 17, 18” in
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<bibRefCitation id="EFF0A0F1EB395745FD84DAB8FCA2F860" author="Chen" box="[637,841,1967,1995]" etAl="et al." firstAuthor="Chen" pageId="31" pageNumber="32" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. & Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005" yearSuffix="b">
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Chen
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<emphasis id="B9150112EB395745FD46DAB9FD1BF860" box="[703,752,1968,1994]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
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. 2005b
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BDEDD00EB39577AFF3EDADBFD07FF7D" blockId="31.[151,1437,1895,2031]" lastBlockId="32.[151,1437,153,324]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<emphasis id="B9150112EB395745FF3EDADBFDE1F844" bold="true" box="[199,522,2003,2030]" pageId="31" pageNumber="32">Supplementary description</emphasis>
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(not repeating characters common to
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<taxonomicName id="4C61A683EB395745FC52DADDFBB7F844" authorityName="Gong, Zhu & Chen" authorityYear="2018" box="[939,1116,2004,2031]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zhangyuanae">
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<emphasis id="B9150112EB395745FC52DADDFC57F844" box="[939,956,2005,2030]" italics="true" pageId="31" pageNumber="32">P</emphasis>
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.
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<emphasis id="B9150112EB395745FC31DADCFBB7F844" box="[968,1116,2004,2030]" italics="true" pageId="31" pageNumber="32">zhangyuanae</emphasis>
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</taxonomicName>
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). Supracervical setae 20–22. Apical, leaf-like expansion of arista apically pointed or with minute, awn-like projection. Cibarial, medial sensilla 7–8 per side. All tarsi with gray tarsomeres IV and V.
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</paragraph>
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<paragraph id="8BDEDD00EB06577AFF3EDDE8FF11FEB5" blockId="32.[151,1437,153,324]" pageId="32" pageNumber="33">
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|
<emphasis id="B9150112EB06577AFF3EDDE8FE25FF51" bold="true" box="[199,462,224,251]" pageId="32" pageNumber="33">Specimens examined.</emphasis>
|
|||
|
<materialsCitation id="3B09D75DEB06577AFE25DDE9FF1DFEB5" ID-GBIF-Occurrence="2636268388" collectingDate="2004-05-02" collectionCode="SEHU" collectorName="H. - W. Chen" country="China" location="Guangdong" pageId="32" pageNumber="33" specimenCount="5" specimenCount-male="5" stateProvince="Guangdong" typeStatus="paratype">
|
|||
|
<collectingCountry id="F3769D90EB06577AFE25DDE9FDC8FF51" box="[476,547,225,251]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
|||
|
:
|
|||
|
<specimenCount id="9D671689EB06577AFDCCDDE8FDB3FF51" box="[565,600,224,251]" pageId="32" pageNumber="33" type="male">5♂</specimenCount>
|
|||
|
<typeStatus id="54DA63A2EB06577AFD9EDDE9FD3EFF51" box="[615,725,225,251]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
|
|||
|
, Nanlling, Luyuan,
|
|||
|
<collectingRegion id="49A513E2EB06577AFC33DDE9FBBBFF51" box="[970,1104,225,251]" country="China" name="Guangdong" pageId="32" pageNumber="33">Guangdong</collectingRegion>
|
|||
|
,
|
|||
|
<date id="FFDFFBC0EB06577AFB9BDDE9FB2FFF51" box="[1122,1220,225,251]" pageId="32" pageNumber="33" value="2004-05-02">
|
|||
|
<collectingDate id="EF9B0228EB06577AFB9BDDE9FB2FFF51" box="[1122,1220,225,251]" pageId="32" pageNumber="33" value="2004-05-02">2.v.2004</collectingDate>
|
|||
|
</date>
|
|||
|
,
|
|||
|
<collectorName id="2694B8D6EB06577AFB21DDE9FA8FFF51" box="[1240,1380,225,251]" pageId="32" pageNumber="33">H.-W. Chen</collectorName>
|
|||
|
leg. (
|
|||
|
<collectionCode id="ED7045C5EB06577AFF66DC0DFF05FEB5" box="[159,238,261,287]" pageId="32" pageNumber="33">SEHU</collectionCode>
|
|||
|
)
|
|||
|
</materialsCitation>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C37B8E8BEB06577AFF3EDC20FCF7FEEE" box="[199,796,296,324]" pageId="32" pageNumber="33" type="distribution">
|
|||
|
<paragraph id="8BDEDD00EB06577AFF3EDC20FCF7FEEE" blockId="32.[151,1437,153,324]" box="[199,796,296,324]" pageId="32" pageNumber="33">
|
|||
|
<emphasis id="B9150112EB06577AFF3EDC20FE8AFEE9" bold="true" box="[199,353,296,323]" pageId="32" pageNumber="33">Distribution.</emphasis>
|
|||
|
<collectingCountry id="F3769D90EB06577AFE9EDC21FE41FEE9" box="[359,426,297,323]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
|||
|
(
|
|||
|
<collectingRegion id="49A513E2EB06577AFE40DC21FDE5FEE9" box="[441,526,297,323]" country="China" name="Jiangxi" pageId="32" pageNumber="33">Jiangxi</collectingRegion>
|
|||
|
,
|
|||
|
<collectingRegion id="49A513E2EB06577AFDE3DC21FD4BFEE9" box="[538,672,297,323]" country="China" name="Guangdong" pageId="32" pageNumber="33">Guangdong</collectingRegion>
|
|||
|
,
|
|||
|
<collectingRegion id="49A513E2EB06577AFD53DC21FCE4FEEE" box="[682,783,297,324]" country="China" name="Guangxi" pageId="32" pageNumber="33">Guangxi</collectingRegion>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|