420 lines
51 KiB
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420 lines
51 KiB
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<document id="9CEDD4EF1A78267CA8A4C9976FCEA6B3" ID-DOI="10.5281/zenodo.186240" ID-GBIF-Dataset="2f626f91-281e-48c5-9b52-7052663e307b" ID-ISSN="1175-5326" ID-Zenodo-Dep="186240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461048445079" checkinUser="plazi" docAuthor="Domínguez, Michel & Moreno, Luis V." docDate="2009" docId="03DE87B5FFD50D498BF928C158C8F9C8" docLanguage="en" docName="zt02028p066.pdf" docOrigin="Zootaxa 2028" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Typhlops golyathi Domínguez & Moreno, 2009, sp. nov." docType="treatment" docVersion="10" lastPageNumber="63" masterDocId="FFE7FFCDFFD70D4D8B6E28565A30FFF4" masterDocTitle="Taxonomy of the Cuban blind snakes (Scolecophidia, Typhlopidae), with the description of a new large species" masterLastPageNumber="66" masterPageNumber="59" pageNumber="61" updateTime="1698577881630" updateUser="plazi">
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<mods:titleInfo id="8688FBED2F56367AB49F9824FF9D272E">
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<mods:title id="7F156DE260030886BE570D5264144F37">Taxonomy of the Cuban blind snakes (Scolecophidia, Typhlopidae), with the description of a new large species</mods:title>
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<mods:namePart id="D595BAE7DBEBA853DC8B2449F0031215">Domínguez, Michel</mods:namePart>
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<mods:namePart id="A120613E173D3FD015639EBD3A701C7E">Moreno, Luis V.</mods:namePart>
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<mods:typeOfResource id="A62DD3F883A07FB44E66F0D770431556">text</mods:typeOfResource>
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<mods:title id="42301DEEEC0090B2BD471E455AB8B334">Zootaxa</mods:title>
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<mods:part id="776BDBB0809AED7CAABF9EB3D8BFF0E4">
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<mods:date id="ED427D677890AB8CE105F5ADA0F702DC">2009</mods:date>
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<mods:detail id="6FFD6EB0CD8836C96046DBBE6492167F" type="volume">
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<mods:number id="9A947C112E3CFD823B3BF15FC0EF8029">2028</mods:number>
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<mods:classification id="3760E97000F1540AA76C9030A0815217">journal article</mods:classification>
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<mods:identifier id="F6452A7BD7EB683A6678D47FFA2145BF" type="DOI">10.5281/zenodo.186240</mods:identifier>
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<mods:identifier id="B7D1C4283018B8CD76BBA247AFC40D0A" type="Zenodo-Dep">186240</mods:identifier>
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<treatment id="03DE87B5FFD50D498BF928C158C8F9C8" ID-DOI="http://doi.org/10.5281/zenodo.5629856" ID-GBIF-Taxon="119611375" ID-Zenodo-Dep="5629856" LSID="urn:lsid:plazi:treatment:03DE87B5FFD50D498BF928C158C8F9C8" httpUri="http://treatment.plazi.org/id/03DE87B5FFD50D498BF928C158C8F9C8" lastPageId="4" lastPageNumber="63" pageId="2" pageNumber="61">
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<subSubSection id="C36D6528FFD50D4F8BF928C15B96FF20" pageId="2" pageNumber="61" type="nomenclature">
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<paragraph id="8BC836A3FFD50D4F8BF928C15BE2FF45" blockId="2.[151,466,151,212]" box="[151,466,151,177]" pageId="2" pageNumber="61">
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<heading id="D08081CFFFD50D4F8BF928C15BE2FF45" bold="true" box="[151,466,151,177]" fontSize="11" level="1" pageId="2" pageNumber="61" reason="1">
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<emphasis id="B903EAB1FFD50D4F8BF928C15BE2FF45" bold="true" box="[151,466,151,177]" pageId="2" pageNumber="61">
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<taxonomicName id="4C774D20FFD50D4F8BF928C15B5DFF45" box="[151,365,151,177]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="golyathi" status="sp. nov.">
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<emphasis id="B903EAB1FFD50D4F8BF928C15B5DFF45" bold="true" box="[151,365,151,177]" italics="true" pageId="2" pageNumber="61">Typhlops golyathi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057CAFFD50D4F8A1B28C15BE2FF45" box="[373,466,151,177]" pageId="2" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836A3FFD50D4F8BF928EC5B96FF20" blockId="2.[151,466,151,212]" box="[151,422,186,212]" pageId="2" pageNumber="61">
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(
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<tableCitation id="C6F50318FFD50D4F8BF128EC5B33FF20" box="[159,259,186,212]" captionStart="TABLE 1" captionStartId="2.[151,240,766,790]" captionTargetBox="[151,1437,964,1347]" captionTargetPageId="2" captionText="TABLE 1. Comparison among eight Cuban Typhlops species of T. biminiensis species group using selected characters: A, T. golyathi; B, T. anchaurus; C, T. anousius; D, T. arator; E, T. contorhinus; F, T. notorachius; G, T. perimychus; H, T. satelles. Data for species B to G were obtained from Thomas and Hedges (2007), and for species C from Domínguez and Moreno (in press)." httpUri="http://table.plazi.org/id/DF08662BFFD50D4F8BF92AA85B6DFC89" pageId="2" pageNumber="61" tableUuid="DF08662BFFD50D4F8BF92AA85B6DFC89">Tables 1</tableCitation>
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,
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<tableCitation id="C6F50318FFD50D4F8A7F28EC5B12FF20" box="[273,290,186,212]" captionStart="TABLE 2" captionStartId="3.[151,239,936,960]" captionTargetBox="[151,1436,1022,1629]" captionTargetPageId="3" captionText="TABLE 2. Comparison between Typhlops golyathi and Typhlops arator. Data for T. arator were obtained from Thomas" httpUri="http://table.plazi.org/id/DF08662BFFD40D4E8BF92BFE5FACFC34" pageId="2" pageNumber="61" tableUuid="DF08662BFFD40D4E8BF92BFE5FACFC34">2</tableCitation>
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;
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<figureCitation id="134C2A26FFD50D4F8A4128EC5B4FFF20" box="[303,383,186,212]" captionStart="FIGURE 1" captionStartId="4.[151,255,1264,1288]" captionTargetBox="[475,1112,219,1222]" captionTargetId="figure@4.[472,1115,197,1239]" captionTargetPageId="4" captionText="FIGURE 1. Typhlops golyathi (holotype) dorsal and lateral views, showing the head scutellation (bar = 2 mm)." httpUri="https://zenodo.org/record/186241/files/figure.png" pageId="2" pageNumber="61">Figs. 1</figureCitation>
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,
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<figureCitation id="134C2A26FFD50D4F8AE028EC5BAEFF20" box="[398,414,186,212]" captionStart="FIGURE 2" captionStartId="5.[151,256,1312,1336]" captionTargetBox="[158,1424,521,1280]" captionTargetId="figure@5.[151,1436,514,1288]" captionTargetPageId="5" captionText="FIGURE 2. Geographic distribution of the Typhlops biminiensis species group in Cuba. Closed star = T. golyathi sp. nov.; closed square = T. anchaurus; mixed square = T. anousius; closed rhombus = T. arator; mixed circle = T. contorhinus; closed non – inverse triangles = T. notorachius; closed circles = T. perimychus; closed inverse triangle = T. satelles." httpUri="https://zenodo.org/record/186242/files/figure.png" pageId="2" pageNumber="61">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6528FFD50D4F8BF929515EDDFEBC" pageId="2" pageNumber="61" type="materials_examined">
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<paragraph id="8BC836A3FFD50D4F8BF929515EDDFEBC" blockId="2.[151,1436,263,715]" pageId="2" pageNumber="61">
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<emphasis id="B903EAB1FFD50D4F8BF929515B23FED5" bold="true" box="[151,275,263,289]" pageId="2" pageNumber="61">
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<typeStatus id="54CC8801FFD50D4F8BF929515B3EFED5" box="[151,270,263,289]" pageId="2" pageNumber="61" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male
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<collectionCode id="ED66AE66FFD50D4F8A0F295E5BF5FED6" box="[353,453,264,290]" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="2" pageNumber="61">CZACC</collectionCode>
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4.5385 from Valle de
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<collectingCountry id="F3607633FFD50D4F89BA295E5959FED6" box="[724,873,264,290]" name="Saint Vincent and the Grenadines" pageId="2" pageNumber="61">San Vicente</collectingCountry>
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, Viñales Municipality, Pinar del Río Province,
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<collectingCountry id="F3607633FFD50D4F8BF929785AE8FEBC" box="[151,216,302,328]" name="Cuba" pageId="2" pageNumber="61">Cuba</collectingCountry>
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,
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<geoCoordinate id="EE435064FFD50D4F8B8D29785B44FEBC" box="[227,372,302,328]" direction="north" orientation="latitude" pageId="2" pageNumber="61" precision="15" value="83.71667">83º43'00'' N</geoCoordinate>
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,
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<geoCoordinate id="EE435064FFD50D4F8A1129785823FEBC" box="[383,531,302,328]" direction="west" orientation="longitude" pageId="2" pageNumber="61" precision="15" value="-22.683332">22º41'00'' W</geoCoordinate>
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,
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<quantity id="4C8F9B46FFD50D4F897129785858FEBC" box="[543,616,302,328]" metricMagnitude="2" metricUnit="m" metricValue="1.1199999999999999" pageId="2" pageNumber="61" unit="m" value="112.0">112 m</quantity>
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elevation, collected on
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<date id="FFC91063FFD50D4F8811297859CFFEBC" box="[895,1023,302,328]" pageId="2" pageNumber="61" value="2003-04">April 2003</date>
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by Roberto Alonso.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6528FFD50D4F8BA829035867FC44" pageId="2" pageNumber="61" type="diagnosis">
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<paragraph id="8BC836A3FFD50D4F8BA829035B01FD3F" blockId="2.[151,1436,263,715]" pageId="2" pageNumber="61">
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<emphasis id="B903EAB1FFD50D4F8BA829035B76FE9B" bold="true" box="[198,326,341,367]" pageId="2" pageNumber="61">Diagnosis.</emphasis>
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A large species (
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<quantity id="4C8F9B46FFD50D4F897D29035847FE9B" box="[531,631,341,367]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="61" unit="mm" value="360.0">360 mm</quantity>
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maximum SVL) with moderate body size (57.8 TL/MBD) of the
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<taxonomicName id="4C774D20FFD50D4F8EE829035B11FE61" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="B903EAB1FFD50D4F8EE829035B11FE61" italics="true" pageId="2" pageNumber="61">T. biminiensis</emphasis>
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</taxonomicName>
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species group that presents more middorsal scales (> 625) than any other West Indian species (
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<tableCitation id="C6F50318FFD50D4F8BF129F45AC9FE48" box="[159,249,418,444]" captionStart="TABLE 1" captionStartId="2.[151,240,766,790]" captionTargetBox="[151,1437,964,1347]" captionTargetPageId="2" captionText="TABLE 1. Comparison among eight Cuban Typhlops species of T. biminiensis species group using selected characters: A, T. golyathi; B, T. anchaurus; C, T. anousius; D, T. arator; E, T. contorhinus; F, T. notorachius; G, T. perimychus; H, T. satelles. Data for species B to G were obtained from Thomas and Hedges (2007), and for species C from Domínguez and Moreno (in press)." httpUri="http://table.plazi.org/id/DF08662BFFD50D4F8BF92AA85B6DFC89" pageId="2" pageNumber="61" tableUuid="DF08662BFFD50D4F8BF92AA85B6DFC89">Table 1</tableCitation>
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). It also presents a posterior reduction of four longitudinal scale rows, this characteristic is unique among species of the group. The closest species is
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<taxonomicName id="4C774D20FFD50D4F8981299F5965FE16" box="[751,853,457,482]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="arator">
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<emphasis id="B903EAB1FFD50D4F8981299F5965FE16" box="[751,853,457,482]" italics="true" pageId="2" pageNumber="61">T. arator</emphasis>
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</taxonomicName>
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. Both species are large Cuban blind snakes, have the same number of scale rows at the anterior part of the body (26) and a greater number of middorsal scales than any other West Indian species (
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<tableCitation id="C6F50318FFD50D4F892B2A4058AEFDC4" box="[581,670,534,560]" captionStart="TABLE 1" captionStartId="2.[151,240,766,790]" captionTargetBox="[151,1437,964,1347]" captionTargetPageId="2" captionText="TABLE 1. Comparison among eight Cuban Typhlops species of T. biminiensis species group using selected characters: A, T. golyathi; B, T. anchaurus; C, T. anousius; D, T. arator; E, T. contorhinus; F, T. notorachius; G, T. perimychus; H, T. satelles. Data for species B to G were obtained from Thomas and Hedges (2007), and for species C from Domínguez and Moreno (in press)." httpUri="http://table.plazi.org/id/DF08662BFFD50D4F8BF92AA85B6DFC89" pageId="2" pageNumber="61" tableUuid="DF08662BFFD50D4F8BF92AA85B6DFC89">Table 1</tableCitation>
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). Also, both species are distributed to the west of the remaining species of the
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<taxonomicName id="4C774D20FFD50D4F8A2D2A6B5BD9FDA2" box="[323,489,573,598]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="B903EAB1FFD50D4F8A2D2A6B5BD9FDA2" box="[323,489,573,598]" italics="true" pageId="2" pageNumber="61">T. biminiensis</emphasis>
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</taxonomicName>
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species group (
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<figureCitation id="134C2A26FFD50D4F89C82A6B58C0FDA3" box="[678,752,573,599]" captionStart="FIGURE 2" captionStartId="5.[151,256,1312,1336]" captionTargetBox="[158,1424,521,1280]" captionTargetId="figure@5.[151,1436,514,1288]" captionTargetPageId="5" captionText="FIGURE 2. Geographic distribution of the Typhlops biminiensis species group in Cuba. Closed star = T. golyathi sp. nov.; closed square = T. anchaurus; mixed square = T. anousius; closed rhombus = T. arator; mixed circle = T. contorhinus; closed non – inverse triangles = T. notorachius; closed circles = T. perimychus; closed inverse triangle = T. satelles." httpUri="https://zenodo.org/record/186242/files/figure.png" pageId="2" pageNumber="61">Fig. 2</figureCitation>
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). However, the new species can be distinguished from
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<taxonomicName id="4C774D20FFD50D4F8EE82A6B5AD2FD89" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="arator">
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<emphasis id="B903EAB1FFD50D4F8EE82A6B5AD2FD89" italics="true" pageId="2" pageNumber="61">T. arator</emphasis>
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</taxonomicName>
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by its posterior scale row reduction (4 rows vs. 2 rows), total middorsals (> 625 vs. <580), relative tail length (3% vs. 2%), snout and rostral patterns, ocular size (OLM/OW 0.8 vs. 0.5) and sinuosity (0.8 vs. 0.2) (
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<tableCitation id="C6F50318FFD50D4F8BF12AE75B33FD3F" box="[159,259,689,715]" captionStart="TABLE 1" captionStartId="2.[151,240,766,790]" captionTargetBox="[151,1437,964,1347]" captionTargetPageId="2" captionText="TABLE 1. Comparison among eight Cuban Typhlops species of T. biminiensis species group using selected characters: A, T. golyathi; B, T. anchaurus; C, T. anousius; D, T. arator; E, T. contorhinus; F, T. notorachius; G, T. perimychus; H, T. satelles. Data for species B to G were obtained from Thomas and Hedges (2007), and for species C from Domínguez and Moreno (in press)." httpUri="http://table.plazi.org/id/DF08662BFFD50D4F8BF92AA85B6DFC89" pageId="2" pageNumber="61" tableUuid="DF08662BFFD50D4F8BF92AA85B6DFC89">Tables 1</tableCitation>
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,
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<tableCitation id="C6F50318FFD50D4F8A7F2AE75B12FD3F" box="[273,290,689,715]" captionStart="TABLE 2" captionStartId="3.[151,239,936,960]" captionTargetBox="[151,1436,1022,1629]" captionTargetPageId="3" captionText="TABLE 2. Comparison between Typhlops golyathi and Typhlops arator. Data for T. arator were obtained from Thomas" httpUri="http://table.plazi.org/id/DF08662BFFD40D4E8BF92BFE5FACFC34" pageId="2" pageNumber="61" tableUuid="DF08662BFFD40D4E8BF92BFE5FACFC34">2</tableCitation>
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).
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</paragraph>
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<caption id="DF08662BFFD50D4F8BF92AA85B6DFC89" ID-Table-UUID="DF08662BFFD50D4F8BF92AA85B6DFC89" httpUri="http://table.plazi.org/id/DF08662BFFD50D4F8BF92AA85B6DFC89" pageId="2" pageNumber="61" targetBox="[151,1437,964,1347]" targetIsTable="true" targetPageId="2">
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<paragraph id="8BC836A3FFD50D4F8BF92AA85B6DFC89" blockId="2.[151,1437,766,944]" pageId="2" pageNumber="61">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8BF92AA85B34FCE2" bold="true" box="[151,260,766,790]" pageId="2" pageNumber="61">TABLE 1</emphasis>
|
|||
|
|
. Comparison among eight Cuban
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F89192AA858E7FCE1" box="[631,727,766,789]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F89192AA858E7FCE1" box="[631,727,766,789]" italics="true" pageId="2" pageNumber="61">Typhlops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species of
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F883F2AA859D6FCE1" box="[849,998,766,789]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="biminiensis">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F883F2AA859D6FCE1" box="[849,998,766,789]" italics="true" pageId="2" pageNumber="61">T. biminiensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group using selected characters: A,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F8BD92B775B19FCCC" box="[183,297,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="golyathi">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8BD92B775B19FCCC" box="[183,297,801,824]" italics="true" pageId="2" pageNumber="61">T. golyathi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; B,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F8A382B775BD1FCCC" box="[342,481,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="anchaurus">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8A382B775BD1FCCC" box="[342,481,801,824]" italics="true" pageId="2" pageNumber="61">T. anchaurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; C,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F89602B7758B7FCCC" box="[526,647,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="anousius">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F89602B7758B7FCCC" box="[526,647,801,824]" italics="true" pageId="2" pageNumber="61">T. anousius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; D,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F89DB2B775923FCCC" box="[693,787,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="arator">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F89DB2B775923FCCC" box="[693,787,801,824]" italics="true" pageId="2" pageNumber="61">T. arator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; E,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F88512B7759E8FCCC" box="[831,984,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="contorhinus">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F88512B7759E8FCCC" box="[831,984,801,824]" italics="true" pageId="2" pageNumber="61">T. contorhinus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; F,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F8F6E2B775EA9FCCC" box="[1024,1177,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="notorachius">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8F6E2B775EA9FCCC" box="[1024,1177,801,824]" italics="true" pageId="2" pageNumber="61">T. notorachius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; G,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F8FAB2B775F69FCCC" box="[1221,1369,801,824]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="perimychus">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8FAB2B775F69FCCC" box="[1221,1369,801,824]" italics="true" pageId="2" pageNumber="61">T. perimychus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; H,
|
|||
|
|
<taxonomicName id="4C774D20FFD50D4F8EE62B775AD7FCAF" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="satelles">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8EE62B775AD7FCAF" italics="true" pageId="2" pageNumber="61">T. satelles</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Data for species B to G were obtained from Thomas and Hedges (2007), and for species C from Domínguez and Moreno (in press).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC836A3FFD50D4F8BF12BCF5867FC44" blockId="2.[151,1437,766,944]" box="[159,599,921,944]" pageId="2" pageNumber="61">Character Species</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<paragraph id="8BC836A3FFD50D4F8BF12BB85F1FFAB7" pageId="2" pageNumber="61">
|
|||
|
|
<table id="F977C403FFD5F2B28BF92B925FADFAB7" box="[151,1437,964,1347]" gridcols="9" gridrows="8" pageId="2" pageNumber="61">
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92B925FADFBF1" box="[151,1437,964,1029]" gridrow="0" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92B925BE4FBF1" box="[151,468,964,1029]" gridcol="0" gridrow="0" pageId="2" pageNumber="61">Maximum TL (mm)</th>
|
|||
|
|
<th id="76965D9DFFD5F2B289682B92581EFBF1" box="[518,558,964,1029]" gridcol="1" gridrow="0" pageId="2" pageNumber="61">A 371</th>
|
|||
|
|
<th id="76965D9DFFD5F2B289352B9258B3FBF1" box="[603,643,964,1029]" gridcol="2" gridrow="0" pageId="2" pageNumber="61">B 240</th>
|
|||
|
|
<th id="76965D9DFFD5F2B289DE2B92593EFBF1" box="[688,782,964,1029]" gridcol="3" gridrow="0" pageId="2" pageNumber="61">C 278</th>
|
|||
|
|
<th id="76965D9DFFD5F2B2885F2B9259BEFBF1" box="[817,910,964,1029]" gridcol="4" gridrow="0" pageId="2" pageNumber="61">D 460</th>
|
|||
|
|
<th id="76965D9DFFD5F2B288DC2B9259EAFBF1" box="[946,986,964,1029]" gridcol="5" gridrow="0" pageId="2" pageNumber="61">E 316</th>
|
|||
|
|
<th id="76965D9DFFD5F2B28F792B925E45FBF1" box="[1047,1141,964,1029]" gridcol="6" gridrow="0" pageId="2" pageNumber="61">F 301</th>
|
|||
|
|
<th id="76965D9DFFD5F2B28FF62B925EC5FBF1" box="[1176,1269,964,1029]" gridcol="7" gridrow="0" pageId="2" pageNumber="61">G 280</th>
|
|||
|
|
<th id="76965D9DFFD5F2B28E782B925FADFBF1" box="[1302,1437,964,1029]" gridcol="8" gridrow="0" pageId="2" pageNumber="61">H 350</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92C4A5FADFBC7" box="[151,1437,1052,1075]" gridrow="1" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92C4A5BE4FBC7" box="[151,468,1052,1075]" gridcol="0" gridrow="1" pageId="2" pageNumber="61">Maximum TA (mm)</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682C4A581EFBC7" box="[518,558,1052,1075]" gridcol="1" gridrow="1" pageId="2" pageNumber="61">11</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352C4A58B3FBC7" box="[603,643,1052,1075]" gridcol="2" gridrow="1" pageId="2" pageNumber="61">5</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2C4A593EFBC7" box="[688,782,1052,1075]" gridcol="3" gridrow="1" pageId="2" pageNumber="61">5</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2C4A59BEFBC7" box="[817,910,1052,1075]" gridcol="4" gridrow="1" pageId="2" pageNumber="61">10</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2C4A59EAFBC7" box="[946,986,1052,1075]" gridcol="5" gridrow="1" pageId="2" pageNumber="61">6</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792C4A5E45FBC7" box="[1047,1141,1052,1075]" gridcol="6" gridrow="1" pageId="2" pageNumber="61">5</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62C4A5EC5FBC7" box="[1176,1269,1052,1075]" gridcol="7" gridrow="1" pageId="2" pageNumber="61">6</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782C4A5FADFBC7" box="[1302,1437,1052,1075]" gridcol="8" gridrow="1" pageId="2" pageNumber="61">7</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92C1F5FADFB94" box="[151,1437,1097,1120]" gridrow="2" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92C1F5BE4FB94" box="[151,468,1097,1120]" gridcol="0" gridrow="2" pageId="2" pageNumber="61">TL/TA</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682C1F581EFB94" box="[518,558,1097,1120]" gridcol="1" gridrow="2" pageId="2" pageNumber="61">34</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352C1F58B3FB94" box="[603,643,1097,1120]" gridcol="2" gridrow="2" pageId="2" pageNumber="61">48</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2C1F593EFB94" box="[688,782,1097,1120]" gridcol="3" gridrow="2" pageId="2" pageNumber="61">55–56</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2C1F59BEFB94" box="[817,910,1097,1120]" gridcol="4" gridrow="2" pageId="2" pageNumber="61">46–48</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2C1F59EAFB94" box="[946,986,1097,1120]" gridcol="5" gridrow="2" pageId="2" pageNumber="61">53</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792C1F5E45FB94" box="[1047,1141,1097,1120]" gridcol="6" gridrow="2" pageId="2" pageNumber="61">48–75</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62C1F5EC5FB94" box="[1176,1269,1097,1120]" gridcol="7" gridrow="2" pageId="2" pageNumber="61">46–85</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782C1F5FADFB94" box="[1302,1437,1097,1120]" gridcol="8" gridrow="2" pageId="2" pageNumber="61">55–61</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92C205FADFB79" box="[151,1437,1142,1165]" gridrow="3" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92C205BE4FB79" box="[151,468,1142,1165]" gridcol="0" gridrow="3" pageId="2" pageNumber="61">TL/MBD</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682C20581EFB79" box="[518,558,1142,1165]" gridcol="1" gridrow="3" pageId="2" pageNumber="61">58</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352C2058B3FB79" box="[603,643,1142,1165]" gridcol="2" gridrow="3" pageId="2" pageNumber="61">63</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2C20593EFB79" box="[688,782,1142,1165]" gridcol="3" gridrow="3" pageId="2" pageNumber="61">45–62</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2C2059BEFB79" box="[817,910,1142,1165]" gridcol="4" gridrow="3" pageId="2" pageNumber="61">51–58</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2C2059EAFB79" box="[946,986,1142,1165]" gridcol="5" gridrow="3" pageId="2" pageNumber="61">63</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792C205E45FB79" box="[1047,1141,1142,1165]" gridcol="6" gridrow="3" pageId="2" pageNumber="61">45–57</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62C205EC5FB79" box="[1176,1269,1142,1165]" gridcol="7" gridrow="3" pageId="2" pageNumber="61">41–59</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782C205FADFB79" box="[1302,1437,1142,1165]" gridcol="8" gridrow="3" pageId="2" pageNumber="61">62–75</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92CF25FADFB4F" box="[151,1437,1188,1211]" gridrow="4" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92CF25BE4FB4F" box="[151,468,1188,1211]" gridcol="0" gridrow="4" pageId="2" pageNumber="61">Middorsal scale counts</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682CF2581EFB4F" box="[518,558,1188,1211]" gridcol="1" gridrow="4" pageId="2" pageNumber="61">629</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352CF258B3FB4F" box="[603,643,1188,1211]" gridcol="2" gridrow="4" pageId="2" pageNumber="61">514</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2CF2593EFB4F" box="[688,782,1188,1211]" gridcol="3" gridrow="4" pageId="2" pageNumber="61">460–513</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2CF259BEFB4F" box="[817,910,1188,1211]" gridcol="4" gridrow="4" pageId="2" pageNumber="61">578–579</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2CF259EAFB4F" box="[946,986,1188,1211]" gridcol="5" gridrow="4" pageId="2" pageNumber="61">502</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792CF25E45FB4F" box="[1047,1141,1188,1211]" gridcol="6" gridrow="4" pageId="2" pageNumber="61">475–529</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62CF25EC5FB4F" box="[1176,1269,1188,1211]" gridcol="7" gridrow="4" pageId="2" pageNumber="61">453–496</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782CF25FADFB4F" box="[1302,1437,1188,1211]" gridcol="8" gridrow="4" pageId="2" pageNumber="61">514–527</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92C875FADFB1C" box="[151,1437,1233,1256]" gridrow="5" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92C875BE4FB1C" box="[151,468,1233,1256]" gridcol="0" gridrow="5" pageId="2" pageNumber="61">Anterior scale rows</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682C87581EFB1C" box="[518,558,1233,1256]" gridcol="1" gridrow="5" pageId="2" pageNumber="61">26</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352C8758B3FB1C" box="[603,643,1233,1256]" gridcol="2" gridrow="5" pageId="2" pageNumber="61">24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2C87593EFB1C" box="[688,782,1233,1256]" gridcol="3" gridrow="5" pageId="2" pageNumber="61">24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2C8759BEFB1C" box="[817,910,1233,1256]" gridcol="4" gridrow="5" pageId="2" pageNumber="61">26</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2C8759EAFB1C" box="[946,986,1233,1256]" gridcol="5" gridrow="5" pageId="2" pageNumber="61">24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792C875E45FB1C" box="[1047,1141,1233,1256]" gridcol="6" gridrow="5" pageId="2" pageNumber="61">24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62C875EC5FB1C" box="[1176,1269,1233,1256]" gridcol="7" gridrow="5" pageId="2" pageNumber="61">22, 24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782C875FADFB1C" box="[1302,1437,1233,1256]" gridcol="8" gridrow="5" pageId="2" pageNumber="61">22, 24</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92CA85FADFAE1" box="[151,1437,1278,1301]" gridrow="6" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92CA85BE4FAE1" box="[151,468,1278,1301]" gridcol="0" gridrow="6" pageId="2" pageNumber="61">Posterior scale rows</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682CA8581EFAE1" box="[518,558,1278,1301]" gridcol="1" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352CA858B3FAE1" box="[603,643,1278,1301]" gridcol="2" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2CA8593EFAE1" box="[688,782,1278,1301]" gridcol="3" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2CA859BEFAE1" box="[817,910,1278,1301]" gridcol="4" gridrow="6" pageId="2" pageNumber="61">24</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2CA859EAFAE1" box="[946,986,1278,1301]" gridcol="5" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792CA85E45FAE1" box="[1047,1141,1278,1301]" gridcol="6" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62CA85EC5FAE1" box="[1176,1269,1278,1301]" gridcol="7" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782CA85FADFAE1" box="[1302,1437,1278,1301]" gridcol="8" gridrow="6" pageId="2" pageNumber="61">22</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD5F2B28BF92D7A5FADFAB7" box="[151,1437,1324,1347]" gridrow="7" pageId="2" pageNumber="61">
|
|||
|
|
<th id="76965D9DFFD5F2B28BF92D7A5BE4FAB7" box="[151,468,1324,1347]" gridcol="0" gridrow="7" pageId="2" pageNumber="61">Scale rows reduction (% TL)</th>
|
|||
|
|
<td id="76965D9DFFD5F2B289682D7A581EFAB7" box="[518,558,1324,1347]" gridcol="1" gridrow="7" pageId="2" pageNumber="61">42</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289352D7A58B3FAB7" box="[603,643,1324,1347]" gridcol="2" gridrow="7" pageId="2" pageNumber="61">31</td>
|
|||
|
|
<td id="76965D9DFFD5F2B289DE2D7A593EFAB7" box="[688,782,1324,1347]" gridcol="3" gridrow="7" pageId="2" pageNumber="61">2–8</td>
|
|||
|
|
<td id="76965D9DFFD5F2B2885F2D7A59BEFAB7" box="[817,910,1324,1347]" gridcol="4" gridrow="7" pageId="2" pageNumber="61">34–41</td>
|
|||
|
|
<td id="76965D9DFFD5F2B288DC2D7A59EAFAB7" box="[946,986,1324,1347]" gridcol="5" gridrow="7" pageId="2" pageNumber="61">31</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28F792D7A5E45FAB7" box="[1047,1141,1324,1347]" gridcol="6" gridrow="7" pageId="2" pageNumber="61">15–40</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28FF62D7A5EC5FAB7" box="[1176,1269,1324,1347]" gridcol="7" gridrow="7" pageId="2" pageNumber="61">50</td>
|
|||
|
|
<td id="76965D9DFFD5F2B28E782D7A5FADFAB7" box="[1302,1437,1324,1347]" gridcol="8" gridrow="7" pageId="2" pageNumber="61">11</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<subSubSection id="C36D6528FFD50D4E8BAB2DD958EDF8CC" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="description">
|
|||
|
|
<paragraph id="8BC836A3FFD50D4E8BAB2DD95E1DFC81" blockId="2.[151,1437,1423,2003]" lastBlockId="3.[151,1437,152,885]" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
|
|||
|
|
<emphasis id="B903EAB1FFD50D4F8BAB2DD95B6CFA5D" bold="true" box="[197,348,1423,1449]" pageId="2" pageNumber="61">Description.</emphasis>
|
|||
|
|
Male with large size (
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F890B2DC658F8FA5E" box="[613,712,1424,1450]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="61" unit="mm" value="360.0">360 mm</quantity>
|
|||
|
|
SVL),
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F884E2DC65943FA5E" box="[800,883,1424,1450]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="61" unit="mm" value="11.0">11 mm</quantity>
|
|||
|
|
TA,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F88C02DC65E20FA5E" box="[942,1040,1424,1450]" metricMagnitude="-1" metricUnit="m" metricValue="3.71" pageId="2" pageNumber="61" unit="mm" value="371.0">371 mm</quantity>
|
|||
|
|
TL, 33.7 TL/TA, moderate body,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F8BF92DE05AC0FA24" box="[151,240,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="61" unit="mm" value="4.2">4.2 mm</quantity>
|
|||
|
|
ABD,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F8A2C2DE05BACFA24" box="[322,412,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="61" unit="mm" value="6.4">6.4 mm</quantity>
|
|||
|
|
MBD,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F8A9D2DE0587CFA24" box="[499,588,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="61" unit="mm" value="5.3">5.3 mm</quantity>
|
|||
|
|
PBD,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F89F42DE058C3FA24" box="[666,755,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="61" unit="mm" value="3.7">3.7 mm</quantity>
|
|||
|
|
TD, 88.3 TL/ABD, 58.0 TL/MBD, 70.0 TL/PBD, 3.0 TA/ TD. Tail with a small apical spine and three scales bordering the vent opening. Head non–distinct from neck and ogival in dorsal view, broader than long (1.06 HWM/HL), narrowing to the apex, 0.89 HWE/HWM and 0.52 HWN/HWM, strongly dorso–ventrally depressed. Snout lightly sharp–pointed, narrow (0.50
|
|||
|
|
<collectingCountry id="F3607633FFD50D4F8E332E7C5FA3F9B0" box="[1373,1427,1578,1604]" name="India" pageId="2" pageNumber="61">IND</collectingCountry>
|
|||
|
|
/ HWM) and ogival to rounded head in outline. Rostral broad in dorsal view, broader than long (1.03 RWD1/ RLD2, 0.92 RWD1/RLD1), curved–sided (1.01 RWD2/RWD1), lightly umboed, not flared on anterior apex, nearly parallel–sided midrostrally, with sharp–pointed to rounded posterior apex, almost reaching interocular level,
|
|||
|
|
<quantity id="4C8F9B46FFD50D4F8B8C2E9D5B7DF911" box="[226,333,1739,1765]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="61" unit="mm" value="0.08">0.08 mm</quantity>
|
|||
|
|
RED (0.04 RED/RLD2); broad rostral in ventral view, broader than long, without anterior constriction, anteriorly equally as broad as in dorsal view, obliquely–sided with strong posterior constriction near the mouth, 1.90 ((RWD1 + RTL)/TL)*100. Nasal completely divided; subtriangular superior nasal, with shallow concavity in posterior margin and narrowly in contact with anterior margin of the supraocular; broad nostrils, not visible in dorsal view and much closer to rostral margin than preocular margin; short superior nasal suture and nostrils obliquely oriented (45º angle approximately) to the body longitudinal axis in lateral view; inferior nasal suture with a pronounced curve and contacting in the midpoint of second supralabial; nasal–preocular suture coincident with second–third supralabials suture; preocular–ocular suture coincident with third–fourth supralabials suture; postnasal pattern divergent (0.61 PPNW/APNW, 0.63 RWD1/APNW). Single preocular with "S" pattern, twice as wide as long (0.51 RC/PW), smaller than superior nasal and ocular, in contact with second and third supralabials (
|
|||
|
|
<quantity id="4C8F9B46FFD40D4E89DB29595910FEDD" box="[693,800,271,297]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="62" unit="mm" value="0.28">0.28 mm</quantity>
|
|||
|
|
PSL), sinuous preocular–ocular suture (0.72 PS, 0.79 OS). Polygonal ocular and longer than wide (0.72 OLM/OW); black eyes,
|
|||
|
|
<quantity id="4C8F9B46FFD40D4E8F7729635EB4FEBB" box="[1049,1156,309,335]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="62" unit="mm" value="0.28">0.28 mm</quantity>
|
|||
|
|
ED, at preocular notch level,
|
|||
|
|
<quantity id="4C8F9B46FFD40D4E8BB0290A5B76FE82" box="[222,326,348,374]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="3" pageNumber="62" unit="mm" value="2.63">2.63 mm</quantity>
|
|||
|
|
IOD, and distant from the snout, 0.50 SED/HL; postocular located between parietal and fourth supralabial, with straight posterior margin. Frontal similar to supraocular, both slightly longer than postfrontal, interparietal and interoccipital; nuchal similar to the costals; supraocular obliquely oriented (45º angle approximately) to the head midline; two parietals extending towards the head midline and two occipitals nearly transversely oriented to body axis and parietals. Scale posterior borders are sharp–pointed to rounded in frontal, supraocular, interparietal, interoccipital and nuchal; lightly truncated in postfrontal and parietals, and lightly rounded in occipitals. Four supralabials with T–III imbrication pattern; second supralabial 1.5 times the size of first, in contact with nasal superior and lower edge of preocular; third supralabial twice the size of the first, in contact with preocular and lower edge of ocular; fourth supralabial is the largest, more than four times the first, with convex posterior margin. Four infralabials; second lightly larger than first; third twice times the first; fourth infralabial is the largest, 2.5 times the first. Middorsal scale count, 629; 26 scale rows, reducing posteriorly to: 25 scale rows at 29 % TL or 231th middorsal scales level, 24 scale rows at 30 % TL or 235th middorsal scales level, 23 scale rows at 39 % TL or 285th middorsal scales level, and finally to 22 at 42 % TL or 298th middorsal scales level. Dorsocaudals, 20 and ventrocaudals, 22.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF08662BFFD40D4E8BF92BFE5FACFC34" ID-Table-UUID="DF08662BFFD40D4E8BF92BFE5FACFC34" box="[151,1436,936,960]" httpUri="http://table.plazi.org/id/DF08662BFFD40D4E8BF92BFE5FACFC34" pageId="3" pageNumber="62" targetBox="[151,1436,1022,1629]" targetIsTable="true" targetPageId="3">
|
|||
|
|
<paragraph id="8BC836A3FFD40D4E8BF92BFE5FACFC34" blockId="3.[151,1436,936,994]" box="[151,1436,936,960]" pageId="3" pageNumber="62">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8BF92BFE5B32FC34" bold="true" box="[151,258,936,960]" pageId="3" pageNumber="62">TABLE 2</emphasis>
|
|||
|
|
. Comparison between
|
|||
|
|
<taxonomicName id="4C774D20FFD40D4E8A922BFF588AFC34" box="[508,698,937,960]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="golyathi">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8A922BFF588AFC34" box="[508,698,937,960]" italics="true" pageId="3" pageNumber="62">Typhlops golyathi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C774D20FFD40D4E89812BFF59A9FC34" box="[751,921,937,960]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="arator">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E89812BFF59A9FC34" box="[751,921,937,960]" italics="true" pageId="3" pageNumber="62">Typhlops arator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Data for
|
|||
|
|
<taxonomicName id="4C774D20FFD40D4E8F682BFF5E55FC34" box="[1030,1125,937,960]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="arator">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8F682BFF5E55FC34" box="[1030,1125,937,960]" italics="true" pageId="3" pageNumber="62">T. arator</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were obtained from Thomas
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC836A3FFD40D4E8BF92B9D5B59FC16" blockId="3.[151,1436,936,994]" box="[151,361,971,994]" pageId="3" pageNumber="62">
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFE64B52FFD40D4E8BAA2B9D5B55FC16" author="Hedges" box="[196,357,971,994]" pageId="3" pageNumber="65" refString="Thomas, R. & Hedges, S. B. (2007) Eleven new species of snakes of the Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba. Z ootaxa, 1400, 1 - 26." type="journal article" year="2007">Hedges (2007)</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC836A3FFD40D4E8BF12BA85E5BF9A9" pageId="3" pageNumber="62">
|
|||
|
|
<table id="F977C403FFD4F2B28BF92BA85FACF9A9" box="[151,1436,1022,1629]" gridcols="3" gridrows="9" pageId="3" pageNumber="62">
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92BA85FACFB9E" box="[151,1436,1022,1130]" gridrow="0" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92BA85BEAFB9E" box="[151,474,1022,1130]" gridcol="0" gridrow="0" pageId="3" pageNumber="62">Character Dorsal snout pattern</th>
|
|||
|
|
<th id="76965D9DFFD4F2B289BA2BA859A7FB9E" box="[724,919,1022,1130]" gridcol="1" gridrow="0" pageId="3" pageNumber="62">
|
|||
|
|
Species
|
|||
|
|
<taxonomicName id="4C774D20FFD40D4E89BA2C7F592DFB9E" authority="Ogival" authorityName="Ogival" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="golyathi">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E89BA2C7F5975FBB4" box="[724,837,1065,1088]" italics="true" pageId="3" pageNumber="62">T. golyathi</emphasis>
|
|||
|
|
Ogival
|
|||
|
|
</taxonomicName>
|
|||
|
|
to rounded
|
|||
|
|
</th>
|
|||
|
|
<th id="76965D9DFFD4F2B28F522BA85FACFB9E" box="[1084,1436,1022,1130]" gridcol="2" gridrow="0" pageId="3" pageNumber="62">
|
|||
|
|
<taxonomicName id="4C774D20FFD40D4E8F522C7F5EB5FB9E" authority="Ogival" authorityName="Ogival" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="arator">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8F522C7F5EAAFBB4" box="[1084,1178,1065,1088]" italics="true" pageId="3" pageNumber="62">T. arator</emphasis>
|
|||
|
|
Ogival
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92CD75FACFB6C" box="[151,1436,1153,1176]" gridrow="1" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92CD75BEAFB6C" box="[151,474,1153,1176]" gridcol="0" gridrow="1" pageId="3" pageNumber="62">Lateral snout pattern</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2CD759A7FB6C" box="[724,919,1153,1176]" gridcol="1" gridrow="1" pageId="3" pageNumber="62">Pointed</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522CD75FACFB6C" box="[1084,1436,1153,1176]" gridcol="2" gridrow="1" pageId="3" pageNumber="62">Pointed to rounded</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92CF85FACFB31" box="[151,1436,1198,1221]" gridrow="2" pageId="3" pageNumber="62" rowspan-1="1" rowspan-2="1">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92CF85BEAFB31" box="[151,474,1198,1221]" gridcol="0" gridrow="2" pageId="3" pageNumber="62">Rostral pattern in dorsal view</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92C8D5FACFB06" box="[151,1436,1243,1266]" gridrow="3" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92C8D5BEAFB06" box="[151,474,1243,1266]" gridcol="0" gridrow="3" pageId="3" pageNumber="62">Acuminate posteriorly</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2C8D59A7FB06" box="[724,919,1243,1266]" gridcol="1" gridrow="3" pageId="3" pageNumber="62">Weakly</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522C8D5FACFB06" box="[1084,1436,1243,1266]" gridcol="2" gridrow="3" pageId="3" pageNumber="62">Strongly</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92D5F5FACFAD4" box="[151,1436,1289,1312]" gridrow="4" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92D5F5BEAFAD4" box="[151,474,1289,1312]" gridcol="0" gridrow="4" pageId="3" pageNumber="62">Umbo</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2D5F59A7FAD4" box="[724,919,1289,1312]" gridcol="1" gridrow="4" pageId="3" pageNumber="62">Weakly</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522D5F5FACFAD4" box="[1084,1436,1289,1312]" gridcol="2" gridrow="4" pageId="3" pageNumber="62">Strongly</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92D605FACFAB9" box="[151,1436,1334,1357]" gridrow="5" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92D605BEAFAB9" box="[151,474,1334,1357]" gridcol="0" gridrow="5" pageId="3" pageNumber="62">Flared on anterior apex</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2D6059A7FAB9" box="[724,919,1334,1357]" gridcol="1" gridrow="5" pageId="3" pageNumber="62">No</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522D605FACFAB9" box="[1084,1436,1334,1357]" gridcol="2" gridrow="5" pageId="3" pageNumber="62">Strongly</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92D355FACFA8E" box="[151,1436,1379,1402]" gridrow="6" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92D355BEAFA8E" box="[151,474,1379,1402]" gridcol="0" gridrow="6" pageId="3" pageNumber="62">Parallel sides midrostral</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2D3559A7FA8E" box="[724,919,1379,1402]" gridcol="1" gridrow="6" pageId="3" pageNumber="62">Nearly</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522D355FACFA8E" box="[1084,1436,1379,1402]" gridcol="2" gridrow="6" pageId="3" pageNumber="62">No</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92DC75FACF9C4" box="[151,1436,1425,1584]" gridrow="7" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92DC75BEAF9C4" box="[151,474,1425,1584]" gridcol="0" gridrow="7" pageId="3" pageNumber="62">RWD1/RLD2 RWD2/RWD1 ((RWD1 + RTL)/TL)*100 OLM/OW</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2DC759A7F9C4" box="[724,919,1425,1584]" gridcol="1" gridrow="7" pageId="3" pageNumber="62">1.03 1.01 1.90 0.80</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522DC75FACF9C4" box="[1084,1436,1425,1584]" gridcol="2" gridrow="7" pageId="3" pageNumber="62">
|
|||
|
|
0.97 1.04 2.46
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8F522E4E5E7BF9C4" bold="true" box="[1084,1099,1560,1584]" pageId="3" pageNumber="62">≈</emphasis>
|
|||
|
|
0.5
|
|||
|
|
</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="354734E1FFD4F2B28BF92E105FACF9A9" box="[151,1436,1606,1629]" gridrow="8" pageId="3" pageNumber="62">
|
|||
|
|
<th id="76965D9DFFD4F2B28BF92E105BEAF9A9" box="[151,474,1606,1629]" gridcol="0" gridrow="8" pageId="3" pageNumber="62">OS</th>
|
|||
|
|
<td id="76965D9DFFD4F2B289BA2E1059A7F9A9" box="[724,919,1606,1629]" gridcol="1" gridrow="8" pageId="3" pageNumber="62">0.79</td>
|
|||
|
|
<td id="76965D9DFFD4F2B28F522E105FACF9A9" box="[1084,1436,1606,1629]" gridcol="2" gridrow="8" pageId="3" pageNumber="62">0.19</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC836A3FFD40D4E8BAB2EFC58EDF8CC" blockId="3.[151,1437,1706,1928]" pageId="3" pageNumber="62">Coloration in alcohol dark brown dorsum and light brown venter. Number of dark brown scale rows (in dorsal view) counted at neck, midbody and before vent levels are seven, 11 and 11, respectively. Whitish snout: rostral, nasals, preocular, anterior portion of frontal, mid–inferior of ocular and postocular, and four supralabials. Tail unspotted along ventral surface.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6528FFD40D498BAB2F115F77FA1A" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="etymology">
|
|||
|
|
<paragraph id="8BC836A3FFD40D4E8BAB2F1159C4F87C" blockId="3.[151,1437,1706,1928]" pageId="3" pageNumber="62">
|
|||
|
|
<emphasis id="B903EAB1FFD40D4E8BAB2F115B65F895" bold="true" box="[197,341,1863,1889]" pageId="3" pageNumber="62">Etymology.</emphasis>
|
|||
|
|
Patronymic of Golyath (Later Latin translated from Hebrew), biblical giant from book 1st Samuel, Old Testament. Name in allusion to the large size of this species.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF08662BFFD30D498BF92CA65F0CFAF3" box="[151,1340,1264,1288]" httpUri="https://zenodo.org/record/186241/files/figure.png" pageId="4" pageNumber="63" targetBox="[475,1112,219,1222]" targetPageId="4">
|
|||
|
|
<paragraph id="8BC836A3FFD30D498BF92CA65F0CFAF3" blockId="4.[151,1340,1264,1288]" box="[151,1340,1264,1288]" pageId="4" pageNumber="63">
|
|||
|
|
<emphasis id="B903EAB1FFD30D498BF92CA65B29FAFC" bold="true" box="[151,281,1264,1288]" pageId="4" pageNumber="63">FIGURE 1.</emphasis>
|
|||
|
|
<taxonomicName id="4C774D20FFD30D498A4E2CA65BEDFAF3" box="[288,477,1264,1287]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="golyathi">
|
|||
|
|
<emphasis id="B903EAB1FFD30D498A4E2CA65BEDFAF3" box="[288,477,1264,1287]" italics="true" pageId="4" pageNumber="63">Typhlops golyathi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(holotype) dorsal and lateral views, showing the head scutellation (bar = 2 mm).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC836A3FFD30D498BAB2D6F5F77FA1A" blockId="4.[151,1437,1337,1596]" pageId="4" pageNumber="63">
|
|||
|
|
<emphasis id="B903EAB1FFD30D498BAB2D6F5BA2FAA7" bold="true" box="[197,402,1337,1363]" pageId="4" pageNumber="63">Natural History.</emphasis>
|
|||
|
|
The animal was collected under a stone, during the day. This region reaches 28 – 30 ºC maximum temperature in summer and 14 – 16 ºC minimum temperature in winter. It receives approximately 1
|
|||
|
|
<quantity id="4C8F9B46FFD30D498BF92DD15ACDFA55" box="[151,253,1415,1441]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="63" unit="mm" value="800.0">800 mm</quantity>
|
|||
|
|
of annual precipitation (Instituto Cubano de Geodesia y Cartografía, 1978). The vegetation is characterized by semideciduous and evergreen forests, and vegetation complex of “mogote” – peculiar karst mountains called “mogotes” in Spanish, from Western and Central
|
|||
|
|
<collectingCountry id="F3607633FFD30D4988C42D8259D8FA1A" box="[938,1000,1492,1518]" name="Cuba" pageId="4" pageNumber="63">Cuba</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFE64B52FFD30D4988992D825F0AFA1A" author="Capote" box="[1015,1338,1492,1518]" pageId="4" pageNumber="65" refString="Capote, R. P. & Berazain, R. (1984) Clasificacion de las formaciones vegetales de Cuba. Revista del Jardin Botanico Nacional de Cuba, 5, 27 - 75." type="journal article" year="1984">Capote and Berazaín, 1984</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6528FFD30D498BAB2DAC58C8F9C8" pageId="4" pageNumber="63" type="distribution">
|
|||
|
|
<paragraph id="8BC836A3FFD30D498BAB2DAC58C8F9C8" blockId="4.[151,1437,1337,1596]" pageId="4" pageNumber="63">
|
|||
|
|
<emphasis id="B903EAB1FFD30D498BAB2DAC5B58F9E1" bold="true" box="[197,360,1530,1557]" pageId="4" pageNumber="63">Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="4C774D20FFD30D498A1F2DAD5877F9E0" box="[369,583,1531,1556]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="golyathi">
|
|||
|
|
<emphasis id="B903EAB1FFD30D498A1F2DAD5877F9E0" box="[369,583,1531,1556]" italics="true" pageId="4" pageNumber="63">Typhlops golyathi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is known only from the
|
|||
|
|
<typeStatus id="54CC8801FFD30D49881C2DAD5995F9E1" box="[882,933,1531,1557]" pageId="4" pageNumber="63">type</typeStatus>
|
|||
|
|
–locality of Valle de
|
|||
|
|
<collectingCountry id="F3607633FFD30D498FCC2DAD5F06F9E1" box="[1186,1334,1531,1557]" name="Saint Vincent and the Grenadines" pageId="4" pageNumber="63">San Vicente</collectingCountry>
|
|||
|
|
, Viñales Municipality, Pinar del Río Province,
|
|||
|
|
<collectingCountry id="F3607633FFD30D49893D2E7458A1F9C8" box="[595,657,1570,1596]" name="Cuba" pageId="4" pageNumber="63">Cuba</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="134C2A26FFD30D4989CC2E7458D9F9C8" box="[674,745,1570,1596]" captionStart="FIGURE 2" captionStartId="5.[151,256,1312,1336]" captionTargetBox="[158,1424,521,1280]" captionTargetId="figure@5.[151,1436,514,1288]" captionTargetPageId="5" captionText="FIGURE 2. Geographic distribution of the Typhlops biminiensis species group in Cuba. Closed star = T. golyathi sp. nov.; closed square = T. anchaurus; mixed square = T. anousius; closed rhombus = T. arator; mixed circle = T. contorhinus; closed non – inverse triangles = T. notorachius; closed circles = T. perimychus; closed inverse triangle = T. satelles." httpUri="https://zenodo.org/record/186242/files/figure.png" pageId="4" pageNumber="63">Fig. 2</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|