plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/A3/09/03A3096E3A11FFFAF462FC164D33C18B.xml

457 lines
51 KiB
XML
Raw Normal View History

added 03
2024-06-21 12:22:17 +02:00
<document id="BC46068B270B43BE74E43905BE29E71F" ID-CLB-Dataset="5496" ID-DOI="10.11646/zootaxa.296.1.1" ID-GBIF-Dataset="3ddea826-544c-452c-994e-01c1a4973a4b" ID-ISSN="1175­5334" ID-Zenodo-Dep="5014425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624382618413" checkinUser="felipe" docAuthor="Shelley, Rowland M." docDate="2003" docId="03A3096E3A11FFFAF462FC164D33C18B" docLanguage="en" docName="zt00296.pdf" docOrigin="Zootaxa 296 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.296.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Retrorsia leonardi Shelley 2003, new species" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FF9A71163A15FFF2F56AFFFA4C5BC255" masterDocTitle="A new polydesmid milliped genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett and Gardner, 1968, and a summary of the family in Western North America (Polydesmida: Polydesmidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="5" updateTime="1699020097571" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="174F4AB3796EB9EEE8DE93B3FC094ECA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A5FD4D9F2F97BEA71EA3F7CC18A4AE84">
<mods:title id="D33ADC61A30DAEB240C822DF835DEF82">A new polydesmid milliped genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett and Gardner, 1968, and a summary of the family in Western North America (Polydesmida: Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="87674B912C5C6EC92254C5E7331CD397" type="personal">
<mods:role id="4BF9E80188AC0A0B5D8E5A743D19FC33">
<mods:roleTerm id="70C3EE052C4B274BE5414CD6F8113205">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="434B72D5147B81A9935CDCF9C9F54C84">Shelley, Rowland M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="67A5D9B5DC2108EDF323E9C7C336D53B">text</mods:typeOfResource>
<mods:relatedItem id="09AC24FFF0308E8F604DB546983086AB" type="host">
<mods:titleInfo id="EC98B145C1EBEC6B97C191B85E7A1309">
<mods:title id="563AAF04BE53627E01E1527DB0FA3339">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7278FDC2A3608136517B4D828319DACD">
<mods:date id="12BFE17A84DF562D8DE3B468F87291E3">2003</mods:date>
<mods:detail id="9C7F69BB740CB96241C90ECA0BC96223" type="pubDate">
<mods:number id="90FE47D84640893B53C4A939AD3E7851">2003-09-16</mods:number>
</mods:detail>
<mods:detail id="5803400968555BE400B4DED644F59812" type="volume">
<mods:number id="DE2A576A0594ACEA8B6A3DB034B0C3F3">296</mods:number>
</mods:detail>
<mods:detail id="26EA5F2D0A49D60B783CA83F081207C4" type="issue">
<mods:number id="29948369B35232A29674C0A4808427D8">1</mods:number>
</mods:detail>
<mods:extent id="2FD612616D4EA540ACB91C908F0013C0" unit="page">
<mods:start id="7391C736BF38572F58F1FAEF9B2C5F8A">1</mods:start>
<mods:end id="E8FF8144DD98F20E972976C22C8AB2A6">12</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="F98A77BC4C93C3C66E8B7CBF27B70FED">
<mods:url id="72C1A72AA881356976751BA256EE1673">https://biotaxa.org/Zootaxa/article/view/zootaxa.296.1.1</mods:url>
</mods:location>
<mods:classification id="144E6CF545DDBDD74128214DE24ADF22">journal article</mods:classification>
<mods:identifier id="F6E60316DAA901383CAF5918B184E743" type="CLB-Dataset">5496</mods:identifier>
<mods:identifier id="7D89D48CCCA51E9445FA29F66E278C70" type="DOI">10.11646/zootaxa.296.1.1</mods:identifier>
<mods:identifier id="A197714E10ED452AFD3BC54F026FFA9E" type="GBIF-Dataset">3ddea826-544c-452c-994e-01c1a4973a4b</mods:identifier>
<mods:identifier id="317C3EB2C2D88B49E60A91B4FE521130" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="0910DFA2B93AA8C880EFFEC6416AB044" type="Zenodo-Dep">5014425</mods:identifier>
<mods:identifier id="61DFB6076A57A76DD7327A04A5C013B8" type="ZooBank">1C2F5E8B-8D60-49F1-B6CE-E13FFDCD67D3</mods:identifier>
</mods:mods>
<treatment id="03A3096E3A11FFFAF462FC164D33C18B" ID-DOI="http://doi.org/10.5281/zenodo.5095716" ID-GBIF-Taxon="183531920" ID-Zenodo-Dep="5095716" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B" httpUri="http://treatment.plazi.org/id/03A3096E3A11FFFAF462FC164D33C18B" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<subSubSection id="C310EBF33A11FFF6F462FC164ED8C653" box="[264,643,1004,1030]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BB5B8783A11FFF6F462FC164ED8C653" blockId="4.[264,643,1004,1070]" box="[264,643,1004,1030]" pageId="4" pageNumber="5">
<heading id="D0FD0F143A11FFF6F462FC164ED8C653" bold="true" box="[264,643,1004,1030]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C0AC3FB3A11FFF6F462FC164DBEC653" ID-CoL="78RVF" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[264,485,1004,1030]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="leonardi" status="sp. nov.">
<emphasis id="B97E646A3A11FFF6F462FC164DBEC653" bold="true" box="[264,485,1004,1030]" italics="true" pageId="4" pageNumber="5">Retrorsia leonardi</emphasis>
</taxonomicName>
,
<emphasis id="B97E646A3A11FFF6F499FC164ED8C653" bold="true" box="[499,643,1004,1030]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A24DD9113A11FFF6F499FC164ED8C653" box="[499,643,1004,1030]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310EBF33A11FFFAF462FBEE4D33C18B" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB5B8783A11FFF6F462FBEE4D2FC67B" blockId="4.[264,643,1004,1070]" box="[264,372,1044,1070]" pageId="4" pageNumber="5">
<figureCitation id="1331A4FD3A11FFF6F462FBEE4D2FC67B" box="[264,372,1044,1070]" captionStart="FIGURES 3–4" captionStartId="5.[264,382,1352,1376]" captionTargetBox="[276,1313,637,1342]" captionTargetId="figure-270@5.[273,1313,633,1348]" captionTargetPageId="5" captionText="FIGURES 3–4. Retrorsia leonardi. 3, right gonopod of holotype, medial view (drawn at 200x magnification). 4, telopodite of the same, lateral view (drawn at 400x magnification). C, coxa; E, endomerite; P, pulvillus; PF, prefemur; T, telopodite; TT, tibiotarsus." figureDoi="http://doi.org/10.5281/zenodo.5014435" httpUri="https://zenodo.org/record/5014435/files/figure.png" pageId="4" pageNumber="5">Figs. 3–4</figureCitation>
</paragraph>
<paragraph id="8BB5B8783A11FFF6F462FBB64F61C703" blockId="4.[264,1324,1100,1886]" pageId="4" pageNumber="5">
<emphasis id="B97E646A3A11FFF6F462FBB64DE5C630" box="[264,446,1100,1125]" italics="true" pageId="4" pageNumber="5">Type specimens</emphasis>
.
<materialsCitation id="3B62B2253A11FFF6F4A6FBB64F81C6E3" ID-GBIF-Occurrence="3321340308" collectingDate="2002-12-14" collectionCode="NCSM" collectorName="Male &amp; W. P. Leonard" county="Multnomah County" location="Columbia River Gorge National Scenic Area" municipality="Ainsworth State Park" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<collectorName id="26FFDDAE3A11FFF6F4A6FBB64E53C633" box="[460,520,1100,1126]" pageId="4" pageNumber="5">Male</collectorName>
<typeStatus id="54B106DA3A11FFF6F765FBB64E2DC633" box="[527,630,1100,1126]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED1B20BD3A11FFF6F7EFFBB64E87C633" box="[645,732,1100,1126]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="5" type="Museum">NCSM</collectionCode>
) collected by
<collectorName id="26FFDDAE3A11FFF6F614FBB6487AC633" box="[894,1057,1100,1126]" pageId="4" pageNumber="5">W. P. Leonard</collectorName>
on
<date id="FFB49EB83A11FFF6F127FBB64971C633" box="[1101,1322,1100,1126]" pageId="4" pageNumber="5" value="2002-12-14">
<collectingDate id="EFF067503A11FFF6F127FBB64971C633" box="[1101,1322,1100,1126]" pageId="4" pageNumber="5" value="2002-12-14">14 December 2002</collectingDate>
</date>
at
<collectingMunicipality id="6BD122023A11FFF6F44FFB8E4E7BC6DB" box="[293,544,1140,1166]" pageId="4" pageNumber="5">Ainsworth State Park</collectingMunicipality>
in the
<location id="8ED5EEA33A11FFF6F71BFB8E48D8C6DB" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A11FFF6F71BFB8E48D8C6DB" box="[625,1155,1140,1166]" county="Multnomah County" municipality="Ainsworth State Park" name="Columbia River Gorge National Scenic Area" pageId="4" pageNumber="5" stateProvince="Oregon">Columbia River Gorge National Scenic Area</location>
, ca.
<quantity id="4CF2159D3A11FFF6F1D2FB8E48A7C6DB" box="[1208,1276,1140,1166]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="4" pageNumber="5" unit="mi" value="30.0">30 mi</quantity>
(
<quantity id="4CF2159D3A11FFF6F066FB8E4D74C6E3" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="5" unit="km" value="48.0">48 km</quantity>
) E of downtown
<location id="8ED5EEA33A11FFF6F77AFB664E2DC6E3" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A11FFF6F77AFB664E2DC6E3" box="[528,630,1180,1206]" county="Multnomah County" municipality="Ainsworth State Park" name="Portland" pageId="4" pageNumber="5" stateProvince="Oregon">Portland</location>
,
<collectingCounty id="62D4C0F43A11FFF6F7EFFB664F2BC6E3" box="[645,880,1180,1206]" pageId="4" pageNumber="5">Multnomah County</collectingCounty>
,
<collectingRegion id="49CE769A3A11FFF6F614FB664F81C6E3" box="[894,986,1180,1206]" country="United States of America" name="Oregon" pageId="4" pageNumber="5">Oregon</collectingRegion>
</materialsCitation>
.
<materialsCitation id="3B62B2253A11FFF6F698FB664F18C753" ID-GBIF-Occurrence="3321340304" collectingDate="1968-01-21" collectionCode="CAS" collectorName="Male &amp; D. R. Malcolm" location="Helvetia" municipality="Bishop Road" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
<collectorName id="26FFDDAE3A11FFF6F698FB664875C6E3" box="[1010,1070,1180,1206]" pageId="4" pageNumber="5">Male</collectorName>
<typeStatus id="54B106DA3A11FFF6F152FB6648C6C6E3" box="[1080,1181,1180,1206]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED1B20BD3A11FFF6F1DAFB6648B7C6E3" box="[1200,1260,1180,1206]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="5" type="Museum">CAS</collectionCode>
) collected by
<collectorName id="26FFDDAE3A11FFF6F413FB3E4E73C68B" box="[377,552,1220,1246]" pageId="4" pageNumber="5">D. R. Malcolm</collectorName>
on
<date id="FFB49EB83A11FFF6F738FB3E4F49C68B" box="[594,786,1220,1246]" pageId="4" pageNumber="5" value="1968-01-21">
<collectingDate id="EFF067503A11FFF6F738FB3E4F49C68B" box="[594,786,1220,1246]" pageId="4" pageNumber="5" value="1968-01-21">21 January 1968</collectingDate>
</date>
along
<collectingMunicipality id="6BD122023A11FFF6F60BFB3E4FA7C68B" box="[865,1020,1220,1246]" pageId="4" pageNumber="5">Bishop Road</collectingMunicipality>
,
<quantity id="4CF2159D3A11FFF6F16DFB3E4860C68B" box="[1031,1083,1220,1246]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="4" pageNumber="5" unit="mi" value="2.0">2 mi</quantity>
(3.2 km) N
<location id="8ED5EEA33A11FFF6F1A3FB3E4977C68B" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A11FFF6F1A3FB3E4977C68B" box="[1225,1324,1220,1246]" municipality="Bishop Road" name="Helvetia" pageId="4" pageNumber="5" stateProvince="Washington">Helvetia</location>
[ca.
<quantity id="4CF2159D3A11FFF6F456FB164D29C753" box="[316,370,1260,1286]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="4" pageNumber="5" unit="mi" value="8.0">8 mi</quantity>
{12.8 km} NW Portland],
<collectingRegion id="49CE769A3A11FFF6F7D2FB164F18C753" box="[696,835,1260,1286]" country="United States of America" name="Washington" pageId="4" pageNumber="5">Washington</collectingRegion>
</materialsCitation>
<materialsCitation id="3B62B2253A11FFF6F626FB164857C753" ID-GBIF-Occurrence="3321340301" box="[844,1036,1260,1286]" collectorName="County" location="Oregon" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Oregon" typeStatus="paratype">
<collectorName id="26FFDDAE3A11FFF6F626FB164FF8C753" box="[844,931,1260,1286]" pageId="4" pageNumber="5">County</collectorName>
,
<collectingRegion id="49CE769A3A11FFF6F6DAFB164857C753" box="[944,1036,1260,1286]" country="United States of America" name="Oregon" pageId="4" pageNumber="5">Oregon</collectingRegion>
</materialsCitation>
.
<materialsCitation id="3B62B2253A11FFF6F14AFB164F6DC703" ID-GBIF-Occurrence="3321340309" collectingDate="1972-01-08" collectionCode="CAS, NCSM, VMNH" collectorName="Six &amp; E. Benedict" county="Columbia County" location="Clatskanie" pageId="4" pageNumber="5" specimenCount="4" specimenCount-female="4" stateProvince="Oregon" typeStatus="paratype">
<collectorName id="26FFDDAE3A11FFF6F14AFB16481CC753" box="[1056,1095,1260,1286]" pageId="4" pageNumber="5">Six</collectorName>
male and
<specimenCount id="9D0C73F13A11FFF6F1AFFB164977C753" box="[1221,1324,1260,1286]" pageId="4" pageNumber="5" type="female">4 female</specimenCount>
<typeStatus id="54B106DA3A11FFF6F462FAEE4D23C77B" box="[264,376,1300,1326]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED1B20BD3A11FFF6F4E0FAEE4D9CC77B" box="[394,455,1300,1326]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="5" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1B20BD3A11FFF6F4BFFAEE4E70C77B" box="[469,555,1300,1326]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="4" pageNumber="5" type="Museum">NCSM</collectionCode>
,
<collectionCode id="ED1B20BD3A11FFF6F752FAEE4ECFC77B" box="[568,660,1300,1326]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="4" pageNumber="5">VMNH</collectionCode>
) collected by
<collectorName id="26FFDDAE3A11FFF6F654FAEE4F93C77B" box="[830,968,1300,1326]" pageId="4" pageNumber="5">E. Benedict</collectorName>
on
<date id="FFB49EB83A11FFF6F69DFAEE48EBC77B" box="[1015,1200,1300,1326]" pageId="4" pageNumber="5" value="1972-01-08">
<collectingDate id="EFF067503A11FFF6F69DFAEE48EBC77B" box="[1015,1200,1300,1326]" pageId="4" pageNumber="5" value="1972-01-08">
8 January
<quantity id="4CF2159D3A11FFF6F119FAEE48EBC77B" box="[1139,1200,1300,1326]" metricMagnitude="6" metricUnit="m" metricValue="3.1741091712000005" pageId="4" pageNumber="5" unit="mi" value="1972.3">1972</quantity>
</collectingDate>
</date>
, 3 mi (4.8 km) SW
<location id="8ED5EEA33A11FFF6F418FAC64DAAC703" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A11FFF6F418FAC64DAAC703" box="[370,497,1340,1366]" county="Columbia County" name="Clatskanie" pageId="4" pageNumber="5" stateProvince="Oregon">Clatskanie</location>
,
<collectingCounty id="62D4C0F43A11FFF6F496FAC64E95C703" box="[508,718,1340,1366]" pageId="4" pageNumber="5">Columbia County</collectingCounty>
,
<collectingRegion id="49CE769A3A11FFF6F7B0FAC64F6DC703" box="[730,822,1340,1366]" country="United States of America" name="Oregon" pageId="4" pageNumber="5">Oregon</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB5B8783A11FFF6F452FA9E4F59C7A3" blockId="4.[264,1324,1100,1886]" pageId="4" pageNumber="5">
<emphasis id="B97E646A3A11FFF6F452FA9E4DF4C728" box="[312,431,1380,1405]" italics="true" pageId="4" pageNumber="5">Diagnosis</emphasis>
. Stem of gonopodal telopodite long, slender, and curvilinear, widening slightly distad at level of endomerite but not produced into a discrete flange; pulvillus rounded; tibiotarsus bent strongly dorsad, apically broad; endomerite short and subtriangular, oriented transversely to telopodal stem.
</paragraph>
<paragraph id="8BB5B8783A11FFF6F452F9FE4FEDC4EB" blockId="4.[264,1324,1100,1886]" pageId="4" pageNumber="5">
<emphasis id="B97E646A3A11FFF6F452F9FE4DF0C448" box="[312,427,1540,1565]" italics="true" pageId="4" pageNumber="5">
<typeStatus id="54B106DA3A11FFF6F452F9FE4DFCC448" box="[312,423,1540,1565]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
.
</emphasis>
Head densely covered with short, fine, parallel­sided setae; epicranial suture distinct; genae not extending beyond adjacent cranial margins. Paranota broad with strongly serrate, notched margins on segments 1–6, becoming slightly narrower on segment 7 and progressively less dentate caudad. Epiproct short and broad, with two tufts of long, terminal setae. Paraprocts with slight marginal rims.
</paragraph>
<paragraph id="8BB5B8783A11FFF7F452F9364D6AC01B" blockId="4.[264,1324,1100,1886]" lastBlockId="5.[264,1323,284,590]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Gonopodal aperture obchordate, caudal margin extending backwards between 9th legs, sides flush with metazonal surface. Gonopods
<emphasis id="B97E646A3A11FFF6F6F8F90E4F84C558" box="[914,991,1780,1805]" italics="true" pageId="4" pageNumber="5">in situ</emphasis>
with telopodites extending anteriad in subparallel arrangement over anterior margin of aperture and sternum of segment 6. Gonopod structure as follows (
<figureCitation id="1331A4FD3A11FFF6F7BDF8BE4F1CC50B" box="[727,839,1860,1886]" captionStart="FIGURES 3–4" captionStartId="5.[264,382,1352,1376]" captionTargetBox="[276,1313,637,1342]" captionTargetId="figure-270@5.[273,1313,633,1348]" captionTargetPageId="5" captionText="FIGURES 3–4. Retrorsia leonardi. 3, right gonopod of holotype, medial view (drawn at 200x magnification). 4, telopodite of the same, lateral view (drawn at 400x magnification). C, coxa; E, endomerite; P, pulvillus; PF, prefemur; T, telopodite; TT, tibiotarsus." figureDoi="http://doi.org/10.5281/zenodo.5014435" httpUri="https://zenodo.org/record/5014435/files/figure.png" pageId="4" pageNumber="5">Figs. 3–4</figureCitation>
): Coxa large, expanding into ventral lobe extending for 1/4 of length of telopodite. Prefemur expanding distad into short, rounded lobe. Telopodite relatively narrow, curving gently ventrad for most of length, distal expansion short and narrow, pulvillus small and rounded, widely separated from expansion, hairs continuing distad along ridge on medial margin; tibiotarsus curving strongly dorsad, directed caudad, folded distad, apically broad; endomerite short and subtriangular, oriented transversely across stem, expanding into two narrowly rounded lobes. Prostatic groove apparently without loop or chamber, opening on medial surface proximal to pulvillus.
</paragraph>
<caption id="DF75E8F03A10FFF7F462FAB24FBAC7F9" ID-DOI="http://doi.org/10.5281/zenodo.5014435" ID-Zenodo-Dep="5014435" httpUri="https://zenodo.org/record/5014435/files/figure.png" pageId="5" pageNumber="6" startId="5.[264,382,1352,1376]" targetBox="[276,1313,637,1342]" targetPageId="5">
<paragraph id="8BB5B8783A10FFF7F462FAB24FBAC7F9" blockId="5.[264,1323,1352,1452]" pageId="5" pageNumber="6">
<emphasis id="B97E646A3A10FFF7F462FAB24DEBC735" bold="true" box="[264,432,1352,1376]" pageId="5" pageNumber="6">FIGURES 3–4</emphasis>
.
<taxonomicName id="4C0AC3FB3A10FFF7F4A3FAB34ED4C735" authorityName="Shelley" authorityYear="2003" box="[457,655,1353,1376]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B97E646A3A10FFF7F4A3FAB34ED4C735" box="[457,655,1353,1376]" italics="true" pageId="5" pageNumber="6">Retrorsia leonardi</emphasis>
</taxonomicName>
. 3, right gonopod of holotype, medial view (drawn at 200x magnification). 4, telopodite of the same, lateral view (drawn at 400x magnification). C, coxa; E, endomerite; P, pulvillus; PF, prefemur; T, telopodite; TT, tibiotarsus.
</paragraph>
</caption>
<paragraph id="8BB5B8783A10FFF7F452FA1E484FC7A8" blockId="5.[264,1323,1507,1893]" box="[312,1044,1507,1533]" pageId="5" pageNumber="6">
<emphasis id="B97E646A3A10FFF7F452FA1E4DB5C7A8" box="[312,494,1508,1533]" italics="true" pageId="5" pageNumber="6">
Male
<typeStatus id="54B106DA3A10FFF7F411FA1E4DB5C7A8" box="[379,494,1508,1533]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
</emphasis>
. Agreeing closely with
<typeStatus id="54B106DA3A10FFF7F665FA194F2EC7A8" box="[783,885,1507,1533]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
in all details.
</paragraph>
<paragraph id="8BB5B8783A10FFF7F452F9F64EABC490" blockId="5.[264,1323,1507,1893]" pageId="5" pageNumber="6">
<emphasis id="B97E646A3A10FFF7F452F9F64DCDC470" box="[312,406,1548,1573]" italics="true" pageId="5" pageNumber="6">Ecology</emphasis>
.
<materialsCitation id="3B62B2253A10FFF7F4C4F9F14EB7C490" ID-GBIF-Occurrence="3321340315" county="County" location="Pacific" municipality="Their" pageId="5" pageNumber="6" specimenCount="2" stateProvince="Washington" typeStatus="holotype">
The
<typeStatus id="54B106DA3A10FFF7F489F9F14E12C470" box="[483,585,1547,1573]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
was collected on a north­facing slope, probably in sword fern litter (
<taxonomicName id="4C0AC3FB3A10FFF7F43BF9CE4E0AC418" authorityName="C. Presl" authorityYear="1836" baseAuthorityName="Kaulfuss" box="[337,593,1588,1613]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="munitum">
<emphasis id="B97E646A3A10FFF7F43BF9CE4E0AC418" box="[337,593,1588,1613]" italics="true" pageId="5" pageNumber="6">Polystichum munitum</emphasis>
</taxonomicName>
); the
<typeStatus id="54B106DA3A10FFF7F7FFF9C94EA1C418" box="[661,762,1587,1613]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
from
<collectingRegion id="49CE769A3A10FFF7F629F9C94F95C418" box="[835,974,1587,1613]" country="United States of America" name="Washington" pageId="5" pageNumber="6">Washington</collectingRegion>
<collectingCounty id="62D4C0F43A10FFF7F6BCF9C94877C418" box="[982,1068,1587,1613]" pageId="5" pageNumber="6">County</collectingCounty>
was extracted from a berlesate of mixed conifer and deciduous duff.
<collectingMunicipality id="6BD122023A10FFF7F65FF9A14F2EC420" box="[821,885,1627,1653]" pageId="5" pageNumber="6">Their</collectingMunicipality>
discoveries in winter along with several new chordeumatidans (
<bibRefCitation id="EF9BC5893A10FFF7F723F9794F05C4C8" author="Shear, W. A. &amp; Leonard, W. P." box="[585,862,1667,1693]" pageId="5" pageNumber="6" pagination="1 - 11" refId="ref5530" refString="Shear, W. A. &amp; Leonard, W. P. (2003) Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa, 243, 1 - 11." type="journal article" year="2003">Shear &amp; Leonard, 2003</bibRefCitation>
) testifies to the need for collecting during cool weather in the
<location id="8ED5EEA33A10FFF7F776F9514E36C490" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A10FFF7F776F9514E36C490" box="[540,621,1707,1733]" county="County" municipality="Their" name="Pacific" pageId="5" pageNumber="6" stateProvince="Washington">Pacific</location>
northwest
</materialsCitation>
.
</paragraph>
<paragraph id="8BB5B8783A10FFF7F452F92E4F55C568" blockId="5.[264,1323,1507,1893]" pageId="5" pageNumber="6">
<emphasis id="B97E646A3A10FFF7F452F92E4D9CC4B8" box="[312,455,1748,1773]" italics="true" pageId="5" pageNumber="6">Distribution</emphasis>
. Occurring along a
<quantity id="4CF2159D3A10FFF7F7D7F9294F59C4B8" box="[701,770,1747,1773]" metricMagnitude="5" metricUnit="m" metricValue="1.1265408" pageId="5" pageNumber="6" unit="mi" value="70.0">70 mi</quantity>
(
<quantity id="4CF2159D3A10FFF7F679F9294F34C4B8" box="[787,879,1747,1773]" metricMagnitude="5" metricUnit="m" metricValue="1.1199999999999999" pageId="5" pageNumber="6" unit="km" value="112.0">112 km</quantity>
) stretch in the lower Columbia River Valley,
<collectingRegion id="49CE769A3A10FFF7F40FF9014D9BC540" box="[357,448,1787,1813]" country="United States of America" name="Oregon" pageId="5" pageNumber="6">Oregon</collectingRegion>
, from the Gorge in Multnomah County westward to Columbia County some
<quantity id="4CF2159D3A10FFF7F427F8D94DCAC568" box="[333,401,1827,1853]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="5" pageNumber="6" unit="mi" value="30.0">30 mi</quantity>
(
<quantity id="4CF2159D3A10FFF7F4CAF8D94DB5C568" box="[416,494,1827,1853]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="6" unit="km" value="48.0">48 km</quantity>
) from the river’s mouth.
</paragraph>
<paragraph id="8BB5B8783A10FFF4F452F8B64DF7C30B" blockId="5.[264,1323,1507,1893]" lastBlockId="6.[264,1323,284,470]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B97E646A3A10FFF7F452F8B64DE8C530" box="[312,435,1868,1893]" italics="true" pageId="5" pageNumber="6">Etymology</emphasis>
. I am pleased to name this species for the collector, William P. Leonard, in recognition of his discoveries of new milliped species and genera in the northwestern
<collectingCountry id="F31DF8E83A13FFF4F462FEBE4DF3C30B" box="[264,424,324,350]" name="United States of America" pageId="6" pageNumber="7">United States</collectingCountry>
.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F452FE964F92C383" blockId="6.[264,1323,284,470]" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F452FE964DC5C3D0" box="[312,414,364,389]" italics="true" pageId="6" pageNumber="7">Remarks</emphasis>
. The gonopod is so minute that even after clearing in lactic acid for a few days and viewing at 400x under a compound microscope, I could not discern whether or not there is a distal loop or chamber on the prostatic groove.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F462FDD34EC7C016" blockId="6.[264,668,553,619]" box="[264,668,553,579]" pageId="6" pageNumber="7">
<taxonomicName id="4C0AC3FB3A13FFF4F462FDD34DA4C016" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[264,511,553,579]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="benedictae" status="sp. nov.">
<emphasis id="B97E646A3A13FFF4F462FDD34DA4C016" bold="true" box="[264,511,553,579]" italics="true" pageId="6" pageNumber="7">Retrorsia benedictae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24DD9113A13FFF4F765FDD34EC7C016" box="[527,668,553,579]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB5B8783A13FFF4F462FDAB4D2BC03E" blockId="6.[264,668,553,619]" box="[264,368,593,619]" pageId="6" pageNumber="7">
<figureCitation id="1331A4FD3A13FFF4F462FDAB4D2BC03E" box="[264,368,593,619]" captionStart="FIGURES 5–8" captionStartId="7.[264,382,1747,1771]" captionTargetBox="[478,1109,394,1719]" captionTargetId="figure-86@7.[454,1116,391,1097]" captionTargetPageId="7" captionText="FIGURES 5–8. Retrorsia benedictae (all drawn at 400x magnification) 5, telopodite of left gonopod of holotype, medial view. 6, the same, lateral view. 7, telopodite of left gonopod of male from Pacific County, Washington, medial view. 8, the same, lateral view. F, tibiotarsal flange; other abbreviations as in figs. 3–4." figureDoi="http://doi.org/10.5281/zenodo.5014437" httpUri="https://zenodo.org/record/5014437/files/figure.png" pageId="6" pageNumber="7">Figs. 5­8</figureCitation>
</paragraph>
<paragraph id="8BB5B8783A13FFF4F462FD764885C14B" blockId="6.[264,1324,652,1878]" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F462FD764DE4C0F0" box="[264,447,652,677]" italics="true" pageId="6" pageNumber="7">Type specimens</emphasis>
.
<materialsCitation id="3B62B2253A13FFF4F4BCFD764881C14B" ID-GBIF-Occurrence="3321340318" collectingDate="1971-11-27" collectionCode="CAS, NCSM" collectorName="E. M. Benedict &amp; SE Olney" county="Clatsop County" location="North Fork Klaskanne River" municipality="Male" pageId="6" pageNumber="7" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Oregon" typeStatus="holotype">
Male
<typeStatus id="54B106DA3A13FFF4F773FD764EDBC0F3" box="[537,640,652,678]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED1B20BD3A13FFF4F7FAFD764E97C0F3" box="[656,716,652,678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="7" type="Museum">CAS</collectionCode>
) and
<specimenCount id="9D0C73F13A13FFF4F664FD764F06C0F3" box="[782,861,652,678]" pageId="6" pageNumber="7" type="male">3 male</specimenCount>
and
<specimenCount id="9D0C73F13A13FFF4F6FDFD764FA5C0F3" box="[919,1022,652,678]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
<typeStatus id="54B106DA3A13FFF4F16FFD76482EC0F3" box="[1029,1141,652,678]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED1B20BD3A13FFF4F1ECFD764898C0F3" box="[1158,1219,652,678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="7" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED1B20BD3A13FFF4F1A5FD76497FC0F3" box="[1231,1316,652,678]" country="USA" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="6" pageNumber="7" type="Museum">NCSM</collectionCode>
) collected by
<collectorName id="26FFDDAE3A13FFF4F4F5FD4E4E0AC09B" box="[415,593,692,718]" pageId="6" pageNumber="7">E. M. Benedict</collectorName>
on
<date id="FFB49EB83A13FFF4F715FD4E4F39C09B" box="[639,866,692,718]" pageId="6" pageNumber="7" value="1971-11-27">
<collectingDate id="EFF067503A13FFF4F715FD4E4F39C09B" box="[639,866,692,718]" pageId="6" pageNumber="7" value="1971-11-27">27 November 1971</collectingDate>
</date>
along OR hwy.
<quantity id="4CF2159D3A13FFF4F14EFD4E48CFC09B" box="[1060,1172,692,718]" metricMagnitude="5" metricUnit="m" metricValue="3.255702912" pageId="6" pageNumber="7" unit="mi" value="202.3">202, 3 mi</quantity>
(4.8 km)
<collectorName id="26FFDDAE3A13FFF4F060FD4E4D0AC0A3" pageId="6" pageNumber="7">SE Olney</collectorName>
,
<collectingCounty id="62D4C0F43A13FFF4F435FD264E41C0A3" box="[351,538,732,758]" pageId="6" pageNumber="7">Clatsop County</collectingCounty>
,
<collectingRegion id="49CE769A3A13FFF4F74DFD264ED8C0A3" box="[551,643,732,758]" country="United States of America" name="Oregon" pageId="6" pageNumber="7">Oregon</collectingRegion>
.
<collectingMunicipality id="6BD122023A13FFF4F7F3FD264E8EC0A3" box="[665,725,732,758]" pageId="6" pageNumber="7">Male</collectingMunicipality>
<typeStatus id="54B106DA3A13FFF4F7B5FD264F1FC0A3" box="[735,836,732,758]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED1B20BD3A13FFF4F63FFD264FCAC0A3" box="[853,913,732,758]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="7" type="Museum">CAS</collectionCode>
) taken by same collector on same date
<quantity id="4CF2159D3A13FFF4F42DFCFE4D26C14B" box="[327,381,772,798]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="6" pageNumber="7" unit="mi" value="6.0">6 mi</quantity>
(9.6 km)
<collectorName id="26FFDDAE3A13FFF4F49AFCFE4E3AC14B" box="[496,609,772,798]" pageId="6" pageNumber="7">SE Olney</collectorName>
and the
<location id="8ED5EEA33A13FFF4F7AFFCFE484EC14B" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A13FFF4F7AFFCFE484EC14B" box="[709,1045,772,798]" county="Clatsop County" municipality="Male" name="North Fork Klaskanne River" pageId="6" pageNumber="7" stateProvince="Oregon">North Fork Klaskanne River</location>
,
<location id="8ED5EEA33A13FFF4F14BFCFE4881C14B" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A13FFF4F14BFCFE4881C14B" box="[1057,1242,772,798]" county="Clatsop County" municipality="Male" name="Clatsop County" pageId="6" pageNumber="7" stateProvince="Oregon">Clatsop County</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F452FCD64F8FC1EB" blockId="6.[264,1324,652,1878]" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F452FCD64DF4C110" box="[312,431,812,837]" italics="true" pageId="6" pageNumber="7">Diagnosis</emphasis>
. Stem of gonopodal telopodite relatively broad, curving moderately ventrad, with distinct flange distad at level of endomerite; pulvillus oblong; tibiotarsus curving gently dorsad, apically acuminate; endomerite relatively long, breadth and configuration variable, oriented along axis of telopodal stem.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F452FC364854C663" blockId="6.[264,1324,652,1878]" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F452FC364DF8C1B0" box="[312,419,972,997]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="54B106DA3A13FFF4F452FC364DF8C1B0" box="[312,419,972,997]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
. Somatic features as in
<taxonomicName id="4C0AC3FB3A13FFF4F7ACFC364F11C1B0" authorityName="Shelley" authorityYear="2003" box="[710,842,972,997]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B97E646A3A13FFF4F7ACFC364F11C1B0" box="[710,842,972,997]" italics="true" pageId="6" pageNumber="7">R. leonardi</emphasis>
</taxonomicName>
except paranotal margins lightly dentate on segments 1–6, becoming slightly narrower on segment 7 and progressively less dentate caudad. Epiproct with two pairs of relatively short terminal setae.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F452FBBE4F2FC46B" blockId="6.[264,1324,652,1878]" pageId="6" pageNumber="7">
Gonopodal aperture rounded, caudal margin not extending backwards between 9th legs. Gonopods
<emphasis id="B97E646A3A13FFF4F4B4FB964E70C6D0" box="[478,555,1132,1157]" italics="true" pageId="6" pageNumber="7">in situ</emphasis>
extending ventrad in aperture in subparallel arrangement, not extending over anterior margin of aperture or sternum of segment 6. Gonopod structure as follows (
<figureCitation id="1331A4FD3A13FFF4F41EFB464DB3C683" box="[372,488,1212,1238]" captionStart="FIGURES 5–8" captionStartId="7.[264,382,1747,1771]" captionTargetBox="[478,1109,394,1719]" captionTargetId="figure-86@7.[454,1116,391,1097]" captionTargetPageId="7" captionText="FIGURES 5–8. Retrorsia benedictae (all drawn at 400x magnification) 5, telopodite of left gonopod of holotype, medial view. 6, the same, lateral view. 7, telopodite of left gonopod of male from Pacific County, Washington, medial view. 8, the same, lateral view. F, tibiotarsal flange; other abbreviations as in figs. 3–4." figureDoi="http://doi.org/10.5281/zenodo.5014437" httpUri="https://zenodo.org/record/5014437/files/figure.png" pageId="6" pageNumber="7">Figs. 5–6</figureCitation>
): Coxa large, with horizontal, triangular projection directed ventrad, without ventral lobe. Prefemur with margin gently sinusoid, without ventral lobe. Telopodite relatively broad, curving moderately ventrad for most of length, distal expansion in form of discrete concave flange with subacuminate corners, demarcated from tibiotarsus by strong indentation, pulvillus oblong, expanding apically, arising at base of flange; tibiotarsus curving gently dorsad, proximal corner slightly expanded, apically linear and acuminate; endomerite long, apically acuminate, oriented along axis of telopodal stem, curving generally subparallel to tibiotarsus but shorter than latter, with short, triangular, medial and lateral lobes, former overlapping telopodal stem at level of base of tibiotarsus. Prostatic groove with distinct distal chamber.
</paragraph>
<paragraph id="8BB5B8783A13FFF4F452F9B64818C433" blockId="6.[264,1324,652,1878]" box="[312,1091,1612,1638]" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F452F9B64DB5C430" box="[312,494,1612,1637]" italics="true" pageId="6" pageNumber="7">
Male
<typeStatus id="54B106DA3A13FFF4F411F9B64DB5C430" box="[379,494,1612,1637]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
</emphasis>
. Agreeing closely with
<typeStatus id="54B106DA3A13FFF4F665F9B64F2EC433" box="[783,885,1612,1638]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
in all particulars.
</paragraph>
<paragraph id="8BB5B8783A13FFF5F452F98E4D01C30B" blockId="6.[264,1324,652,1878]" lastBlockId="7.[264,1321,284,350]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B97E646A3A13FFF4F452F98E4DFEC4D8" box="[312,421,1652,1677]" italics="true" pageId="6" pageNumber="7">Variation</emphasis>
.
<materialsCitation id="3B62B2253A13FFF5F4D1F98E4D0DC30B" ID-GBIF-Occurrence="3321340312" lastPageId="7" lastPageNumber="8" location="The" municipality="The" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<collectingMunicipality id="6BD122023A13FFF4F4D1F98E4DB2C4DB" box="[443,489,1652,1678]" pageId="6" pageNumber="7">The</collectingMunicipality>
gonopods of the
<collectingRegion id="49CE769A3A13FFF4F7DEF98E4F64C4DB" box="[692,831,1652,1678]" country="United States of America" name="Washington" pageId="6" pageNumber="7">Washington</collectingRegion>
specimen (
<figureCitation id="1331A4FD3A13FFF4F6ACF98E486FC4DB" box="[966,1076,1652,1678]" captionStart="FIGURES 5–8" captionStartId="7.[264,382,1747,1771]" captionTargetBox="[478,1109,394,1719]" captionTargetId="figure-86@7.[454,1116,391,1097]" captionTargetPageId="7" captionText="FIGURES 5–8. Retrorsia benedictae (all drawn at 400x magnification) 5, telopodite of left gonopod of holotype, medial view. 6, the same, lateral view. 7, telopodite of left gonopod of male from Pacific County, Washington, medial view. 8, the same, lateral view. F, tibiotarsal flange; other abbreviations as in figs. 3–4." figureDoi="http://doi.org/10.5281/zenodo.5014437" httpUri="https://zenodo.org/record/5014437/files/figure.png" pageId="6" pageNumber="7">Figs. 7–8</figureCitation>
) differ from those of the
<collectingRegion id="49CE769A3A13FFF4F45EF9664DD7C4E3" box="[308,396,1692,1718]" country="United States of America" name="Oregon" pageId="6" pageNumber="7">Oregon</collectingRegion>
males in that the flange is less concave and has a sublinear margin, the pulvillus is apically rounded and not expanded, and the proximal corner of the tibiotarsus is pointed but not produced into a lobe.
<location id="8ED5EEA33A13FFF4F7AFF9164EA8C553" LSID="urn:lsid:plazi:treatment:03A3096E3A11FFFAF462FC164D33C18B:8ED5EEA33A13FFF4F7AFF9164EA8C553" box="[709,755,1772,1798]" municipality="The" name="The" pageId="6" pageNumber="7" stateProvince="Washington">The</location>
most variable aspect is the endomerite, which is blade­like and broader than that of the
<typeStatus id="54B106DA3A13FFF4F7BEF8EE4F65C57B" box="[724,830,1812,1838]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
. It also lies parallel to the telopodal stem instead of the tibiotarsus and expands apically into two lobes that are similar to those in
<taxonomicName id="4C0AC3FB3A13FFF5F07BF8C64D31C360" authorityName="Shelley" authorityYear="2003" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" lastPageId="7" lastPageNumber="8" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B97E646A3A13FFF5F07BF8C64D31C360" italics="true" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">R. leonardi</emphasis>
</taxonomicName>
, though the terminal one is slightly produced and the proximomedial lobe is absent
</materialsCitation>
.
</paragraph>
<caption id="DF75E8F03A12FFF5F462F9294E60C50E" ID-DOI="http://doi.org/10.5281/zenodo.5014437" ID-Zenodo-Dep="5014437" httpUri="https://zenodo.org/record/5014437/files/figure.png" pageId="7" pageNumber="8" startId="7.[264,382,1747,1771]" targetBox="[478,1109,394,1719]" targetPageId="7">
<paragraph id="8BB5B8783A12FFF5F462F9294E60C50E" blockId="7.[264,1323,1747,1883]" pageId="7" pageNumber="8">
<emphasis id="B97E646A3A12FFF5F462F9294DF6C4BE" bold="true" box="[264,429,1747,1771]" pageId="7" pageNumber="8">FIGURES 5–8</emphasis>
.
<taxonomicName id="4C0AC3FB3A12FFF5F4ABF92E4EC4C4BE" authorityName="Shelley" authorityYear="2003" box="[449,671,1748,1771]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B97E646A3A12FFF5F4ABF92E4EC4C4BE" box="[449,671,1748,1771]" italics="true" pageId="7" pageNumber="8">Retrorsia benedictae</emphasis>
</taxonomicName>
(all drawn at 400x magnification) 5, telopodite of left gonopod of holotype, medial view. 6, the same, lateral view. 7, telopodite of left gonopod of male from Pacific County, Washington, medial view. 8, the same, lateral view. F, tibiotarsal flange; other abbreviations as in figs. 3–4.
</paragraph>
</caption>
<paragraph id="8BB5B8783A1DFFFAF452FEE64ED4C023" blockId="8.[264,1323,284,990]" pageId="8" pageNumber="9">
<emphasis id="B97E646A3A1DFFFAF452FEE64DCDC360" box="[312,406,284,309]" italics="true" pageId="8" pageNumber="9">Ecology</emphasis>
. The
<typeStatus id="54B106DA3A1DFFFAF489FEE64E12C363" box="[483,585,284,310]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
and associated
<typeStatus id="54B106DA3A1DFFFAF66EFEE64F2FC363" box="[772,884,284,310]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
were collected at ca.
<quantity id="4CF2159D3A1DFFFAF11FFEE64898C363" box="[1141,1219,284,310]" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="8" pageNumber="9" unit="ft" value="400.0">400 ft.</quantity>
(
<quantity id="4CF2159D3A1DFFFAF1B9FEE6497BC363" box="[1235,1312,284,310]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="m" value="120.0">120 m</quantity>
) elevation, and the other
<typeStatus id="54B106DA3A1DFFFAF74AFEBE4EDEC30B" box="[544,645,324,350]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
was at
<quantity id="4CF2159D3A1DFFFAF7B6FEBE4F73C30B" box="[732,808,324,350]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="8" pageNumber="9" unit="ft" value="800.0">800 ft.</quantity>
(
<quantity id="4CF2159D3A1DFFFAF65DFEBE4FDDC30B" box="[823,902,324,350]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="9" unit="m" value="240.0">240 m</quantity>
); no habitat information is available for either site. The
<collectingRegion id="49CE769A3A1DFFFAF487FE964E23C3D3" box="[493,632,364,390]" country="United States of America" name="Washington" pageId="8" pageNumber="9">Washington</collectingRegion>
individual was encountered at
<quantity id="4CF2159D3A1DFFFAF68EFE96486BC3D3" box="[996,1072,364,390]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="8" pageNumber="9" unit="ft" value="900.0">900 ft.</quantity>
(
<quantity id="4CF2159D3A1DFFFAF155FE9648D0C3D3" box="[1087,1163,364,390]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="9" unit="m" value="270.0">270 m</quantity>
) elevation on a gentle slope in the Willapa Hills on
<collectingRegion id="49CE769A3A1DFFFAF78AFE6E4FEBC3FB" box="[736,944,404,430]" country="United States of America" name="Washington" pageId="8" pageNumber="9">Washington State</collectingRegion>
Forest lands. The habitat is a young, mixed coastal forest of Douglas­fir (
<taxonomicName id="4C0AC3FB3A1DFFFAF678FE464846C380" box="[786,1053,444,469]" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="menziesii">
<emphasis id="B97E646A3A1DFFFAF678FE464846C380" box="[786,1053,444,469]" italics="true" pageId="8" pageNumber="9">Pseudotsuga menziesii</emphasis>
</taxonomicName>
) and western hemlock (
<taxonomicName id="4C0AC3FB3A1DFFFAF478FE1E4DAEC3A8" box="[274,501,484,509]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="B97E646A3A1DFFFAF478FE1E4DAEC3A8" box="[274,501,484,509]" italics="true" pageId="8" pageNumber="9">Tsuga heterophylla</emphasis>
</taxonomicName>
). Mature trees have been harvested twice on this site, so this is a “third growth” forest approximately 40 years old. Conifers dominate but are mixed with red alder (
<taxonomicName id="4C0AC3FB3A1DFFFAF4E8FDCE4E4BC018" box="[386,528,564,589]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="rubra">
<emphasis id="B97E646A3A1DFFFAF4E8FDCE4E4BC018" box="[386,528,564,589]" italics="true" pageId="8" pageNumber="9">Alnus rubra</emphasis>
</taxonomicName>
) and other hardwoods; the understory vegetation is sparse, consisting primarily of ferns and shrubs.
</paragraph>
<paragraph id="8BB5B8783A1DFFFAF452FD7E4D1AC143" blockId="8.[264,1323,284,990]" pageId="8" pageNumber="9">
<emphasis id="B97E646A3A1DFFFAF452FD7E4D9CC0C8" box="[312,455,644,669]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
. Occurring near the mouth of the Columbia River in northeastern
<collectingRegion id="49CE769A3A1DFFFAF1BAFD7E4973C0CB" box="[1232,1320,644,670]" country="United States of America" name="Oregon" pageId="8" pageNumber="9">Oregon</collectingRegion>
and some
<quantity id="4CF2159D3A1DFFFAF417FD564D9BC093" box="[381,448,684,710]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="8" pageNumber="9" unit="mi" value="25.0">25 mi</quantity>
(
<quantity id="4CF2159D3A1DFFFAF4A5FD564E40C093" box="[463,539,684,710]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="km" value="40.0">40 km</quantity>
) to the north in coastal
<collectingRegion id="49CE769A3A1DFFFAF641FD564FEDC093" box="[811,950,684,710]" country="United States of America" name="Washington" pageId="8" pageNumber="9">Washington</collectingRegion>
(
<figureCitation id="1331A4FD3A1DFFFAF6AFFD564857C093" box="[965,1036,684,710]" captionStart="FIGURE 2" captionStartId="3.[264,367,1802,1826]" captionTargetBox="[521,1074,1077,1791]" captionTargetId="figure-225@3.[502,1075,1077,1791]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Retrorsia; dots, R. leonardi; triangles, R. benedictae. OR, Oregon; WA, Washington." figureDoi="http://doi.org/10.5281/zenodo.5014433" httpUri="https://zenodo.org/record/5014433/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
); the area is about
<quantity id="4CF2159D3A1DFFFAF18FFD564973C093" box="[1253,1320,684,710]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="8" pageNumber="9" unit="mi" value="25.0">25 mi</quantity>
(
<quantity id="4CF2159D3A1DFFFAF47AFD2E4D06C0BB" box="[272,349,724,750]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="km" value="40.0">40 km</quantity>
) west of that of
<taxonomicName id="4C0AC3FB3A1DFFFAF776FD2E4EC5C0B8" authorityName="Shelley" authorityYear="2003" box="[540,670,724,749]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B97E646A3A1DFFFAF776FD2E4EC5C0B8" box="[540,670,724,749]" italics="true" pageId="8" pageNumber="9">R. leonardi</emphasis>
</taxonomicName>
. In addition to the
<typeStatus id="54B106DA3A1DFFFAF6E8FD2E4F99C0BB" box="[898,962,724,750]" pageId="8" pageNumber="9">types</typeStatus>
, the following sample is available:
</paragraph>
<paragraph id="8BB5B8783A1DFFFAF452FCDE4E03C133" blockId="8.[264,1323,284,990]" pageId="8" pageNumber="9">
<materialsCitation id="3B62B2253A1DFFFAF452FCDE4E0FC133" ID-GBIF-Occurrence="3321340302" collectingDate="2002-12-17" collectionCode="CAS" collectorName="J. Ziegltrum &amp; R. Shoals" county="Pacific Co." location="Pacific Co." pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Washington">
<collectingRegion id="49CE769A3A1DFFFAF452FCDE4DA6C16B" box="[312,509,804,830]" country="United States of America" name="Washington" pageId="8" pageNumber="9">WASHINGTON</collectingRegion>
:
<emphasis id="B97E646A3A1DFFFAF760FCDE4ECCC168" box="[522,663,804,829]" italics="true" pageId="8" pageNumber="9">
<collectingCounty id="62D4C0F43A1DFFFAF760FCDE4EC9C168" box="[522,658,804,829]" pageId="8" pageNumber="9">Pacific Co.</collectingCounty>
,
</emphasis>
<quantity id="4CF2159D3A1DFFFAF7CBFCDE4E83C16B" box="[673,728,804,830]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="8" pageNumber="9" unit="mi" value="2.0">2 mi</quantity>
(3.2 km)
<collectorName id="26FFDDAE3A1DFFFAF639FCDE4FABC16B" box="[851,1008,804,830]" pageId="8" pageNumber="9">NW Menlow</collectorName>
,
<specimenCount id="9D0C73F13A1DFFFAF694FCDF4849C162" box="[1022,1042,805,823]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
,
<date id="FFB49EB83A1DFFFAF149FCDE4952C16B" box="[1059,1289,804,830]" pageId="8" pageNumber="9" value="2002-12-17">
<collectingDate id="EFF067503A1DFFFAF149FCDE4952C16B" box="[1059,1289,804,830]" pageId="8" pageNumber="9" value="2002-12-17">17 December 2002</collectingDate>
</date>
,
<collectorName id="26FFDDAE3A1DFFFAF072FCDE4DDBC133" pageId="8" pageNumber="9">J. Ziegltrum</collectorName>
,
<collectorName id="26FFDDAE3A1DFFFAF4E6FCB64DA7C133" box="[396,508,844,870]" pageId="8" pageNumber="9">R. Shoals</collectorName>
(
<collectionCode id="ED1B20BD3A1DFFFAF766FCB64E10C133" box="[524,587,844,870]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="9" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB5B8783A1DFFFAF452FC8E4D33C18B" blockId="8.[264,1323,284,990]" pageId="8" pageNumber="9">
<emphasis id="B97E646A3A1DFFFAF452FC8E4DE8C1D8" box="[312,435,884,909]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
. This species is named for the collector of the
<typeStatus id="54B106DA3A1DFFFAF161FC8E4829C1DB" box="[1035,1138,884,910]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
and
<typeStatus id="54B106DA3A1DFFFAF1D9FC8E497DC1DB" box="[1203,1318,884,910]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
, Ellen M. Benedict, in recognition of her efforts in collecting minute litter organisms in
<collectingRegion id="49CE769A3A1DFFFAF462FC3E4D38C18B" box="[264,355,964,990]" country="United States of America" name="Oregon" pageId="8" pageNumber="9">Oregon</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink