treatments-xml/data/03/BD/EF/03BDEF36F504941DFFAB47F4DDED8A09.xml

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<mods:title id="FA58509A95D3E4B002C846EFB63732EC">A revision of the Malagasy endemic genus Adetomyrma (Hymenoptera: Formicidae: Amblyoponinae)</mods:title>
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<subSubSection id="C30E0DABF504941FFFAB47F4DD638DC8" box="[151,567,1468,1495]" pageId="9" pageNumber="10" type="nomenclature">
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<taxonomicName id="4C1425A3F504941FFFAB47F4DE868DC9" ID-CoL="CL4D9" authority="Yoshimura &amp; Fisher 2012" box="[151,466,1469,1495]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aureocuprea" status="sp. nov.">
<emphasis id="B9608232F504941FFFAB47F4DE868DC9" bold="true" box="[151,466,1469,1495]" italics="true" pageId="9" pageNumber="10">Adetomyrma aureocuprea</emphasis>
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(
<figureCitation id="132F42A5F504941FFFA34796DFA18DE9" box="[159,245,1503,1527]" captionStart="FIGURES 19 – 20" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[151,1435,1102,1635]" captionTargetId="figure@7.[151,1436,1102,1650]" captionTargetPageId="7" captionText="FIGURES 19 – 20. Paramere of Adetomyrma males in lateral view. 19, A. caudapinniger (CASENT 0244415: holotype); 20, A. aureocuprea (CASENT 0227991: holotype). 19, a broad and distinct lobe is present on posterodorsal portion of the paramere; 20, a broad and distinct lobe is absent on posterodorsal portion of the paramere." httpUri="https://zenodo.org/record/213825/files/figure.png" pageId="9" pageNumber="10">Figs 20</figureCitation>
, 
<figureCitation id="132F42A5F504941FFE3D4796DE748DE9" box="[257,288,1503,1527]" captionStart="FIGURES 27 – 28" captionStartId="9.[151,264,817,840]" captionTargetBox="[151,1436,313,786]" captionTargetId="figure@9.[151,1436,313,802]" captionTargetPageId="9" captionText="FIGURES 27 – 28. Eye of Adetomyrma males in full-face view. 27, A. cilium (CASENT 0007808: holotype); 28, A. aureocuprea (CASENT 0227991: holotype). 27, hairs on the eye longer than 0.5 × length of horizontal diameter of the mid ocellus; 28, hairs on the eye are shorter than 0.25 × length of horizontal diameter of the mid ocellus." httpUri="https://zenodo.org/record/213828/files/figure.png" pageId="9" pageNumber="10">28</figureCitation>
, 
<figureCitation id="132F42A5F504941FFE174796DE1E8DE9" box="[299,330,1503,1527]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="9" pageNumber="10">29</figureCitation>
, 
<figureCitation id="132F42A5F504941FFE694796DE278DE9" box="[341,371,1503,1527]" captionStart="FIGURES 38 – 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 – 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="9" pageNumber="10">38</figureCitation>
, 
<figureCitation id="132F42A5F504941FFE434796DEC98DE9" box="[383,413,1503,1527]" captionStart="FIGURES 47 – 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 – 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="9" pageNumber="10">47</figureCitation>
, 
<figureCitation id="132F42A5F504941FFE954796DE938DE9" box="[425,455,1503,1527]" captionStart="FIGURES 56 – 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 – 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="9" pageNumber="10">56</figureCitation>
, 
<figureCitation id="132F42A5F504941FFEEF4796DEA48DE9" box="[467,496,1503,1527]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="9" pageNumber="10">64</figureCitation>
)
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<collectingMunicipality id="6BCFC45AF504941FFC9B446EDB6D8E21" box="[935,1081,1575,1599]" pageId="9" pageNumber="10">Fianarantsoa</collectingMunicipality>
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354° N
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alt., gallery forest, 
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<collectorName id="26E13BF6F504941FFBC84402DA088E7D" box="[1268,1372,1611,1635]" pageId="9" pageNumber="10">Griswold</collectorName>
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<collectorName id="26E13BF6F504941FFA584404DAC88E7A" box="[1380,1436,1612,1636]" pageId="9" pageNumber="10">et al.</collectorName>
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leg. [
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<typeStatus id="54AFE082F504941FFFFB44DDDE158EB2" box="[199,321,1684,1708]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
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<specimenCode id="DBB2F65BF504941FFB6844DADA768EB5" box="[1108,1314,1683,1707]" pageId="9" pageNumber="10">CASENT0490920</specimenCode>
[
<collectionCode id="ED05C6E5F504941FFA0F44DADAC58EB5" box="[1331,1425,1683,1707]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="10">MHNG</collectionCode>
]
</materialsCitation>
, 
<materialsCitation id="3B7C547DF504941FFFAB44FEDE818ED1" ID-GBIF-Occurrence="3446753315" box="[151,469,1719,1743]" collectingDate="2003-01-27" collectingDateMax="2003-01-31" collectingDateMin="2003-01-27" collectingMethod="at light" collectionCode="MCZC" collectorName="Fisher &amp; Griswold &amp; et al." country="Madagascar" elevation="725" latitude="-22.481667" location="Parc National d'Isalo" longLatPrecision="125" longitude="45.461666" municipality="Fianarantsoa" pageId="9" pageNumber="10" specimenCode="CASENT0490924" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
CASENT0490924 [
<collectionCode id="ED05C6E5F504941FFE4F44FEDE9E8ED1" box="[371,458,1719,1743]" pageId="9" pageNumber="10">MCZC</collectionCode>
],
</materialsCitation>
with same data as 
<typeStatus id="54AFE082F504941FFD9644FEDC5B8ED1" box="[682,783,1719,1743]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF504941CFFFB4495DE188A22" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8BAB5E20F504941FFFFB4495DD4A8EED" blockId="9.[151,1436,1575,2031]" box="[199,542,1755,1780]" pageId="9" pageNumber="10">
<emphasis id="B9608232F504941FFFFB4495DEF28EEA" bold="true" box="[199,422,1756,1780]" pageId="9" pageNumber="10">Worker and queen</emphasis>
unknown.
</paragraph>
<paragraph id="8BAB5E20F504941FFFFB4549DD588F25" blockId="9.[151,1436,1575,2031]" pageId="9" pageNumber="10">
<emphasis id="B9608232F504941FFFFB4549DEF08F06" bold="true" box="[199,420,1791,1816]" pageId="9" pageNumber="10">Male. Description.</emphasis>
Measurements: 
<typeStatus id="54AFE082F504941FFD5D44B6DD938F09" box="[609,711,1791,1815]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
. HL 0.57, HW 0.87, SL 0.16, EL 0.43, WL 1.42, MnW 0.9, CI 152.4, SI 18, EI 74.8, MnI 103.9.
</paragraph>
<paragraph id="8BAB5E20F504941FFFF54501DCC68F9D" blockId="9.[151,1436,1575,2031]" pageId="9" pageNumber="10">
HL 0.44–0.63, HW 0.59–0.91, SL 0.11–0.17, EL 0.3–0.45, WL 1.03–1.54, MnW 0.63–0.88, CI 135.6–145.7, SI 17.9–19.6, EI 67.4–75, MnI 97.1–106.3 (
<specimenCount id="9D1295A9F504941FFDB84522DC5A8F9D" box="[644,782,1899,1923]" pageId="9" pageNumber="10" type="generic">5 specimens</specimenCount>
measured).
</paragraph>
<paragraph id="8BAB5E20F504941CFFF545D9DA5E88CA" blockId="9.[151,1436,1575,2031]" lastBlockId="10.[151,1437,151,2012]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Eye large and prominent, varied in size, but posterior margin not exceeding posterior margin of mid ocellus in full-face view (
<figureCitation id="132F42A5F504941FFE7845FADEC78FD5" box="[324,403,1971,1995]" captionStart="FIGURES 47 – 55" captionStartId="14.[151,264,1683,1706]" captionTargetBox="[151,1435,651,1651]" captionTargetId="figure@14.[151,1436,651,1663]" captionTargetPageId="14" captionText="FIGURES 47 – 55. Head of Adetomyrma males in full-face view. 47, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 48, A. bressleri sp. nov. (CASENT 0008693); 49, A. caputleae sp. nov. (CASENT 0079552); 50, A. cassis sp. nov. (CASENT 0163620: holotype); 51, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 52, A. cilium sp. nov. (CASENT 0007808: holotype); 53, A. clarivida sp. nov. (CASENT 0064259: holotype); 54, A. goblin sp. nov. (CASENT 0084070); 55, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213831/files/figure.png" pageId="9" pageNumber="10">Fig 47</figureCitation>
). Distance between lateral ocellus and eye about same as diameter of lateral ocellus. Palpal formula 3,3 (three maxillary and three labial). Notaulus absent on mesoscutum. Parapsidal line usually unclear, sometimes weakly impressed but never pigmented with darker color (
<figureCitation id="132F42A5F507941CFC8442DEDB5C88B1" box="[952,1032,151,175]" captionStart="FIGURES 38 – 46" captionStartId="13.[151,264,1423,1446]" captionTargetBox="[151,1432,685,1400]" captionTargetId="figure@13.[151,1436,685,1416]" captionTargetPageId="13" captionText="FIGURES 38 – 46. Males of Adetomyrma in dorsal view. 38, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 39, A. bressleri sp. nov. (CASENT 0008693); 40, A. caputleae sp. nov. (CASENT 0079552); 41, A. cassis sp. nov. (CASENT 0163620: holotype); 42, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 43, A. cilium sp. nov. (CASENT 0007808: holotype); 44, A. clarivida sp. nov. (CASENT 0064259: holotype); 45, A. goblin sp. nov. (CASENT 0084070); 46, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213830/files/figure.png" pageId="10" pageNumber="11">Fig 38</figureCitation>
). Anterior margin of petiole longer than dorsal margin in lateral view (
<figureCitation id="132F42A5F507941CFD2242F5DD3888CA" box="[542,620,188,212]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="10" pageNumber="11">Fig 29</figureCitation>
). Subpetiolar process not developed, without hairs (
<figureCitation id="132F42A5F507941CFB8C42F5DBAA88CA" box="[1200,1278,188,212]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="10" pageNumber="11">Fig 29</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F507941CFFF54296DE3C89B2" blockId="10.[151,1437,151,2012]" pageId="10" pageNumber="11">
Left and right parameres not overlapping or narrowly overlapping on dorsal small part of basimere (
<figureCitation id="132F42A5F507941CFA7C4296DAC488E9" box="[1344,1424,223,247]" captionStart="FIGURES 19 – 20" captionStartId="7.[151,264,1672,1695]" captionTargetBox="[151,1435,1102,1635]" captionTargetId="figure@7.[151,1436,1102,1650]" captionTargetPageId="7" captionText="FIGURES 19 – 20. Paramere of Adetomyrma males in lateral view. 19, A. caudapinniger (CASENT 0244415: holotype); 20, A. aureocuprea (CASENT 0227991: holotype). 19, a broad and distinct lobe is present on posterodorsal portion of the paramere; 20, a broad and distinct lobe is absent on posterodorsal portion of the paramere." httpUri="https://zenodo.org/record/213825/files/figure.png" pageId="10" pageNumber="11">Fig 20</figureCitation>
). No distinct projection or lobe present on posterodorsal portion of paramere. Basal ring not reduced, covering whole anterior margin of paramere in lateral view. Basal projection on cuspis clear but not extraordinarily well developed. Aedeagus in lateral view as in 
<figureCitation id="132F42A5F507941CFD2C4305DDD2897A" box="[528,646,332,356]" captionStart="FIGURES 56 – 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 – 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="10" pageNumber="11">Figure 56</figureCitation>
: distal portion narrowed distal to ventral projection, apical margin relatively sharp, but without small projection on its posteroventral portion, posteroventral margin of ventral projection convex.
</paragraph>
<paragraph id="8BAB5E20F507941CFFF543FEDE188A22" blockId="10.[151,1437,151,2012]" pageId="10" pageNumber="11">
Hair on compound eyes short, about 0.25× diameter of mid ocellus (
<figureCitation id="132F42A5F507941CFCF343FEDB4A89D1" box="[975,1054,439,463]" captionStart="FIGURES 27 – 28" captionStartId="9.[151,264,817,840]" captionTargetBox="[151,1436,313,786]" captionTargetId="figure@9.[151,1436,313,802]" captionTargetPageId="9" captionText="FIGURES 27 – 28. Eye of Adetomyrma males in full-face view. 27, A. cilium (CASENT 0007808: holotype); 28, A. aureocuprea (CASENT 0227991: holotype). 27, hairs on the eye longer than 0.5 × length of horizontal diameter of the mid ocellus; 28, hairs on the eye are shorter than 0.25 × length of horizontal diameter of the mid ocellus." httpUri="https://zenodo.org/record/213828/files/figure.png" pageId="10" pageNumber="11">Fig 28</figureCitation>
). With mesofemur in dorsal view, anterior face with dense appressed hairs, and sometimes several longer subdecumbent hairs on basal portion. Ventral margin of eye not edged with darker pigment or punctures. Body bicolored, head brown and remainder yellow (
<figureCitation id="132F42A5F507941CFFCE406DDE148A22" box="[242,320,548,572]" captionStart="FIGURES 29 – 37" captionStartId="12.[151,264,1414,1437]" captionTargetBox="[151,1435,447,1381]" captionTargetId="figure@12.[151,1436,447,1407]" captionTargetPageId="12" captionText="FIGURES 29 – 37. Males of Adetomyrma in lateral view. 29, A. aureocuprea sp. nov. (CASENT 0227991: holotype); 30, A. bressleri sp. nov. (CASENT 0008693); 31, A. caputleae sp. nov. (CASENT 0079552); 32, A. cassis sp. nov. (CASENT 0163620: holotype); 33, A. caudapinniger sp. nov. (CASENT 0244415: holotype); 34, A. cilium sp. nov. (CASENT 0007808: holotype); 35, A. clarivida sp. nov. (CASENT 0064259: holotype); 36, A. goblin sp. nov. (CASENT 0084070); 37, A. venatrix (CASENT 0151606)." httpUri="https://zenodo.org/record/213829/files/figure.png" pageId="10" pageNumber="11">Fig 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF507941CFFF54001DBAE8A9A" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BAB5E20F507941CFFF54001DBAE8A9A" blockId="10.[151,1437,151,2012]" pageId="10" pageNumber="11">
<emphasis id="B9608232F507941CFFF54001DE1B8A7E" bold="true" box="[201,335,584,608]" pageId="10" pageNumber="11">Etymology.</emphasis>
This species name is derived from the Latin words aureus (golden) and cupreus (coppery), referring to the body coloration. The species epithet is treated as a noun in apposition, and thus invariant.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF507941CFFF540C6DDB98AB9" box="[201,749,655,680]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BAB5E20F507941CFFF540C6DDB98AB9" blockId="10.[151,1437,151,2012]" box="[201,749,655,680]" pageId="10" pageNumber="11">
<emphasis id="B9608232F507941CFFF540C6DE368AB6" bold="true" box="[201,354,655,680]" pageId="10" pageNumber="11">Distribution.</emphasis>
<collectingCountry id="F3031EB0F507941CFE5540C6DD618AB9" box="[361,565,655,679]" name="Madagascar" pageId="10" pageNumber="11">MADAGASCAR</collectingCountry>
: as in 
<figureCitation id="132F42A5F507941CFD4640C6DDBD8AB9" box="[634,745,655,679]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="10" pageNumber="11">Figure 64</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF507941CFFF540FADE438F52" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BAB5E20F507941CFFF540FADE438F52" blockId="10.[151,1437,151,2012]" pageId="10" pageNumber="11">
<emphasis id="B9608232F507941CFFF540FADD7A8AD2" bold="true" box="[201,558,691,716]" pageId="10" pageNumber="11">Additional material examined</emphasis>
: in addition to the 
<typeStatus id="54AFE082F507941CFC3F40FCDC678AD3" box="[771,819,693,717]" pageId="10" pageNumber="11">type</typeStatus>
material, specimens from the following localities were examined in this study: 
<collectingCountry id="F3031EB0F507941CFE8D409EDD288AF1" box="[433,636,727,751]" name="Madagascar" pageId="10" pageNumber="11">MADAGASCAR</collectingCountry>
. Parc National Montagne d'Ambre [1st campsite], 
<geoCoordinate id="EE2038E7F507941CFBF2409EDA308AF1" box="[1230,1380,727,751]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-12.5145">12° 30.87´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA49409EDFAD8B0A" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="49.18133">49° 10.88´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFE3840B5DE1E8B0A" box="[260,330,764,788]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="10" pageNumber="11" unit="m" value="960.0">960 m</quantity>
alt., rainforest; Ampasindava, Forêt d'Ambilanivy, 
<quantity id="4CECF3C5F507941CFCB440B5DC818B0A" box="[904,981,764,788]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="10" pageNumber="11" unit="km" value="3.9">3.9 km</quantity>
181° S Ambaliha, 
<geoCoordinate id="EE2038E7F507941CFB9540B5DA628B0A" box="[1193,1334,764,788]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-13.798667">13° 47.92´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA7E40B5DFFF8B26" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="48.161667">48° 9.7´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFF8A4156DFAA8B26" box="[182,254,799,824]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="m" value="600.0">600 m</quantity>
alt., rainforest; Sofia Region, district of Port-Berger, Ambovomamy 
<quantity id="4CECF3C5F507941CFB354156DB058B29" box="[1033,1105,799,823]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="20.0">20 km</quantity>
N of Port-Berger, 
<geoCoordinate id="EE2038E7F507941CFA194156DFFC8B42" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-15.451167">15° 27.07´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFF8A410DDE688B42" box="[182,316,836,860]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="47.613335">47° 36.8´E</geoCoordinate>
, 
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alt., secondary forest on white sandy area; Boeny Region, district of Marovoay, Ampijoroa National Park, 
<quantity id="4CECF3C5F507941CFE7F412EDECF8B61" box="[323,411,871,895]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="11" unit="km" value="160.0">160 km</quantity>
North of Maevatanana on RN 0 4, 
<geoCoordinate id="EE2038E7F507941CFC11412EDCEB8B61" box="[813,959,871,895]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-16.319334">16° 19.16´S</geoCoordinate>
, 
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, 
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alt., deciduous forest; Parc National de Namoroka, 
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317° NW Vilanandro, 
<geoCoordinate id="EE2038E7F507941CFC0B41C5DCE28BBA" box="[823,950,908,932]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-16.406666">16° 24.4´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFCFE41C5DB118BBA" box="[962,1093,908,932]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="45.31">45° 18.6´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFB7341C5DBC28BBA" box="[1103,1174,908,932]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="100.0">100 m</quantity>
alt., tropical dry forest; Boeny Region, district of Soalala Analamanitra forest, 
<quantity id="4CECF3C5F507941CFDC441E6DC688BD9" box="[760,828,943,967]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="11" unit="km" value="14.0">14 km</quantity>
SW of Mitsinjo, 
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, 
<geoCoordinate id="EE2038E7F507941CFB4841E6DBB58BD6" box="[1140,1249,943,968]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="45.7">45° 42´E</geoCoordinate>
, 
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alt., dense dry forest; Betsiboka Bassin, Riv: Mamokomita, Manjokavaradrano, 
<geoCoordinate id="EE2038E7F507941CFCA1419DDB5C8BF2" box="[925,1032,980,1004]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-17.633333">17° 38´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFB28419DDBF18BF2" box="[1044,1189,980,1004]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="46.9055">46° 54.33´E</geoCoordinate>
; Parc National Tsingy de Bemaraha, 
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ESE 123° Antsalova, 
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, 
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, 
<quantity id="4CECF3C5F507941CFCEE41BEDB4D8C0E" box="[978,1049,1015,1040]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="m" value="150.0">150 m</quantity>
alt., tropical dry forest on Tsingy; Menabe Region, district of Morondava, Beroboka village NE of Morondava, Antsarongaza dry forest 
<quantity id="4CECF3C5F507941CFA244655DA328C2A" box="[1304,1382,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="11" unit="km" value="7.5">7.5 km</quantity>
E of Beroboka, 
<geoCoordinate id="EE2038E7F507941CFE2E4676DEF68C49" box="[274,418,1087,1111]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-19.9775">19° 58.65´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFE934676DD168C46" box="[431,578,1087,1112]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.666332">44° 39.98´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFD714676DDD38C46" box="[589,647,1087,1112]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="m" value="50.0">50 m</quantity>
alt., dry forest; Menabe Region, district of Morondava, Beroboka village NE of Morondava, Antsarongaza gallery forest 
<quantity id="4CECF3C5F507941CFDD7462CDC728C62" box="[747,806,1124,1148]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="11" unit="km" value="7.0">7 km</quantity>
E of Beroboka, 
<geoCoordinate id="EE2038E7F507941CFCDF462DDB208C62" box="[995,1140,1124,1148]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-19.9775">19° 58.65´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFBBD462DDA408C62" box="[1153,1300,1124,1148]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.665333">44° 39.92´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFA1E462DDA3E8C62" box="[1314,1386,1124,1148]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="10" pageNumber="11" unit="m" value="128.0">128 m</quantity>
alt., gallery forest; Forêt d'Atsirakambiaty, 
<quantity id="4CECF3C5F507941CFD7046C1DDCF8CBE" box="[588,667,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="10" pageNumber="11" unit="km" value="7.6">7.6 km</quantity>
285° WNW Itremo, 
<geoCoordinate id="EE2038E7F507941CFCB646C1DB598CBE" box="[906,1037,1160,1184]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-20.593334">20° 35.6´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFB2646C1DBCB8CBE" box="[1050,1183,1160,1184]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="46.56333">46° 33.8´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFB9046C1DA568CBE" box="[1196,1282,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="10" pageNumber="11" unit="m" value="1550.0">1550 m</quantity>
alt., montane rainforest; Base Makay, 
<geoCoordinate id="EE2038E7F507941CFE8C46E5DD108CDA" box="[432,580,1196,1220]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-21.222834">21° 13.37´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFD6F46E5DDBD8CDA" box="[595,745,1196,1220]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="45.324833">45° 19.49´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFDCB46E5DC158CDA" box="[759,833,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="11" unit="m" value="490.0">490 m</quantity>
alt., gallery forest on sandy soil; Atsimo Andrefana Region, district of Betioky, 
<quantity id="4CECF3C5F507941CFEE24699DD7D8CF6" box="[478,553,1232,1256]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="30.0">30 km</quantity>
E Betioky, Beza Mahafaly Special Reserve (around Research Station), 
<geoCoordinate id="EE2038E7F507941CFA494699DFA28D12" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.6865">23° 41.19´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFE3F46BDDEC68D12" box="[259,402,1268,1292]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.591">44° 35.46´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFEA146BDDEB08D12" box="[413,484,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="11" unit="m" value="165.0">165 m</quantity>
alt., gallery dry deciduous forest; Parc National d'Isalo, 
<quantity id="4CECF3C5F507941CFB6B46BDDBF08D12" box="[1111,1188,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="10" pageNumber="11" unit="km" value="9.1">9.1 km</quantity>
354° N Ranohira, 
<geoCoordinate id="EE2038E7F507941CFA4946BDDFB88D2E" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-22.481667">22° 28.9´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFFC74751DED68D2E" box="[251,386,1304,1328]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="45.461666">45° 27.7´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFEAD4751DE8F8D2E" box="[401,475,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="10" pageNumber="11" unit="m" value="725.0">725 m</quantity>
alt., gallery forest; 
<quantity id="4CECF3C5F507941CFDFC4751DDAF8D2E" box="[704,763,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="km" value="1.0">1 km</quantity>
E of Isalo National Park Interpretive Center, Fianarantsoa Prov., 
<geoCoordinate id="EE2038E7F507941CFFE34775DE0E8D4A" box="[223,346,1340,1364]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-22.626667">22° 37.6´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFE5A4775DEAC8D4A" box="[358,504,1340,1364]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="45.358166">45° 21.49´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFD3F4775DD1F8D4A" box="[515,587,1340,1364]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="10" pageNumber="11" unit="m" value="885.0">885 m</quantity>
alt., dry wash; Horombe Region, district of Ihosy, Betapia (Border of Fianarantsoa and Tulear): 
<quantity id="4CECF3C5F507941CFEAD4729DE988D66" box="[401,460,1376,1400]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="km" value="9.0">9 km</quantity>
SW of Ilakaka Saphir town, 
<geoCoordinate id="EE2038E7F507941CFC234729DCE68D66" box="[799,946,1376,1400]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-22.628834">22° 37.73´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFCFC4729DB028D66" box="[960,1110,1376,1400]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="45.361168">45° 21.67´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFB584729DBEF8D66" box="[1124,1211,1376,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="10" pageNumber="11" unit="m" value="1036.0">1036 m</quantity>
alt., Uapaca forest; Horombe Region, district of Ihosy, Isalo National Park, 
<quantity id="4CECF3C5F507941CFC3447CDDC1B8D82" box="[776,847,1412,1436]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="m" value="900.0">900 m</quantity>
E of 
<collectionCode id="ED05C6E5F507941CFCB747CDDCB88D82" box="[907,1004,1412,1436]" pageId="10" pageNumber="11">ANGAP</collectionCode>
Interpretation Center, 
<geoCoordinate id="EE2038E7F507941CFBD447CDDA3C8D82" box="[1256,1384,1412,1436]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-22.626667">22° 37.6´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA4947CDDFAD8DDE" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="45.358166">45° 21.49´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFE3847E1DE1F8DDE" box="[260,331,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="10" pageNumber="11" unit="m" value="701.0">701 m</quantity>
alt., open area near stream; Manombo Special Reserve, 
<quantity id="4CECF3C5F507941CFC8347E1DB518DDE" box="[959,1029,1448,1472]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="11" unit="km" value="32.0">32 km</quantity>
SE of Farafangana, 
<geoCoordinate id="EE2038E7F507941CFBD547E1DA3C8DDE" box="[1257,1384,1448,1472]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.021833">23° 1.31´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA4947E1DFBF8DFA" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="47.72">47° 43.2´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFFCA4785DE7B8DFA" box="[246,303,1484,1508]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="11" unit="m" value="36.0">36 m</quantity>
alt., lowland rainforest; Atsimo Andrefana Region, district of Tulear II, Mikea deciduous dry forest 
<quantity id="4CECF3C5F507941CFAB24785DFEF8E16" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="3.0">3 km</quantity>
N Andranomavo village, 
<geoCoordinate id="EE2038E7F507941CFEE347B9DD3A8E16" box="[479,622,1520,1544]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-22.903667">22° 54.22´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFD4747B9DC5F8E16" box="[635,779,1520,1544]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="43.4755">43° 28.53´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFC2447B9DC058E16" box="[792,849,1520,1544]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="m" value="30.0">30 m</quantity>
alt., deciduous dry forest; Fiherenana, 
<geoCoordinate id="EE2038E7F507941CFA3B47B9DAC28E16" box="[1287,1430,1520,1544]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.177">23° 10.62´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFFAB445DDE7D8E32" box="[151,297,1556,1580]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="43.960835">43° 57.65´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFE09445DDE288E32" box="[309,380,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="100.0">100 m</quantity>
alt., gallery forest; Fiherenana, 
<geoCoordinate id="EE2038E7F507941CFDD8445DDC208E32" box="[740,884,1556,1580]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.235332">23° 14.12´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFCBD445DDB468E32" box="[897,1042,1556,1580]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="43.870834">43° 52.25´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFB23445DDB0C8E32" box="[1055,1112,1556,1580]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="m" value="50.0">50 m</quantity>
alt., degraded gallery forest; Sept Lacs, 
<geoCoordinate id="EE2038E7F507941CFE284471DEF08E4E" box="[276,420,1592,1616]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.520834">23° 31.25´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFE8D4471DD618E4E" box="[433,565,1592,1616]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.159668">44° 9.58´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFD7E4471DDDD8E4E" box="[578,649,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="m" value="130.0">130 m</quantity>
alt., gallery forest mixed with spiny thicket trees; Forêt de Mite, 
<quantity id="4CECF3C5F507941CFA574471DFEF8E6A" metricMagnitude="4" metricUnit="m" metricValue="2.07" pageId="10" pageNumber="11" unit="km" value="20.7">20.7 km</quantity>
29° WNW Tongobory, 
<geoCoordinate id="EE2038E7F507941CFEF94415DD078E6A" box="[453,595,1628,1652]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.524166">23° 31.45´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFD634415DDB68E6A" box="[607,738,1628,1652]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.121334">44° 7.28´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFDD14415DC728E6A" box="[749,806,1628,1652]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="11" unit="m" value="75.0">75 m</quantity>
alt., gallery forest; Sept Lacs, 
<geoCoordinate id="EE2038E7F507941CFB464415DA5D8E6A" box="[1146,1289,1628,1652]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.5275">23° 31.65´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA294415DAC38E6A" box="[1301,1431,1628,1652]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.1545">44° 9.27´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFFAB44C9DF9B8E86" box="[151,207,1664,1688]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="11" unit="m" value="70.0">70 m</quantity>
alt., gallery forest; Parcel I, Beza Mahafaly Reserve, near research station, Tulear Province, 
<geoCoordinate id="EE2038E7F507941CFBE644C9DA3C8E86" box="[1242,1384,1664,1688]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.6865">23° 41.19´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFA4944C9DFAF8EA2" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.591">44° 35.46´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFE3B44EDDE1B8EA2" box="[263,335,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="11" unit="m" value="165.0">165 m</quantity>
alt., dry deciduous forest; Parcel II, Beza Mahafaly Reserve, near Bellevue, Tulear Province, 
<geoCoordinate id="EE2038E7F507941CFA4944EDDFAC8EFE" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-23.689833">23° 41.39´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFE394481DECC8EFE" box="[261,408,1736,1760]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="44.5755">44° 34.53´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFE994481DEB98EFE" box="[421,493,1736,1760]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="11" unit="m" value="180.0">180 m</quantity>
alt., spiny forest; Parc National d'Andohahela, Forêt de Manantalinjo, 
<quantity id="4CECF3C5F507941CFA2C4481DA398EFE" box="[1296,1389,1736,1760]" metricMagnitude="4" metricUnit="m" metricValue="3.36" pageId="10" pageNumber="11" unit="km" value="33.6">33.6 km</quantity>
63° ENE Amboasary, 
<quantity id="4CECF3C5F507941CFE5444A5DEEE8F1A" box="[360,442,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="10" pageNumber="11" unit="km" value="7.6">7.6 km</quantity>
99° E Hazofotsy, 
<geoCoordinate id="EE2038E7F507941CFDAF44A5DC7D8F1A" box="[659,809,1772,1796]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-24.817">24° 49.02´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFC0444A5DC968F1A" box="[824,962,1772,1796]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="46.61">46° 36.6´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFCED44A5DB4F8F1A" box="[977,1051,1772,1796]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="m" value="150.0">150 m</quantity>
alt., spiny forest/thicket; Androy Region, district of Tsihombe, 
<quantity id="4CECF3C5F507941CFED54559DD658F36" box="[489,561,1808,1832]" metricMagnitude="4" metricUnit="m" metricValue="7.4" pageId="10" pageNumber="11" unit="km" value="74.0">74 km</quantity>
S of Tsihombe, Cap Ste Marie Reserve, 
<geoCoordinate id="EE2038E7F507941CFB3C4559DBC58F36" box="[1024,1169,1808,1832]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-25.587667">25° 35.26´S</geoCoordinate>
, 
<geoCoordinate id="EE2038E7F507941CFBA24559DA708F36" box="[1182,1316,1808,1832]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="45.163">45° 9.78´E</geoCoordinate>
, 
<quantity id="4CECF3C5F507941CFA0C4559DA3E8F36" box="[1328,1386,1808,1832]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="11" unit="m" value="37.0">37 m</quantity>
alt., spiny bush.
</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF507941DFFF54511DB2B89EA" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BAB5E20F507941CFFF54511DE168FC2" blockId="10.[151,1437,151,2012]" pageId="10" pageNumber="11">
<emphasis id="B9608232F507941CFFF54511DE698F6E" bold="true" box="[201,317,1880,1904]" pageId="10" pageNumber="11">Remarks.</emphasis>
<taxonomicName id="4C1425A3F507941CFE794511DD3E8F6E" box="[325,618,1880,1904]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F507941CFE794511DD3E8F6E" box="[325,618,1880,1904]" italics="true" pageId="10" pageNumber="11">Adetomyrma aureocuprea</emphasis>
</taxonomicName>
is only known from males. 
<taxonomicName id="4C1425A3F507941CFC914511DB0E8F6E" box="[941,1114,1880,1904]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F507941CFC914511DB0E8F6E" box="[941,1114,1880,1904]" italics="true" pageId="10" pageNumber="11">A. aureocuprea</emphasis>
</taxonomicName>
is easily separable from the other 
<taxonomicName id="4C1425A3F507941CFFE14535DE388F8A" box="[221,364,1916,1940]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F507941CFFE14535DE388F8A" box="[221,364,1916,1940]" italics="true" pageId="10" pageNumber="11">Adetomyrma</emphasis>
</taxonomicName>
males by the yellowish body color, no mesoscutal notaulus, poorly developed subpetiolar process, lack of posterodorsal projection or lobe on the paramere, short hairs on the compound eye, and vestigial parapsidal line.
</paragraph>
<paragraph id="8BAB5E20F506941DFFF542DEDB9389D1" blockId="11.[151,1436,151,535]" pageId="11" pageNumber="12">
The males of 
<taxonomicName id="4C1425A3F506941DFE5A42D1DDDE88AE" box="[358,650,152,176]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F506941DFE5A42D1DDDE88AE" box="[358,650,152,176]" italics="true" pageId="11" pageNumber="12">Adetomyrma aureocuprea</emphasis>
</taxonomicName>
display remarkable morphological variation in, for example, the size of the eye and ocelli, head shape, mesonotal shape, petiolar shape, and hairs on body surface. 
<taxonomicName id="4C1425A3F506941DFB8542F5DA3188CA" box="[1209,1381,188,212]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F506941DFB8542F5DA3188CA" box="[1209,1381,188,212]" italics="true" pageId="11" pageNumber="12">A. aureocuprea</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFA4942F5DFE188E9" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 64</figureCitation>
) is completely sympatric with 
<taxonomicName id="4C1425A3F506941DFD2B42A9DDCD88E6" box="[535,665,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bressleri">
<emphasis id="B9608232F506941DFD2B42A9DDCD88E6" box="[535,665,224,248]" italics="true" pageId="11" pageNumber="12">A. bressleri</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFD954296DDAC88E9" box="[681,760,223,247]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 65</figureCitation>
), 
<taxonomicName id="4C1425A3F506941DFC3242A9DCC888E6" box="[782,924,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="caputleae">
<emphasis id="B9608232F506941DFC3242A9DCC888E6" box="[782,924,224,248]" italics="true" pageId="11" pageNumber="12">A. caputleae</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFC904296DCAF88E9" box="[940,1019,223,247]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 66</figureCitation>
), 
<taxonomicName id="4C1425A3F506941DFB2D42A9DB2D88E6" box="[1041,1145,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="goblin">
<emphasis id="B9608232F506941DFB2D42A9DB2D88E6" box="[1041,1145,224,248]" italics="true" pageId="11" pageNumber="12">A. goblin</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFBB54296DB8388E9" box="[1161,1239,223,247]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 71</figureCitation>
), and 
<taxonomicName id="4C1425A3F506941DFA2342A9DAC888E6" box="[1311,1436,224,248]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B9608232F506941DFA2342A9DAC888E6" box="[1311,1436,224,248]" italics="true" pageId="11" pageNumber="12">A. venatrix</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFFA3434DDFA68902" box="[159,242,260,284]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 72</figureCitation>
), and was collected within a 
<quantity id="4CECF3C5F506941DFD66434DDDF38902" box="[602,679,260,284]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="km" value="20.0">20 km</quantity>
radius of 
<taxonomicName id="4C1425A3F506941DFC16434DDCC18902" box="[810,917,260,284]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cilium">
<emphasis id="B9608232F506941DFC16434DDC6F8902" box="[810,827,260,284]" italics="true" pageId="11" pageNumber="12">A</emphasis>
. 
<emphasis id="B9608232F506941DFC73434DDCC18902" box="[847,917,260,284]" italics="true" pageId="11" pageNumber="12">cilium</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFC96434DDCAF8902" box="[938,1019,260,284]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 69</figureCitation>
) and within a 
<quantity id="4CECF3C5F506941DFB88434DDA558902" box="[1204,1281,260,284]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="km" value="70.0">70 km</quantity>
radius of 
<taxonomicName id="4C1425A3F506941DFAB8434DDE6E895E" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="caudapinniger">
<emphasis id="B9608232F506941DFAB8434DDAC18902" box="[1412,1429,260,284]" italics="true" pageId="11" pageNumber="12">A</emphasis>
. 
<emphasis id="B9608232F506941DFFAB4361DE6E895E" box="[151,314,296,320]" italics="true" pageId="11" pageNumber="12">caudapinniger</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFE74436EDEC28921" box="[328,406,295,319]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 68</figureCitation>
). The morphological differences between all species are clear and consistent in each case of sympatric and geographically close localities, even though apparent similarity may be shown to a character of another 
<taxonomicName id="4C1425A3F506941DFFCD4339DED48996" box="[241,384,368,392]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B9608232F506941DFFCD4339DED48996" box="[241,384,368,392]" italics="true" pageId="11" pageNumber="12">Adetomyrma</emphasis>
</taxonomicName>
species collected from distant localities. In addition to the above species, the distribution of 
<taxonomicName id="4C1425A3F506941DFAB84339DE7089B2" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F506941DFAB84339DE7089B2" italics="true" pageId="11" pageNumber="12">A. aureocuprea</emphasis>
</taxonomicName>
is parapatric with 
<taxonomicName id="4C1425A3F506941DFD3643DDDDC789B2" box="[522,659,404,428]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="clarivida">
<emphasis id="B9608232F506941DFD3643DDDDC789B2" box="[522,659,404,428]" italics="true" pageId="11" pageNumber="12">A. clarivida</emphasis>
</taxonomicName>
(
<figureCitation id="132F42A5F506941DFD9B43DDDDAE89B2" box="[679,762,404,428]" captionStart="FIGURES 64 – 72" captionStartId="24.[151,264,1940,1963]" captionTargetBox="[250,1297,193,1910]" captionTargetId="figure@24.[248,1344,169,1924]" captionTargetPageId="24" captionText="FIGURES 64 – 72. Distribution maps for Adetomyrma species. 64, A. aureocuprea sp. nov.; 65, A. bressleri sp. nov.; 66, A. caputleae sp. nov.; 67, A. cassis sp. nov.; 68, A. caudapinniger sp. nov.; 69, A. cilium sp. nov.; 70, A. clarivida sp. nov.; 71, A. goblin sp. nov.; 72, A. venatrix. Star symbols represent the type locality." httpUri="https://zenodo.org/record/213833/files/figure.png" pageId="11" pageNumber="12">Fig 70</figureCitation>
). Separation between 
<taxonomicName id="4C1425A3F506941DFB3D43DDDBE689B2" box="[1025,1202,404,428]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aureocuprea">
<emphasis id="B9608232F506941DFB3D43DDDBE689B2" box="[1025,1202,404,428]" italics="true" pageId="11" pageNumber="12">A. aureocuprea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1425A3F506941DFBCF43DDDA2989B2" box="[1267,1405,404,428]" class="Insecta" family="Formicidae" genus="Adetomyrma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="clarivida">
<emphasis id="B9608232F506941DFBCF43DDDA2989B2" box="[1267,1405,404,428]" italics="true" pageId="11" pageNumber="12">A. clarivida</emphasis>
</taxonomicName>
is strongly supported by the morphological differences observed in the aedeagus (
<figureCitation id="132F42A5F506941DFB2C43FEDBEE89D1" box="[1040,1210,439,463]" captionStart="FIGURES 56 – 63" captionStartId="15.[151,264,1915,1938]" captionTargetBox="[217,1374,179,1882]" captionTargetId="figure@15.[217,1376,179,1893]" captionTargetPageId="15" captionText="FIGURES 56 – 63. Aedeagal plate of Adetomyrma males in lateral view. 56, A. aureocuprea sp. nov. (CASENT 0008678); 57, A. bressleri sp. nov. (CASENT 0218012); 58, A. caputleae sp. nov. (CASENT 0218011); 59, A. caudapinniger sp. nov. (CASENT 0111791); 60, A. cilium sp. nov. (CASENT 0218014); 61, A. clarivida sp. nov. (CASENT 0111802); 62, A. goblin sp. nov. (CASENT 0218010); 63, A. venatrix (CASENT 0063101)." httpUri="https://zenodo.org/record/213832/files/figure.png" pageId="11" pageNumber="12">Figs 56 and 61</figureCitation>
).
</paragraph>
<paragraph id="8BAB5E20F506941DFFF54395DB2B89EA" blockId="11.[151,1436,151,535]" box="[201,1151,476,500]" pageId="11" pageNumber="12">This new species corresponds to the following species code used in previous studies:</paragraph>
</subSubSection>
<subSubSection id="C30E0DABF506941DFFF543B6DDED8A09" box="[201,697,511,535]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BAB5E20F506941DFFF543B6DDED8A09" blockId="11.[151,1436,151,535]" box="[201,697,511,535]" pageId="11" pageNumber="12">
mgm05 (in part): 
<bibRefCitation id="EF8523D1F506941DFEB343B6DDE28A09" author="Yoshimura &amp; Fisher" box="[399,694,511,535]" firstAuthor="Yoshimura" pageId="11" pageNumber="12" pagination="33325" refId="ref20854" refString="Yoshimura, M. &amp; Fisher, B. L. (2012) A revision of male ants of the Malagasy Amblyoponinae (Hymenoptera: Formicidae) with resurrections of the genera Stigmatomma and Xymmer. PLoS ONE, 7, e 33325." type="journal article" year="2012">Yoshimura &amp; Fisher 2012</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>