306 lines
38 KiB
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306 lines
38 KiB
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<document id="AB87A3EB264498AD3B82C8F6046D0007" ID-CLB-Dataset="4637" ID-DOI="10.1515/aemnp-2017-0113" ID-GBIF-Dataset="598ea0a7-8800-412c-b62c-b73757407163" ID-ISSN="0374-1036" ID-Zenodo-Dep="5325777" ID-ZooBank="BB191E4F-DC19-4D2F-9EBF-1EC4D6A5BE52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630151790382" checkinUser="marcus" docAuthor="Plonski, Isidor S." docDate="2017" docId="039E3626F232DD7E66B23B37FD5C6DFC" docLanguage="en" docName="ActaEntMusNatPra.2017.57.suppl.125-131.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57" docSource="http://dx.doi.org/10.1515/aemnp-2017-0113" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Colotrema socotrana Plonski 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="129" masterDocId="FFA74E5EF233DD7A670A3F03FFF46E4F" masterDocTitle="Colotrema socotrana sp. nov. from Socotra Island, with new records of the genus from Yemen (Coleoptera: Malachiidae)" masterLastPageNumber="131" masterPageNumber="125" pageNumber="126" updateTime="1699173374667" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F31FA294044AB36F4EB75E0B8DB10FCF">Colotrema socotrana sp. nov. from Socotra Island, with new records of the genus from Yemen (Coleoptera: Malachiidae)</mods:title>
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<mods:namePart id="2EB82ACED6434CB4C06E55F5B884CDA4">Plonski, Isidor S.</mods:namePart>
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<mods:title id="477A79453AF15F6466A3D1162785AA35">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="131DBBC149FEA5CDBF04EC8450B09C92">
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<mods:date id="9B0DB20F26BC7E9378B63E0B312F577B">2017</mods:date>
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<mods:title id="7F28AFBD30214CE688759F980C044992">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:detail id="FAA1D97309B2FEEE2D67346DC2AC5B91" type="pubDate">
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<mods:number id="82039515C8585CA2F3CA9015FC63EF1B">2017-12-31</mods:number>
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<mods:number id="03EE1D824D4CEBC4D5B579DF078B39B8">57</mods:number>
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<mods:url id="66058018B5BB3EB072541E15D6F262B4">http://dx.doi.org/10.1515/aemnp-2017-0113</mods:url>
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<mods:classification id="0F20EC62083C175717714EA9A832D383">journal article</mods:classification>
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<mods:identifier id="0164D6708D30E5FA74E111CB62001740" type="DOI">10.1515/aemnp-2017-0113</mods:identifier>
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<treatment id="039E3626F232DD7E66B23B37FD5C6DFC" ID-DOI="http://doi.org/10.5281/zenodo.5345917" ID-GBIF-Taxon="185207696" ID-Zenodo-Dep="5345917" LSID="urn:lsid:plazi:treatment:039E3626F232DD7E66B23B37FD5C6DFC" httpUri="http://treatment.plazi.org/id/039E3626F232DD7E66B23B37FD5C6DFC" lastPageId="4" lastPageNumber="129" pageId="1" pageNumber="126">
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<subSubSection id="C32DD4BBF232DD7B66B23B37FD0E6A03" box="[440,762,1076,1100]" pageId="1" pageNumber="126" type="nomenclature">
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<paragraph id="8B888730F232DD7B66B23B37FD0E6A03" blockId="1.[440,762,1076,1100]" box="[440,762,1076,1100]" pageId="1" pageNumber="126">
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<heading id="D0C0305CF232DD7B66B23B37FD0E6A03" bold="true" box="[440,762,1076,1100]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="126" reason="3">
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<emphasis id="B9435B22F232DD7B66B23B37FD0E6A03" bold="true" box="[440,762,1076,1100]" pageId="1" pageNumber="126">
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<taxonomicName id="4C37FCB3F232DD7B66B23B37FD6A6A03" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" box="[440,670,1076,1100]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
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<emphasis id="B9435B22F232DD7B66B23B37FD6A6A03" bold="true" box="[440,670,1076,1100]" italics="true" pageId="1" pageNumber="126">Colotrema socotrana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A270E659F232DD7B65AF3B36FD0E6A03" box="[677,762,1077,1100]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32DD4BBF232DD7B651D3B50FD6E6A29" box="[535,666,1107,1126]" pageId="1" pageNumber="126" type="description">
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<paragraph id="8B888730F232DD7B651D3B50FD6E6A29" blockId="1.[535,666,1107,1126]" box="[535,666,1107,1126]" pageId="1" pageNumber="126">
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(
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<figureCitation id="130C9BB5F232DD7B65173B50FD9E6A29" box="[541,618,1107,1126]" captionStart="Figs 1–4" captionStartId="2.[151,188,1541,1560]" captionTargetBox="[196,1075,222,1482]" captionTargetId="figure-10@2.[170,1108,219,1517]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Colotrema. 1–2 Colotrema socotrana sp. nov.: 1 – female allotype; 2 – male paratype #1; 3 – C. assirensis Wittmer, 1979; 4 – C. testacea wajjensis Wittmer, 1980. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.5325779" httpUri="https://zenodo.org/record/5325779/files/figure.png" pageId="1" pageNumber="126">Figs 1–2</figureCitation>
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,
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<figureCitation id="130C9BB5F232DD7B65793B50FD606A29" box="[627,660,1107,1126]" captionStart="Figs 5–8" captionStartId="3.[113,150,1516,1535]" captionTargetBox="[188,1001,650,1482]" captionTargetId="figure-150@3.[184,1019,624,1510]" captionTargetPageId="3" captionText="Figs 5–8. Details of holotype of Colotrema socotrana sp. nov.5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="1" pageNumber="126">5–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32DD4BBF232DD7B677B3B7EFDD46B2A" pageId="1" pageNumber="126" type="materials_examined">
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<paragraph id="8B888730F232DD7B677B3B7EFD6C6AE4" blockId="1.[113,1089,1149,1381]" pageId="1" pageNumber="126">
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<materialsCitation id="3B5F8D6DF232DD7B677B3B7EFD606AE4" ID-GBIF-Occurrence="3339968302" country="Socotra Island" latitude="12.519444" location="Environment" longLatPrecision="21" longitude="54.123333" municipality="Aloove village" pageId="1" pageNumber="126" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9435B22F232DD7B677B3B7EFF1B6ADF" bold="true" box="[113,239,1149,1168]" pageId="1" pageNumber="126">
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<typeStatus id="548C3992F232DD7B677B3B7EFF6A6ADF" box="[113,158,1149,1168]" pageId="1" pageNumber="126">Type</typeStatus>
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locality.
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</emphasis>
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<location id="8EE8D1EBF232DD7B67FD3B7EFE9C6ADF" LSID="urn:lsid:plazi:treatment:039E3626F232DD7E66B23B37FD5C6DFC:8EE8D1EBF232DD7B67FD3B7EFE9C6ADF" box="[247,360,1149,1168]" country="Socotra Island" latitude="12.519444" longLatPrecision="21" longitude="54.123333" municipality="Aloove village" name="Environment" pageId="1" pageNumber="126">Environment</location>
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(georeferenced on the locality label cited below) of
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<collectingMunicipality id="6BEC1D4AF232DD7B644D3B7EFC3E6ADF" box="[839,970,1149,1168]" pageId="1" pageNumber="126">Aloove village</collectingMunicipality>
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(
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<geoCoordinate id="EE03E1F7F232DD7B64D33B7EFBB46ADF" box="[985,1088,1149,1168]" degrees="12" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="126" precision="15" seconds="10" value="12.519444">12°31′10′′N</geoCoordinate>
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<geoCoordinate id="EE03E1F7F232DD7B677B3B9BFF236AE4" box="[113,215,1176,1195]" degrees="54" direction="east" minutes="07" orientation="longitude" pageId="1" pageNumber="126" precision="15" seconds="24" value="54.123333">54°07′24′′E</geoCoordinate>
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; see
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<bibRefCitation id="EFA6FAC1F232DD7B660B3B9BFE366AE4" author="BEZDEK J. & PURCHART L. & KRAL K. & HULA V." box="[257,450,1176,1195]" pageId="1" pageNumber="126" pagination="1 - 557" refId="ref2564" refString="BEZDEK J., PURCHART L., KRAL K. & HULA V. 2012: List of local Socotran geographical names used in entomological literature. Pp. 27 - 67. In: HAJEK J. & BEZDEK J. (eds): Insect biodiversity of the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52 (Supplementum 2): i - vi + 1 - 557." type="journal article" year="2012">BEZDĚK et al. 2012: 34</bibRefCitation>
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),
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<collectingCountry id="F320C7A0F232DD7B66DB3B9BFDBA6AE4" box="[465,590,1176,1195]" name="Socotra Island" pageId="1" pageNumber="126">Socotra Island</collectingCountry>
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,
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<collectingCountry id="F320C7A0F232DD7B655C3B9BFD606AE4" box="[598,660,1176,1195]" name="Yemen" pageId="1" pageNumber="126">Yemen</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B888730F232DD7B677B3BB1FDD46B2A" blockId="1.[113,1089,1149,1381]" pageId="1" pageNumber="126">
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<emphasis id="B9435B22F232DD7B677B3BB1FF0D6A8A" bold="true" box="[113,249,1202,1221]" pageId="1" pageNumber="126">Type material.</emphasis>
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<typeStatus id="548C3992F232DD7B660B3BB1FEA06A8C" box="[257,340,1202,1221]" pageId="1" pageNumber="126" type="holotype">HOLOTYPE</typeStatus>
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: J (
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<collectionCode id="ED261FF5F232DD7B668A3BB1FE356A8A" box="[384,449,1202,1221]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="126">NMPC</collectionCode>
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), labelled: ‘
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<collectingCountry id="F320C7A0F232DD7B65223BB1FD8E6A8A" box="[552,634,1202,1221]" name="Yemen" pageId="1" pageNumber="126">YEMEN</collectingCountry>
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,
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<collectingCountry id="F320C7A0F232DD7B658F3BB1FCE46A8C" box="[645,784,1202,1221]" name="SOCOTRA ISLAND" pageId="1" pageNumber="126">SOCOTRA ISLAND</collectingCountry>
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/ Aloove area, ALOOVE vill. env. /
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<taxonomicName id="4C37FCB3F232DD7B677B3BCEFEE96AAF" box="[113,285,1229,1248]" class="Magnoliopsida" family="Euphorbiaceae" genus="Jatropha" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="unicostata">
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<emphasis id="B9435B22F232DD7B677B3BCEFEE96AAF" box="[113,285,1229,1248]" italics="true" pageId="1" pageNumber="126">Jatropha unicostata</emphasis>
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</taxonomicName>
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shrubland; / with
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<taxonomicName id="4C37FCB3F232DD7B66B73BCEFD976AAF" box="[445,611,1229,1248]" class="Magnoliopsida" family="Burseraceae" genus="Boswellia" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="126" phylum="Tracheophyta" rank="species" species="elongata">
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<emphasis id="B9435B22F232DD7B66B73BCEFD976AAF" box="[445,611,1229,1248]" italics="true" pageId="1" pageNumber="126">Boswellia elongata</emphasis>
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</taxonomicName>
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trees /
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<date id="FF89A1F0F232DD7B65AF3BCEFCD56AAF" box="[677,801,1229,1248]" pageId="1" pageNumber="126" value="2012-06-19" valueMax="2012-06-20" valueMin="2012-06-19">19.-20.vi.2012</date>
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/
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<geoCoordinate id="EE03E1F7F232DD7B643F3BCEFC646AAF" box="[821,912,1229,1248]" degrees="12" direction="north" minutes="31.2" orientation="latitude" pageId="1" pageNumber="126" precision="92" value="12.52">12°31.2’N</geoCoordinate>
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,
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<geoCoordinate id="EE03E1F7F232DD7B64903BCEFC076AAF" box="[922,1011,1229,1248]" degrees="54" direction="east" minutes="07.4" orientation="longitude" pageId="1" pageNumber="126" precision="92" value="54.123333">54°07.4’E</geoCoordinate>
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,
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<quantity id="4CCF2AD5F232DD7B64F43BCEFBDA6AAF" box="[1022,1070,1229,1248]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="1" pageNumber="126" unit="m" value="221.0">221m</quantity>
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//
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<collectingCountry id="F320C7A0F232DD7B677B3BEBFF4E6AB6" box="[113,186,1256,1275]" name="Yemen" pageId="1" pageNumber="126">SOCOTRA</collectingCountry>
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expedition 2012 / J. Bezděk, J. Hájek,
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<collectionCode id="ED261FF5F232DD7B65073BEBFDE86AB4" box="[525,540,1256,1275]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="126" type="Museum">V</collectionCode>
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. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg.’.
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<typeStatus id="548C3992F232DD7B67A83A01FF076B5C" box="[162,243,1282,1301]" pageId="1" pageNumber="126" type="allotype">ALLOTYPE</typeStatus>
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:
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<specimenCount id="9D314CB9F232DD7B67F53BFFFEFB6B5A" box="[255,271,1276,1301]" pageId="1" pageNumber="126" type="female">♀</specimenCount>
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(
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<collectionCode id="ED261FF5F232DD7B66103A01FEAF6B5A" box="[282,347,1282,1301]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="126">NMPC</collectionCode>
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), same data as
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<typeStatus id="548C3992F232DD7B66D13A01FDDC6B5A" box="[475,552,1282,1301]" pageId="1" pageNumber="126" type="holotype">holotype</typeStatus>
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.
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<typeStatus id="548C3992F232DD7B653B3A01FD7D6B5C" box="[561,649,1282,1301]" pageId="1" pageNumber="126" type="paratype">PARATYPES</typeStatus>
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: 1 J
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<specimenCount id="9D314CB9F232DD7B65B33A01FD1C6B5A" box="[697,744,1276,1301]" pageId="1" pageNumber="126" type="female">2 ♀♀</specimenCount>
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(1 J (
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<collectionCode id="ED261FF5F232DD7B642B3A01FCCA6B5A" box="[801,830,1282,1301]" pageId="1" pageNumber="126">PT</collectionCode>
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#1) and
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<specimenCount id="9D314CB9F232DD7B648B3A01FC556B5A" box="[897,929,1276,1301]" pageId="1" pageNumber="126" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="ED261FF5F232DD7B64A63A01FC336B5A" box="[940,967,1282,1301]" pageId="1" pageNumber="126">PT</collectionCode>
|
|||
|
|
#2) in
|
|||
|
|
<collectionCode id="ED261FF5F232DD7B64F43A01FBC96B5A" box="[1022,1085,1282,1301]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="126">NMPC</collectionCode>
|
|||
|
|
;
|
|||
|
|
<specimenCount id="9D314CB9F232DD7B677B3A1EFF666B60" box="[113,146,1302,1328]" pageId="1" pageNumber="126" type="female">1 ♀</specimenCount>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="ED261FF5F232DD7B67943A1EFF4D6B7F" box="[158,185,1309,1328]" pageId="1" pageNumber="126">PT</collectionCode>
|
|||
|
|
#3) in
|
|||
|
|
<collectionCode id="ED261FF5F232DD7B67F83A1EFED26B7F" box="[242,294,1309,1328]" pageId="1" pageNumber="126">IPVA</collectionCode>
|
|||
|
|
): ‘
|
|||
|
|
<collectingCountry id="F320C7A0F232DD7B66323A1EFE7E6B7F" box="[312,394,1309,1328]" name="Yemen" pageId="1" pageNumber="126">YEMEN</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F320C7A0F232DD7B66993A1EFDE86B61" box="[403,540,1309,1328]" name="SOCOTRA ISLAND" pageId="1" pageNumber="126">SOCOTRA ISLAND</collectingCountry>
|
|||
|
|
/ Noged plain, ABATARO / border of sand dunes and / shrubland
|
|||
|
|
<date id="FF89A1F0F232DD7B677B3A3BFF1A6B04" box="[113,238,1336,1355]" pageId="1" pageNumber="126" value="2012-06-12" valueMax="2012-06-13" valueMin="2012-06-12">12.-13.vi.2012</date>
|
|||
|
|
/
|
|||
|
|
<geoCoordinate id="EE03E1F7F232DD7B67F73A3BFEAD6B04" box="[253,345,1336,1355]" degrees="12" direction="north" minutes="22.1" orientation="latitude" pageId="1" pageNumber="126" precision="92" value="12.368334">12°22.1’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE03E1F7F232DD7B666B3A3BFE316B04" box="[353,453,1336,1355]" degrees="54" direction="east" minutes="03.04" orientation="longitude" pageId="1" pageNumber="126" precision="9" value="54.050667">54°03.04’E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4CCF2AD5F232DD7B66C73A3BFE076B04" box="[461,499,1336,1355]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="126" unit="m" value="20.0">20m</quantity>
|
|||
|
|
//
|
|||
|
|
<collectingCountry id="F320C7A0F232DD7B65023A3BFDA56B06" box="[520,593,1336,1355]" name="Yemen" pageId="1" pageNumber="126">SOCOTRA</collectingCountry>
|
|||
|
|
expedition 2012 / J. Bezděk, J. Hájek,
|
|||
|
|
<collectionCode id="ED261FF5F232DD7B64933A3BFC5C6B04" box="[921,936,1336,1355]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="126" type="Museum">V</collectionCode>
|
|||
|
|
. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg.’.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32DD4BBF232DD7E677B3A7DFDBF6D1C" lastPageId="4" lastPageNumber="129" pageId="1" pageNumber="126" type="description">
|
|||
|
|
<paragraph id="8B888730F232DD7B677B3A7DFD9E6BBA" blockId="1.[113,1089,1406,1589]" pageId="1" pageNumber="126">
|
|||
|
|
<emphasis id="B9435B22F232DD7B677B3A7DFE356BDA" bold="true" box="[113,449,1406,1429]" pageId="1" pageNumber="126">
|
|||
|
|
Description of male
|
|||
|
|
<typeStatus id="548C3992F232DD7B66523A7DFE486BDA" box="[344,444,1406,1429]" pageId="1" pageNumber="126" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Colouration: Head capsule, mouthparts, antennae, pro- and mesothorax, legs, sides and apices of elytra, sternum VIII and pygidium yellow to orange; eyes, metathorax, scutellum and elytra (except sides, apices and suture) black; elytral suture and remaining abdomen lightened to brownish.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B888730F232DD79679A3AFDFE8F6F5C" blockId="1.[113,1089,1406,1589]" lastBlockId="3.[113,1089,220,595]" lastPageId="3" lastPageNumber="128" pageId="1" pageNumber="126">
|
|||
|
|
Head capsule broader than long, with coriaceous microsculpture and fine whitish setae diverging towards compound eyes. Apical maxillary palpomere polygonal in outline (
|
|||
|
|
<figureCitation id="130C9BB5F232DD7B64F8391DFBC7687A" box="[1010,1075,1566,1589]" captionStart="Figs 5–8" captionStartId="3.[113,150,1516,1535]" captionTargetBox="[188,1001,650,1482]" captionTargetId="figure-150@3.[184,1019,624,1510]" captionTargetPageId="3" captionText="Figs 5–8. Details of holotype of Colotrema socotrana sp. nov.5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="1" pageNumber="126">Fig. 5</figureCitation>
|
|||
|
|
). Scapus elongate, clavate, as long as pedicel and antennomere I together; adjoining antennomeres longer than broad.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF48D7B8F231DD78679D3906FCB4687C" ID-DOI="http://doi.org/10.5281/zenodo.5325779" ID-Zenodo-Dep="5325779" httpUri="https://zenodo.org/record/5325779/files/figure.png" pageId="2" pageNumber="127" startId="2.[151,188,1541,1560]" targetBox="[196,1075,222,1482]" targetPageId="2">
|
|||
|
|
<paragraph id="8B888730F231DD78679D3906FCB4687C" blockId="2.[151,1129,1541,1587]" pageId="2" pageNumber="127">
|
|||
|
|
Figs 1–4. Habitus of
|
|||
|
|
<taxonomicName id="4C37FCB3F231DD78664D3906FE556857" authorityName="Wittmer" authorityYear="1979" box="[327,417,1541,1560]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9435B22F231DD78664D3906FE556857" box="[327,417,1541,1560]" italics="true" pageId="2" pageNumber="127">Colotrema</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. 1–2
|
|||
|
|
<taxonomicName id="4C37FCB3F231DD7866C53906FD776857" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" box="[463,643,1541,1560]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
|
|||
|
|
<emphasis id="B9435B22F231DD7866C53906FD776857" box="[463,643,1541,1560]" italics="true" pageId="2" pageNumber="127">Colotrema socotrana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A270E659F231DD78658D3906FD386857" box="[647,716,1541,1560]" pageId="2" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
: 1 – female allotype; 2 – male paratype #1; 3 –
|
|||
|
|
<taxonomicName id="4C37FCB3F231DD78635F3906FE99687C" authority="Wittmer, 1979" authorityName="Wittmer" authorityYear="1979" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="assirensis">
|
|||
|
|
<emphasis id="B9435B22F231DD78635F3906FF1E687C" italics="true" pageId="2" pageNumber="127">C. assirensis</emphasis>
|
|||
|
|
<bibRefCitation id="EFA6FAC1F231DD7867FA3923FE99687C" author="WITTMER W." box="[240,365,1568,1587]" pageId="2" pageNumber="127" pagination="186 - 192" refId="ref2866" refString="WITTMER W. 1979: Insects of Saudi Arabia. Coleoptera: Fam. Malachiidae, Dasytidae, Karumiidae. Fauna of Saudi Arabia 1: 186 - 192." type="journal article" year="1979">Wittmer, 1979</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; 4 –
|
|||
|
|
<taxonomicName id="4C37FCB3F231DD78669C3923FD3F687C" authority="Wittmer, 1980" authorityName="Wittmer" authorityYear="1980" box="[406,715,1568,1587]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="testacea" subSpecies="wajjensis">
|
|||
|
|
<emphasis id="B9435B22F231DD78669C3923FDBD687C" box="[406,585,1568,1587]" italics="true" pageId="2" pageNumber="127">C. testacea wajjensis</emphasis>
|
|||
|
|
<bibRefCitation id="EFA6FAC1F231DD7865443923FD3F687C" author="WITTMER W." box="[590,715,1568,1587]" pageId="2" pageNumber="127" pagination="114 - 118" refId="ref2896" refString="WITTMER W. 1980: Fauna of Saudi Arabia. Coleoptera: Fam. Drilidae, Malachiidae, Prionoceridae (2. Beitrag). Fauna of Saudi Arabia 2: 114 - 118." type="journal article" year="1980">Wittmer, 1980</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Not in scale.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B888730F230DD79679A3E1FFC9F6F9C" blockId="3.[113,1089,220,595]" pageId="3" pageNumber="128">Pronotum transverse, broadest next to apex; apex and base weakly arcuate, sides more strongly arcuate, apical corners rounded, basal corners strongly rounded; side margins distinct; disc moderately convex with coriaceous microsculpture and punctures, whose distance to each other equals 2–3 times their diameter; pubescence similar to that of head capsule. Elytra oblong-ovate; microsculpture, punctures and pubescence similar to those on pronotum; anteapically with circular impression. Front legs with tetramerous tarsi.</paragraph>
|
|||
|
|
<paragraph id="8B888730F230DD79679A3EDFFDEB6C1C" blockId="3.[113,1089,220,595]" pageId="3" pageNumber="128">
|
|||
|
|
Pygidium (
|
|||
|
|
<figureCitation id="130C9BB5F230DD79660D3EDFFEBC6FBC" box="[263,328,476,499]" captionStart="Figs 5–8" captionStartId="3.[113,150,1516,1535]" captionTargetBox="[188,1001,650,1482]" captionTargetId="figure-150@3.[184,1019,624,1510]" captionTargetPageId="3" captionText="Figs 5–8. Details of holotype of Colotrema socotrana sp. nov.5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="3" pageNumber="128">Fig. 6</figureCitation>
|
|||
|
|
) with long basal struts, medially less sclerotized region, and bisinuate apex. Sternite VIII (
|
|||
|
|
<figureCitation id="130C9BB5F230DD79664A3EFFFE896C5C" box="[320,381,508,531]" captionStart="Figs 5–8" captionStartId="3.[113,150,1516,1535]" captionTargetBox="[188,1001,650,1482]" captionTargetId="figure-150@3.[184,1019,624,1510]" captionTargetPageId="3" captionText="Figs 5–8. Details of holotype of Colotrema socotrana sp. nov.5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="3" pageNumber="128">Fig. 7</figureCitation>
|
|||
|
|
) with species-specific apex. Median lobe (
|
|||
|
|
<figureCitation id="130C9BB5F230DD79643C3EFFFC876C5C" box="[822,883,508,531]" captionStart="Figs 5–8" captionStartId="3.[113,150,1516,1535]" captionTargetBox="[188,1001,650,1482]" captionTargetId="figure-150@3.[184,1019,624,1510]" captionTargetPageId="3" captionText="Figs 5–8. Details of holotype of Colotrema socotrana sp. nov.5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="3" pageNumber="128">Fig. 8</figureCitation>
|
|||
|
|
) subparallel at base and in middle, then conically widened, with 3–4 big endophallic sclerites in middle portion, and big bilateral ankylotic setae at apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF48D7B8F230DD79677B3AEFFCA26855" ID-DOI="http://doi.org/10.5281/zenodo.5325783" ID-Zenodo-Dep="5325783" httpUri="https://zenodo.org/record/5325783/files/figure.png" pageId="3" pageNumber="128" startId="3.[113,150,1516,1535]" targetBox="[188,1001,650,1482]" targetPageId="3">
|
|||
|
|
<paragraph id="8B888730F230DD79677B3AEFFCA26855" blockId="3.[113,1089,1516,1562]" pageId="3" pageNumber="128">
|
|||
|
|
Figs 5–8. Details of holotype of
|
|||
|
|
<taxonomicName id="4C37FCB3F230DD7966763AEFFDDB6BB0" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" box="[380,559,1516,1535]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
|
|||
|
|
<emphasis id="B9435B22F230DD7966763AEFFDDB6BB0" box="[380,559,1516,1535]" italics="true" pageId="3" pageNumber="128">Colotrema socotrana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A270E659F230DD7965393AEFFD806BB0" box="[563,628,1516,1535]" pageId="3" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
5 – left maxillary palpus, ventral; 6 – pygidium, ventral view; 7 – sternite VIII, dorsal view; 8 – median lobe, lateral view. Scale bar = 0.2 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B888730F237DD7E679D3FDFFD1A6F1C" blockId="4.[151,1127,220,947]" pageId="4" pageNumber="129">
|
|||
|
|
<emphasis id="B9435B22F237DD7E679D3FDFFEE26EBC" bold="true" box="[151,278,220,243]" pageId="4" pageNumber="129">Variability.</emphasis>
|
|||
|
|
The specimens of the
|
|||
|
|
<typeStatus id="548C3992F237DD7E65003FDFFDCC6EBC" box="[522,568,220,243]" pageId="4" pageNumber="129">type</typeStatus>
|
|||
|
|
series vary considerably in elytral colouration from almost black elytra with only sides and apices near suture yellow (
|
|||
|
|
<figureCitation id="130C9BB5F237DD7E644D3FFFFC776F5C" box="[839,899,252,275]" captionStart="Figs 1–4" captionStartId="2.[151,188,1541,1560]" captionTargetBox="[196,1075,222,1482]" captionTargetId="figure-10@2.[170,1108,219,1517]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Colotrema. 1–2 Colotrema socotrana sp. nov.: 1 – female allotype; 2 – male paratype #1; 3 – C. assirensis Wittmer, 1979; 4 – C. testacea wajjensis Wittmer, 1980. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.5325779" httpUri="https://zenodo.org/record/5325779/files/figure.png" pageId="4" pageNumber="129">Fig. 1</figureCitation>
|
|||
|
|
) to elytra extensively yellow with black colour restricted to longitudinal club-shaped macula on the disc extending from humeral bulges to the apical third of elytra (
|
|||
|
|
<figureCitation id="130C9BB5F237DD7E65AC3E3FFD126F1C" box="[678,742,316,339]" captionStart="Figs 1–4" captionStartId="2.[151,188,1541,1560]" captionTargetBox="[196,1075,222,1482]" captionTargetId="figure-10@2.[170,1108,219,1517]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Colotrema. 1–2 Colotrema socotrana sp. nov.: 1 – female allotype; 2 – male paratype #1; 3 – C. assirensis Wittmer, 1979; 4 – C. testacea wajjensis Wittmer, 1980. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.5325779" httpUri="https://zenodo.org/record/5325779/files/figure.png" pageId="4" pageNumber="129">Fig. 2</figureCitation>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B888730F237DD7E67BF3E5FFE356F9C" blockId="4.[151,1127,220,947]" pageId="4" pageNumber="129">Sexual dimorphism. Males with longer antennae, modified apical maxillary palpomere, anteapical circular elytral impression and tetramerous fore tarsi. Females with shorter antennae, broad and apically truncated apical maxillary palpomere, pentamerous foretarsi, and without elytral impressions.</paragraph>
|
|||
|
|
<paragraph id="8B888730F237DD7E67BF3EDFFDBF6D1C" blockId="4.[151,1127,220,947]" pageId="4" pageNumber="129">
|
|||
|
|
Measurements.AL = 0.72–1.06 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E65003EDFFD9C6FBC" box="[522,616,476,499]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E65673EDFFD3A6FBC" box="[621,718,476,499]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="129" unit="mm" value="0.91">0.91 mm</quantity>
|
|||
|
|
); HW = 0.54–0.65 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E649D3EDFFC016FBC" box="[919,1013,476,499]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E64F03EDFFBAF6FBC" box="[1018,1115,476,499]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="4" pageNumber="129" unit="mm" value="0.59">0.59 mm</quantity>
|
|||
|
|
); IOW = 0.39–0.50 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E66573EFFFE486C5C" box="[349,444,508,531]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E66CB3EFFFDD76C5C" box="[449,547,508,531]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="129" unit="mm" value="0.39">0.39 mm</quantity>
|
|||
|
|
); PW = 0.65–0.76 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E65E73EFFFCB86C5C" box="[749,844,508,531]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E645B3EFFFC476C5C" box="[849,947,508,531]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="129" unit="mm" value="0.65">0.65 mm</quantity>
|
|||
|
|
); PL = 0.37–0.46 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E67943D1FFF0F6C7C" box="[158,251,540,563]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E67F53D1FFEAA6C7C" box="[255,350,540,563]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="129" unit="mm" value="0.39">0.39 mm</quantity>
|
|||
|
|
); EW = 0.91–1.09 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E65283D1FFD8B6C7C" box="[546,639,540,563]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
|||
|
|
<quantity id="4CCF2AD5F237DD7E65893D1FFD156C7C" box="[643,737,540,563]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="129" unit="mm" value="0.96">0.96 mm</quantity>
|
|||
|
|
); EL = 1.17–1.37 (
|
|||
|
|
<typeStatus id="548C3992F237DD7E64963D1FFC0D6C7C" box="[924,1017,540,563]" pageId="4" pageNumber="129" type="holotype">holotype</typeStatus>
|
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|
|
<quantity id="4CCF2AD5F237DD7E64F73D1FFBA86C7C" box="[1021,1116,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="129" unit="mm" value="1.17">1.17 mm</quantity>
|
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|
|
).
|
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|
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<emphasis id="B9435B22F237DD7E679D3D3FFE7A6C1C" bold="true" box="[151,398,572,595]" pageId="4" pageNumber="129">Differential diagnosis.</emphasis>
|
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|
|
Chromatically,
|
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|
|
<taxonomicName id="4C37FCB3F237DD7E653E3D3FFD4F6C1C" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" box="[564,699,572,595]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
|
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|
|
<emphasis id="B9435B22F237DD7E653E3D3FFD4F6C1C" box="[564,699,572,595]" italics="true" pageId="4" pageNumber="129">C. socotrana</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A270E659F237DD7E65CA3D3FFCE56C1C" box="[704,785,572,595]" pageId="4" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
|
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|
|
belongs to a group, whose members possess a completely orange pronotum and predominantly black elytra with yellow sides, apices and suture (the latter to varying extents). The most similar species is
|
|||
|
|
<taxonomicName id="4C37FCB3F237DD7E64B63D7FFEC56CFC" authority="Wittmer, 1991" authorityName="Wittmer" authorityYear="1991" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="barbertonica">
|
|||
|
|
<emphasis id="B9435B22F237DD7E64B63D7FFB926CDC" box="[956,1126,636,659]" italics="true" pageId="4" pageNumber="129">C. barbertonica</emphasis>
|
|||
|
|
<bibRefCitation id="EFA6FAC1F237DD7E679D3D9FFEC56CFC" author="WITTMER W." box="[151,305,668,691]" pageId="4" pageNumber="129" pagination="223 - 251" refId="ref2965" refString="WITTMER W. 1991: The genera Colotes Erichson and Colotrema Wittmer (Coleoptera: Malachiidae) mainly from southern Africa. Annals of the Transvaal Museum 35: 223 - 251." type="journal article" year="1991">Wittmer, 1991</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="F320C7A0F237DD7E66793D9FFE0A6CFC" box="[371,510,668,691]" name="South Africa" pageId="4" pageNumber="129">South Africa</collectingCountry>
|
|||
|
|
, which also possesses a completely orange coloured head capsule, but differs in having bicolorous extremities (cf.
|
|||
|
|
<bibRefCitation id="EFA6FAC1F237DD7E65E13DBFFC496C9C" author="WITTMER W." box="[747,957,700,723]" pageId="4" pageNumber="129" pagination="223 - 251" refId="ref2965" refString="WITTMER W. 1991: The genera Colotes Erichson and Colotrema Wittmer (Coleoptera: Malachiidae) mainly from southern Africa. Annals of the Transvaal Museum 35: 223 - 251." type="journal article" year="1991">WITTMER 1991: 240</bibRefCitation>
|
|||
|
|
). In addition,
|
|||
|
|
<taxonomicName id="4C37FCB3F237DD7E63443DBFFEF66CBC" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
|
|||
|
|
<emphasis id="B9435B22F237DD7E63443DBFFEF66CBC" italics="true" pageId="4" pageNumber="129">C. socotrana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A270E659F237DD7E660D3DDFFEAD6CBC" box="[263,345,732,755]" pageId="4" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is well characterized by the form of the male maxillary palpi (cf.
|
|||
|
|
<bibRefCitation id="EFA6FAC1F237DD7E63023DDFFF246D5C" author="WITTMER W." pageId="4" pageNumber="129" pagination="223 - 251" refId="ref2965" refString="WITTMER W. 1991: The genera Colotes Erichson and Colotrema Wittmer (Coleoptera: Malachiidae) mainly from southern Africa. Annals of the Transvaal Museum 35: 223 - 251." type="journal article" year="1991">WITTMER 1991</bibRefCitation>
|
|||
|
|
: Figs 49–50 for
|
|||
|
|
<taxonomicName id="4C37FCB3F237DD7E668D3DFFFDC06D5C" authorityName="Wittmer" authorityYear="1991" box="[391,564,764,787]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="barbertonica">
|
|||
|
|
<emphasis id="B9435B22F237DD7E668D3DFFFDC06D5C" box="[391,564,764,787]" italics="true" pageId="4" pageNumber="129">C. barbertonica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) and the last abdominal segments. Furthermore,
|
|||
|
|
<taxonomicName id="4C37FCB3F237DD7E63443DFFFEF66D7C" authority="Plonski, 2017" authorityName="Plonski" authorityYear="2017" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="socotrana" status="sp. nov.">
|
|||
|
|
<emphasis id="B9435B22F237DD7E63443DFFFEF66D7C" italics="true" pageId="4" pageNumber="129">C. socotrana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A270E659F237DD7E66023C1FFEAF6D7C" box="[264,347,796,819]" pageId="4" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
can be distinguished from the species occurring in the Arabian Peninsula by the characters given in the key below.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32DD4BBF237DD7E679D3C5FFEAE6DDC" pageId="4" pageNumber="129" type="etymology">
|
|||
|
|
<paragraph id="8B888730F237DD7E679D3C5FFEAE6DDC" blockId="4.[151,1127,220,947]" pageId="4" pageNumber="129">
|
|||
|
|
<emphasis id="B9435B22F237DD7E679D3C5FFEEC6D3C" bold="true" box="[151,280,860,883]" pageId="4" pageNumber="129">Etymology.</emphasis>
|
|||
|
|
The species epithet is a Latin adjective derived from the place of origin and means ‘Socotran’.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32DD4BBF237DD7E679D3C9FFD5C6DFC" box="[151,680,924,947]" pageId="4" pageNumber="129" type="distribution">
|
|||
|
|
<paragraph id="8B888730F237DD7E679D3C9FFD5C6DFC" blockId="4.[151,1127,220,947]" box="[151,680,924,947]" pageId="4" pageNumber="129">
|
|||
|
|
<emphasis id="B9435B22F237DD7E679D3C9FFEDE6DFC" bold="true" box="[151,298,924,947]" pageId="4" pageNumber="129">Distribution.</emphasis>
|
|||
|
|
Endemic to
|
|||
|
|
<collectingCountry id="F320C7A0F237DD7E66BB3C9FFDB96DFC" box="[433,589,924,947]" name="Socotra Island" pageId="4" pageNumber="129">Socotra Island</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F320C7A0F237DD7E655C3C9FFD506DFC" box="[598,676,924,947]" name="Yemen" pageId="4" pageNumber="129">Yemen</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|