<documentid="6CC2E01E0E4A9015C39A247E0430C682"ID-CLB-Dataset="298093"ID-DOI="10.5852/ejt.2024.935.2555"ID-GBIF-Dataset="2f811f90-7810-459b-8230-2a09ee67aa07"ID-ISSN="2118-9773"ID-Zenodo-Dep="11400806"ID-ZooBank="36F07FD1-09B8-4057-86F6-5F6960DC799B"IM.bibliography_approvedBy="jonas"IM.illustrations_approvedBy="jonas"IM.materialsCitations_approvedBy="jonas"IM.metadata_approvedBy="jonas"IM.taxonomicNames_approvedBy="jonas"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="jonas"checkinTime="1717145828877"checkinUser="plazi"docAuthor="Bucher, Manon, Bouteille, Emilien & Cesne, Maxime Le"docDate="2024"docId="E8713702D118FFAB90E3B7CDFA8AF979"docLanguage="en"docName="EJT.2024.935.166-202.pdf"docOrigin="European Journal of Taxonomy 935"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Amberana Distant 1908"docType="key"docVersion="4"lastPageNumber="175"masterDocId="14484F7AD110FFA2905EB56AFFFDFFE9"masterDocTitle="Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar"masterLastPageNumber="202"masterPageNumber="166"pageNumber="174"updateTime="1717617949413"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="AB50F1DB602F8DECC70AA6EBF07FEF89">Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar</mods:title>
<bibRefCitationid="0449FBE5D118FFAA926CB7CDFD1FFD2A"author="Distant W. L."box="[562,738,679,707]"firstAuthor="Distant"journalOrPublisher="Annals and Magazine of Natural History"pageId="8"pageNumber="174"pagination="309 - 323"part="8"refId="ref15002"refString="Distant W. L. 1908. Rhynchotal notes xlv. Annals and Magazine of Natural History (8) 2: 309 - 323. https: // doi. org / 10.1080 / 00222930808692488"title="Rhynchotal notes xlv"type="journal article"year="1908">Distant, 1908</bibRefCitation>
<keyLeadid="DB292521D118FFAA909FB795FA8AFCF1"box="[193,1399,766,793]"pageId="8"pageNumber="174">– Tegmina other than with a red or orange dominant colour ............................................................... 9</keyLead>
<keyLeadid="DB292521D118FFAA90E3B62AFA8AFCB3"box="[189,1399,831,858]"pageId="8"pageNumber="174">2. Thorax with uniform colour, red or black ......................................................................................... 3</keyLead>
<keyLeadid="DB292521D118FFAA909FB609FA8AFC94"box="[193,1399,867,893]"pageId="8"pageNumber="174">– Thorax with more than one colour .................................................................................................... 4</keyLead>
<keyLeadid="DB292521D118FFAA90E3B6CEFA8AFC56"box="[189,1399,932,959]"pageId="8"pageNumber="174">3. Thorax black, total length less than 11.0 mm ................................................................................... 5</keyLead>
<keyLeadid="DB292521D118FFAA909FB6A2FA8AFC0B"box="[193,1399,967,994]"pageId="8"pageNumber="174">– Thorax red, total length longer than 11.0 mm .................................................................................. 7</keyLead>
<bibRefCitationid="0449FBE5D118FFAA94FAB163FA93FBCA"author="Lallemand V."box="[1188,1390,1033,1059]"firstAuthor="Lallemand"journalOrPublisher="Annales de la Societe entomologique de France"pageId="8"pageNumber="174"pagination="280 - 288"part="88"refId="ref15265"refString="Lallemand V. 1920. Cercopides nouveaux de Madagascar. Annales de la Societe entomologique de France 88: 280 - 288."title="Cercopides nouveaux de Madagascar"type="journal article"year="1920">Lallemand, 1920</bibRefCitation>
– Thorax coloured coral with apex of scutellum black .......................................................................... .................................................................
<taxonomicNameid="A7D8FD97D118FFAA9290B125FA8AFB80"authority="Le Cesne & Soulier-Perkins, 2022"authorityName="Le Cesne & Soulier-Perkins"authorityYear="2022"box="[718,1399,1103,1130]"class="Insecta"family="Cercopidae"genus="Amberana"kingdom="Animalia"order="Hemiptera"pageId="8"pageNumber="174"phylum="Arthropoda"rank="species"species="harandranomasina">
<bibRefCitationid="0449FBE5D118FFAA93BAB13AFA8AFB80"author="Le Cesne M. & Soulier-Perkins A."box="[996,1399,1103,1130]"firstAuthor="Le Cesne"journalOrPublisher="Annales de la societe entomologique de France (N. S.)"pageId="8"pageNumber="174"refId="ref15367"refString="Le Cesne M. & Soulier-Perkins A. 2022. Two new species of Amberana Distant, 1908 (Hemiptera: Cicadomorpha: Cercopidae) from Madagascar. Annales de la societe entomologique de France (N. S.). https: // doi. org / 10.1080 / 00379271.2022.2110515"title="Two new species of Amberana Distant, 1908 (Hemiptera: Cicadomorpha: Cercopidae) from Madagascar"type="book"year="2022">Le Cesne & Soulier-Perkins, 2022</bibRefCitation>
<keyLeadid="DB292521D118FFAA90E3B1FAFA8AFB42"box="[189,1399,1168,1195]"pageId="8"pageNumber="174">5. Total length not exceeding 8.0 mm ................................................................................................... 6</keyLead>
– Total length longer than 9.0 mm ............................
<taxonomicNameid="A7D8FD97D118FFAA931FB1DEFA8AFB27"authority="Soulier-Perkins & Le Cesne, 2022"authorityName="Soulier-Perkins & Le Cesne"authorityYear="2022"box="[833,1399,1204,1230]"class="Insecta"family="Cercopidae"genus="Amberana"kingdom="Animalia"order="Hemiptera"pageId="8"pageNumber="174"phylum="Arthropoda"rank="species"species="peyrierasi">
<bibRefCitationid="0449FBE5D118FFAA94E1B1DEFA8AFB27"author="Le Cesne M. & Soulier-Perkins A."box="[1215,1399,1204,1230]"firstAuthor="Le Cesne"journalOrPublisher="Annales de la societe entomologique de France (N. S.)"pageId="8"pageNumber="174"refId="ref15367"refString="Le Cesne M. & Soulier-Perkins A. 2022. Two new species of Amberana Distant, 1908 (Hemiptera: Cicadomorpha: Cercopidae) from Madagascar. Annales de la societe entomologique de France (N. S.). https: // doi. org / 10.1080 / 00379271.2022.2110515"title="Two new species of Amberana Distant, 1908 (Hemiptera: Cicadomorpha: Cercopidae) from Madagascar"type="book"year="2022">Le Cesne, 2022</bibRefCitation>
6. Black border following costal, apical and sutural margins of tegmina, from tip of clavus black border get narrower and follows shortly clavo-corial suture. Sometimes, a black patch can extend from the tip of clavus down to the costal margin .................................................
– Black border on tegmina, starting only from tip of clavus, following apical margin and stopping at mid length of costal margin .....................................................................
7. Black border starting from tip of clavus and following apical and costal margins of tegmina .......... ................................................................................................................
<keyLeadid="DB292521D118FFAA909FB360FA8AF9CD"box="[193,1399,1546,1572]"pageId="8"pageNumber="174">– Black line running along clavo-corial suture ................................................................................... 8</keyLead>
8. Black border all around tegmina, thin black line on clavo-corial suture and a black line on corium starting after M + Cu split and finishing before reaching apex ..................
<bibRefCitationid="0449FBE5D118FFAA94BCB305FA8AF960"author="Jacobi A."box="[1250,1399,1647,1673]"firstAuthor="Jacobi"journalOrPublisher="Voeltzkow's Reise in Ostafrika"pageId="8"pageNumber="174"pagination="519 - 552"part="3"refId="ref15154"refString="Jacobi A. 1917. Die Zikadenfauna Madagascars und der Comoren. Voeltzkow's Reise in Ostafrika 3: 519 - 552."title="Die Zikadenfauna Madagascars und der Comoren"type="journal article"year="1917">Jacobi, 1917</bibRefCitation>
– Larger black border around tegmina but finishing before last third of costal margin, black line on clavo-corial suture large, line on corium not individualized ......................
<bibRefCitationid="0449FBE5D118FFAA94BCB3DFFA8AF926"author="Jacobi A."box="[1250,1399,1717,1743]"firstAuthor="Jacobi"journalOrPublisher="Voeltzkow's Reise in Ostafrika"pageId="8"pageNumber="174"pagination="519 - 552"part="3"refId="ref15154"refString="Jacobi A. 1917. Die Zikadenfauna Madagascars und der Comoren. Voeltzkow's Reise in Ostafrika 3: 519 - 552."title="Die Zikadenfauna Madagascars und der Comoren"type="journal article"year="1917">Jacobi, 1917</bibRefCitation>
<bibRefCitationid="0449FBE5D118FFAA94E0B231FA93F89C"author="Signoret V."box="[1214,1390,1883,1909]"firstAuthor="Signoret"journalOrPublisher="Annales de la Societe entomologique de France"pageId="8"pageNumber="174"pagination="177 - 206"part="8"refId="ref15741"refString="Signoret V. 1860. Faune des hemipteres de Madagascar. 1 ere partie. Homopteres. Annales de la Societe entomologique de France 8 (3): 177 - 206."title="Faune des hemipteres de Madagascar. 1 ere partie. Homopteres"type="journal article"year="1860">Signoret, 1860</bibRefCitation>
<keyLeadid="DB292521D118FFAA909FB215FA8BF871"box="[193,1398,1918,1945]"pageId="8"pageNumber="174">– Black border following apical margin of tegmina ...........................................................................11</keyLead>
11. Tegmina translucent with an orange patch along clavus basal margin and red area along base of costal margin ..................................................................................
<bibRefCitationid="0449FBE5D119FFAB94FAB446FA93FEAF"author="Lallemand V."box="[1188,1390,300,326]"firstAuthor="Lallemand"journalOrPublisher="Annales de la Societe entomologique de France"pageId="9"pageNumber="175"pagination="280 - 288"part="88"refId="ref15265"refString="Lallemand V. 1920. Cercopides nouveaux de Madagascar. Annales de la Societe entomologique de France 88: 280 - 288."title="Cercopides nouveaux de Madagascar"type="journal article"year="1920">Lallemand, 1920</bibRefCitation>
12. Red patch at base of tegmina plus red spot within internal curve of black apical border; vertex and pronotum red .............................................................................
<bibRefCitationid="0449FBE5D119FFAB94EEB4DEFA8AFE27"author="Lallemand V."box="[1200,1399,436,462]"firstAuthor="Lallemand"journalOrPublisher="Memoires de l'Institut royal des Sciences naturelles de Belgique"pageId="9"pageNumber="175"pagination="1 - 193"part="2"refId="ref15288"refString="Lallemand V. 1949. Revision des Cercopinae (Hemiptera Homoptera) Premiere partie. Memoires de l'Institut royal des Sciences naturelles de Belgique 2 (32): 1 - 193."title="Revision des Cercopinae (Hemiptera Homoptera) Premiere partie"type="journal article"year="1949">Lallemand, 1949</bibRefCitation>
– Orange patch at base of tegmina bordered posteriorly by perpendicular black line that follows clavo-corial suture within orange patch; vertex and scutellum black, pronotum orange ............................. ...................................................................................................................
<bibRefCitationid="0449FBE5D119FFAB9493B774FA93FDD1"author="Distant W. L."box="[1229,1390,542,568]"firstAuthor="Distant"journalOrPublisher="Annals and Magazine of Natural History"pageId="9"pageNumber="175"pagination="309 - 323"part="8"refId="ref15002"refString="Distant W. L. 1908. Rhynchotal notes xlv. Annals and Magazine of Natural History (8) 2: 309 - 323. https: // doi. org / 10.1080 / 00222930808692488"title="Rhynchotal notes xlv"type="journal article"year="1908">Distant, 1908</bibRefCitation>
<keyLeadid="DB292521D119FFAB909FB7E9FA8BFD75"box="[193,1398,642,669]"pageId="9"pageNumber="175">– Pronotum black bearing a yellow to orange transversal band ........................................................ 21</keyLead>
<bibRefCitationid="0449FBE5D119FFAB9429B7AEFA8AFD37"author="Soulier-Perkins A. & Kunz G."box="[1143,1399,708,734]"firstAuthor="Soulier-Perkins"journalOrPublisher="Zootaxa"pageId="9"pageNumber="175"pagination="1 - 42"part="3156"refId="ref15805"refString="Soulier-Perkins A. & Kunz G. 2012. Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus. Zootaxa 3156: 1 - 42. https: // doi. org / 10.11646 / zootaxa. 3156.1.1"title="Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus"type="journal article"year="2012">Soulier-Perkins, 2012</bibRefCitation>
<keyLeadid="DB292521D119FFAB909FB78DFA8BFCEB"box="[193,1398,743,770]"pageId="9"pageNumber="175">– Tegmina mainly dark brown to black ............................................................................................. 15</keyLead>
<keyLeadid="DB292521D119FFAB909FB626FA8BFC8C"box="[193,1398,843,870]"pageId="9"pageNumber="175">– Tegmina dark brown to black, bearing 2 or 3 spots of colour ........................................................ 16</keyLead>
<keyLeadid="DB292521D119FFAB90E3B6E7FA8BFC4E"box="[189,1398,909,935]"pageId="9"pageNumber="175">16. Total length not exceeding 7.0 mm ................................................................................................. 17</keyLead>
<keyLeadid="DB292521D119FFAB909FB6DBFA8BFC23"box="[193,1398,944,971]"pageId="9"pageNumber="175">– Total length longer than 7.0 mm ..................................................................................................... 18</keyLead>
<bibRefCitationid="0449FBE5D119FFAB9429B69BFA8AFBE5"author="Soulier-Perkins A. & Kunz G."box="[1143,1399,1009,1036]"firstAuthor="Soulier-Perkins"journalOrPublisher="Zootaxa"pageId="9"pageNumber="175"pagination="1 - 42"part="3156"refId="ref15805"refString="Soulier-Perkins A. & Kunz G. 2012. Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus. Zootaxa 3156: 1 - 42. https: // doi. org / 10.11646 / zootaxa. 3156.1.1"title="Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus"type="journal article"year="2012">Soulier-Perkins, 2012</bibRefCitation>
<bibRefCitationid="0449FBE5D119FFAB9489B17FFA8AFBD9"author="Synave H."box="[1239,1399,1045,1072]"firstAuthor="Synave"journalOrPublisher="Le Naturaliste Malgache"pageId="9"pageNumber="175"pagination="133 - 140"part="9"refId="ref15870"refString="Synave H. 1957. Contribution a l'etude des Cercopidae de Madagascar (Hemiptera, Homoptera). Le Naturaliste Malgache 9: 133 - 140."title="Contribution a l'etude des Cercopidae de Madagascar (Hemiptera, Homoptera)"type="journal article"year="1957">Synave, 1957</bibRefCitation>
<keyLeadid="DB292521D119FFAB90E3B13CFA8BFB99"box="[189,1398,1110,1137]"pageId="9"pageNumber="175">18. Total length inferior to 11.0 mm ..................................................................................................... 19</keyLead>
<bibRefCitationid="0449FBE5D119FFAB9429B113FA8AFB7D"author="Soulier-Perkins A. & Kunz G."box="[1143,1399,1145,1172]"firstAuthor="Soulier-Perkins"journalOrPublisher="Zootaxa"pageId="9"pageNumber="175"pagination="1 - 42"part="3156"refId="ref15805"refString="Soulier-Perkins A. & Kunz G. 2012. Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus. Zootaxa 3156: 1 - 42. https: // doi. org / 10.11646 / zootaxa. 3156.1.1"title="Revision of the Malagasy endemic genus Amberana Distant (Hemiptera, Cercopidae) with description of one new genus"type="journal article"year="2012">Soulier-Perkins, 2012</bibRefCitation>
<bibRefCitationid="0449FBE5D119FFAB94FAB1D1FA93FB3C"author="Lallemand V."box="[1188,1390,1211,1237]"firstAuthor="Lallemand"journalOrPublisher="Annales de la Societe entomologique de France"pageId="9"pageNumber="175"pagination="280 - 288"part="88"refId="ref15265"refString="Lallemand V. 1920. Cercopides nouveaux de Madagascar. Annales de la Societe entomologique de France 88: 280 - 288."title="Cercopides nouveaux de Madagascar"type="journal article"year="1920">Lallemand, 1920</bibRefCitation>
<bibRefCitationid="0449FBE5D119FFAB9486B1B4FA8AFB11"author="Distant W. L."box="[1240,1399,1246,1272]"firstAuthor="Distant"journalOrPublisher="Annals and Magazine of Natural History"pageId="9"pageNumber="175"pagination="309 - 323"part="8"refId="ref15002"refString="Distant W. L. 1908. Rhynchotal notes xlv. Annals and Magazine of Natural History (8) 2: 309 - 323. https: // doi. org / 10.1080 / 00222930808692488"title="Rhynchotal notes xlv"type="journal article"year="1908">Distant, 1908</bibRefCitation>
20. Tegmina bearing only band of colour along anterior claval margin, rest of tegmina dark brown with some red ........................................................................................
<bibRefCitationid="0449FBE5D119FFAB94FAB029FA93FAB4"author="Lallemand V."box="[1188,1390,1347,1373]"firstAuthor="Lallemand"journalOrPublisher="Annales de la Societe entomologique de France"pageId="9"pageNumber="175"pagination="280 - 288"part="88"refId="ref15265"refString="Lallemand V. 1920. Cercopides nouveaux de Madagascar. Annales de la Societe entomologique de France 88: 280 - 288."title="Cercopides nouveaux de Madagascar"type="journal article"year="1920">Lallemand, 1920</bibRefCitation>
<keyLeadid="DB292521D119FFAB909FB00CFA8BFA4A"pageId="9"pageNumber="175">– Tegmina bearing band of colour along anterior claval margin plus two other spots of same colour ..... ......................................................................................................................................................... 21</keyLead>
<keyLeadid="DB292521D119FFAB90E3B0A1FA8BFA0C"box="[189,1398,1482,1509]"pageId="9"pageNumber="175">21. Clypeus black .................................................................................................................................. 22</keyLead>
<bibRefCitationid="0449FBE5D119FFAB9489B345FA93F9A0"author="Jacobi A."box="[1239,1390,1583,1609]"firstAuthor="Jacobi"journalOrPublisher="Voeltzkow's Reise in Ostafrika"pageId="9"pageNumber="175"pagination="519 - 552"part="3"refId="ref15154"refString="Jacobi A. 1917. Die Zikadenfauna Madagascars und der Comoren. Voeltzkow's Reise in Ostafrika 3: 519 - 552."title="Die Zikadenfauna Madagascars und der Comoren"type="journal article"year="1917">Jacobi, 1917</bibRefCitation>
– Tegmina with two longitudinal yellow lines, one on clavus and one on corium ................................ .........................................................................................
<taxonomicNameid="A7D8FD97D119FFAB93DEB31FFB13F979"authority="(Bergroth, 1894)"authorityName="Bucher & Bouteille & Cesne"authorityYear="2024"baseAuthorityName="Bergroth"baseAuthorityYear="1894"box="[896,1262,1653,1680]"class="Insecta"family="Cercopidae"genus="Amberana"kingdom="Animalia"order="Hemiptera"pageId="9"pageNumber="175"phylum="Arthropoda"rank="species"species="vittipennis"status="comb. nov.">
<bibRefCitationid="0449FBE5D119FFAB946FB31FFB18F979"author="Bergroth E."box="[1073,1253,1653,1680]"firstAuthor="Bergroth"journalOrPublisher="Annales de la Societe entomologique de Belgique, Bruxelles"pageId="9"pageNumber="175"pagination="160 - 161"part="38"refId="ref14862"refString="Bergroth E. 1894. Diagnosis Hemipterorum quatuor madagascariensium. Annales de la Societe entomologique de Belgique, Bruxelles 38: 160 - 161."title="Diagnosis Hemipterorum quatuor madagascariensium"type="journal article"year="1894">Bergroth, 1894</bibRefCitation>