157 lines
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157 lines
20 KiB
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<document id="2202BDAFFC9F8E6984D899EAF3C0008B" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D5852FFF9C5991FEDE5675F99B" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) alpercata Silva, Louzada & Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="480" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="480" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="3DC2E56B877DAF52618CA77FCB31F2CB">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="CFFF75F531409D33803E2F35E52C30F7">Silva, Fernando A. B.</mods:namePart>
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</mods:role>
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<mods:namePart id="CDAE553E5A0D0CE457655BB4C1100A0D">Louzada, Júlio</mods:namePart>
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</mods:name>
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<mods:name id="46D85A00602FE5C8D090CDCC74D93814" type="personal">
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<mods:role id="A14B55B5505713AF8FE07D3F595ACD58">
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<mods:roleTerm id="1A323D38168165293CA1E21EEBE77656">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="46DB10D003C034BE58F6F68073D748CD">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:typeOfResource id="83424496BBFA25B761EB434FEE3123F7">text</mods:typeOfResource>
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<mods:title id="2C4EED0F37714FFDAAAD10F74375239D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A93BBE288A959A6A513E5995C5901A3C">
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<mods:date id="39112AA8F1DBB48D9D349876D8A256BA">2015</mods:date>
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<mods:number id="85B8B11BE0AA126E54DF6C50BBB5F95F">3925</mods:number>
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<mods:detail id="637A317FB2300B34BB087507599DDB92" type="issue">
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<mods:number id="1511C5E987DC9034E14FF590A6CE2DA1">4</mods:number>
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<mods:end id="DD1551221C7B9F02D321AC99A06C1212">504</mods:end>
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<mods:classification id="14571CCD116AC45FF3D153041F446C95">journal article</mods:classification>
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<mods:identifier id="05E818AEE3B30CCFDF279813A9D2C7F9" type="DOI">10.11646/zootaxa.3925.4.1</mods:identifier>
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<treatment id="03DE87D5852FFF9C5991FEDE5675F99B" ID-DOI="http://doi.org/10.5281/zenodo.6115736" ID-GBIF-Taxon="119598940" ID-Zenodo-Dep="6115736" LSID="urn:lsid:plazi:treatment:03DE87D5852FFF9C5991FEDE5675F99B" httpUri="http://treatment.plazi.org/id/03DE87D5852FFF9C5991FEDE5675F99B" lastPageNumber="480" pageId="29" pageNumber="480">
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<subSubSection id="C36D6548852FFF9C5991FEDE54FDFEB7" pageId="29" pageNumber="480" type="nomenclature">
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<paragraph id="8BC836C3852FFF9C5991FEDE57B8FE96" blockId="29.[151,759,295,355]" box="[151,759,295,322]" pageId="29" pageNumber="480">
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<heading id="D08081AF852FFF9C5991FEDE57B8FE96" bold="true" box="[151,759,295,322]" fontSize="11" level="1" pageId="29" pageNumber="480" reason="1">
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<emphasis id="B903EAD1852FFF9C5991FEDE57B8FE96" bold="true" box="[151,759,295,322]" pageId="29" pageNumber="480">
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<taxonomicName id="4C774D40852FFF9C5991FEDE572FFE95" ID-CoL="6CJ63" box="[151,608,295,322]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="alpercata" status="sp. nov." subGenus="Aganhyboma">
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<emphasis id="B903EAD1852FFF9C5991FEDE5462FE95" bold="true" box="[151,301,295,321]" italics="true" pageId="29" pageNumber="480">Deltochilum</emphasis>
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(
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<emphasis id="B903EAD1852FFF9C583BFEDE5491FE95" bold="true" box="[317,478,295,321]" italics="true" pageId="29" pageNumber="480">Aganhyboma</emphasis>
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)
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<emphasis id="B903EAD1852FFF9C58F6FEDE572FFE95" bold="true" box="[496,608,295,321]" italics="true" pageId="29" pageNumber="480">alpercata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057AA852FFF9C5B61FED157B8FE96" box="[615,759,296,322]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836C3852FFF9C5991FEB354FDFEB7" blockId="29.[151,759,295,355]" box="[151,434,330,355]" pageId="29" pageNumber="480">
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(
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<figureCitation id="134C2A46852FFF9C5999FEB355BAFEB7" box="[159,245,330,355]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="29" pageNumber="480">Figs 10</figureCitation>
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, 77, 79, 92, 115)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852FFF9C5991FE6B545CFE1B" pageId="29" pageNumber="480" type="etymology">
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<paragraph id="8BC836C3852FFF9C5991FE6B545CFE1B" blockId="29.[151,1437,402,1615]" pageId="29" pageNumber="480">
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<emphasis id="B903EAD1852FFF9C5991FE6B5451FE7F" bold="true" box="[151,286,402,427]" pageId="29" pageNumber="480">Etymology.</emphasis>
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“
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<taxonomicName id="4C774D40852FFF9C5832FE6B54D1FE7F" box="[308,414,402,427]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="alpercata">alpercata</taxonomicName>
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" is a typical footwear of the northeastern region of
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<collectingCountry id="F3607653852FFF9C5D1CFE6B512DFE7F" box="[1050,1122,402,427]" name="Brazil" pageId="29" pageNumber="480">Brazil</collectingCountry>
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. The epithet is a name in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852FFF9C59C1FE2354F5FB17" pageId="29" pageNumber="480" type="description">
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<paragraph id="8BC836C3852FFF9C59C1FE2354F5FB17" blockId="29.[151,1437,402,1615]" pageId="29" pageNumber="480">
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<emphasis id="B903EAD1852FFF9C59C1FE2354EAFE27" bold="true" box="[199,421,474,499]" pageId="29" pageNumber="480">Description. Body.</emphasis>
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Head, pronotum, elytra, and ventral surface black, with a silky sheen. Length>
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<quantity id="4C8F9B26852FFF9C5C43FE2250D7FE26" box="[1349,1432,475,499]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="480" unit="mm" value="14.0">14 mm</quantity>
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.
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<emphasis id="B903EAD1852FFF9C5991FE075591FDC3" bold="true" box="[151,222,510,535]" pageId="29" pageNumber="480">Head.</emphasis>
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Punctures extremely dense and touching; around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae, or with almost inconspicuous setae. Dorsal interocular distance approximately three times eye width (as in Fig. 61).
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<emphasis id="B903EAD1852FFF9C5D78FDBF5191FD8B" bold="true" box="[1150,1246,582,607]" pageId="29" pageNumber="480">Thorax.</emphasis>
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Pronotum black with an opaque sheen or a weak luster (
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<figureCitation id="134C2A46852FFF9C5B6EFD935783FD57" box="[616,716,618,643]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="29" pageNumber="480">Fig. 115</figureCitation>
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). Surface of pronotum with dense foveiform punctures, nearly touching on the sides, anterior angles and posterior portion. Discal punctures somewhat smaller and more dispersed than those of lateral portion. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by less than one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles approximately straight. Hypomera with dense foveiform punctures on anterior and posterior portions (Fig. 79); anteromedial punctures separated by less than one diameter (Fig. 79).
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<emphasis id="B903EAD1852FFF9C587BFCBB549FFC8F" bold="true" box="[381,464,834,859]" pageId="29" pageNumber="480">Elytra.</emphasis>
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Rounded, sides bulging outward (
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<figureCitation id="134C2A46852FFF9C5A60FCBB5686FC8F" box="[870,969,834,859]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="29" pageNumber="480">Fig. 115</figureCitation>
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) and surface slightly irregular (rugose). Elytral striae narrow, carinate margins close and almost touching (as in Figs 74, 77). Interstriae flattened, with foveiform punctures surrounded by four or five shiny microtubercles. Foveiform punctures separated on elytral disc by less than one diameter. Microtubercles of interstriae well delimited and conspicuous, with strong luster; interestrial surface around microtubercles opaque (Fig. 77). Basal carina of seventh interstria short, length about twice the width of the seventh interestria. Basal carina of ninth interstria present in three-quarters of elytra, but it never reaches the anterior limit of apical carina of seventh interstria. Apical carina of third interstria more prominent than apical carinae of second and fourth interstriae. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria less than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852FFF9C59C1FB37507DFADF" pageId="29" pageNumber="480" type="materials_examined">
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<paragraph id="8BC836C3852FFF9C59C1FB3750DEFB33" blockId="29.[151,1437,402,1615]" box="[199,1425,1230,1255]" pageId="29" pageNumber="480">
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<emphasis id="B903EAD1852FFF9C59C1FB3757CCFB33" bold="true" box="[199,643,1230,1255]" pageId="29" pageNumber="480">
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Material studied.
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<typeStatus id="54CC8861852FFF9C589DFB375743FB33" box="[411,524,1230,1255]" pageId="29" pageNumber="480" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3607653852FFF9C5B1EFB3757CCFB33" box="[536,643,1230,1255]" name="Brazil" pageId="29" pageNumber="480">BRAZIL</collectingCountry>
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</emphasis>
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: ALAGOAS: Murici, S. Branca,
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<date id="FFC91003852FFF9C5AFAFB365100FB33" box="[1020,1103,1231,1255]" pageId="29" pageNumber="480" value="1984-05">V.1984</date>
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, F.M. Oliveira (
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<specimenCount id="9D71FD4A852FFF9C5C05FB365064FB32" box="[1283,1323,1231,1255]" pageId="29" pageNumber="480" type="generic">1 ♂</specimenCount>
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<collectionCode id="ED66AE06852FFF9C5C34FB3650CAFB33" box="[1330,1413,1231,1255]" pageId="29" pageNumber="480">CEMT</collectionCode>
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).
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</paragraph>
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<paragraph id="8BC836C3852FFF9C59C1FB0B507DFADF" blockId="29.[151,1437,402,1615]" box="[199,1330,1266,1291]" pageId="29" pageNumber="480">
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<emphasis id="B903EAD1852FFF9C59C1FB0B5465FADF" bold="true" box="[199,298,1266,1291]" pageId="29" pageNumber="480">Habitat.</emphasis>
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<collectingCountry id="F3607653852FFF9C5837FB0B543AFADF" box="[305,373,1266,1291]" name="Brazil" pageId="29" pageNumber="480">Brazil</collectingCountry>
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(Alagoas) (
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<figureCitation id="134C2A46852FFF9C58FCFB0B5702FADF" box="[506,589,1266,1291]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="29" pageNumber="480">Fig. 92</figureCitation>
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), from available data this species inhabits lowland Atlantic forest.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6548852FFF9C59C1FAEF5675F99B" pageId="29" pageNumber="480" type="diagnosis">
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<paragraph id="8BC836C3852FFF9C59C1FAEF570CF9D3" blockId="29.[151,1437,402,1615]" pageId="29" pageNumber="480">
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<emphasis id="B903EAD1852FFF9C59C1FAEF5493FAFB" bold="true" box="[199,476,1302,1327]" pageId="29" pageNumber="480">Diagnosis and remarks</emphasis>
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(within the
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<taxonomicName id="4C774D40852FFF9C5B61FAEE57A1FAFB" box="[615,750,1303,1327]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="finestriatum">
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<emphasis id="B903EAD1852FFF9C5B61FAEE57A1FAFB" box="[615,750,1303,1327]" italics="true" pageId="29" pageNumber="480">finestriatum</emphasis>
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</taxonomicName>
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complex): elytra rounded, sides bulging outward (
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<figureCitation id="134C2A46852FFF9C5C2BFAEF50C0FAFB" box="[1325,1423,1302,1327]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="29" pageNumber="480">Fig. 115</figureCitation>
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); punctures of anteromedial portion of hypomera separated by less than one diameter (Fig. 79); pronotum black with an opaque sheen or a weak luster (
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<figureCitation id="134C2A46852FFF9C5B24FAA757CBFAA3" box="[546,644,1374,1399]" captionStart="FIGURES 111 – 118" captionStartId="50.[151,264,1952,1975]" captionTargetBox="[201,1409,198,1924]" captionTargetId="figure@50.[175,1412,193,1931]" captionTargetPageId="50" captionText="FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244809/files/figure.png" pageId="29" pageNumber="480">Fig. 115</figureCitation>
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); elytral microtubercles well delimited, with a brown sheen (Fig. 77); interestrial surface around microtubercles opaque (Fig. 77). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, in lateral view forming an angle of approximately 45° with phallobase; apex rounded (
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<figureCitation id="134C2A46852FFF9C5801FA33542FFA37" box="[263,352,1482,1507]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="29" pageNumber="480">Fig. 10</figureCitation>
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). Lamella copulatrix (LC): left lobe subtrapezoidal; right lobe subrectangular. Fronto-lateral peripheral sclerite (FLP) as in Fig. 53.
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</paragraph>
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<paragraph id="8BC836C3852FFF9C59C1F9EB5675F99B" blockId="29.[151,1437,402,1615]" pageId="29" pageNumber="480">
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|
Although the aedeagus of
|
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|
<taxonomicName id="4C774D40852FFF9C58F2F9EA57CDF9FF" box="[500,642,1555,1579]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="alpercata" status="sp. nov.">
|
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|
<emphasis id="B903EAD1852FFF9C58F2F9EA57CDF9FF" box="[500,642,1555,1579]" italics="true" pageId="29" pageNumber="480">D. alpercata</emphasis>
|
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|
</taxonomicName>
|
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|
<emphasis id="B903EAD1852FFF9C5B8DF9EB5659F9FF" bold="true" box="[651,790,1554,1579]" pageId="29" pageNumber="480">
|
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|
<taxonomicNameLabel id="A23057AA852FFF9C5B8DF9EB5659F9FF" box="[651,790,1554,1579]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
|
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|
</emphasis>
|
|||
|
and
|
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|
<taxonomicName id="4C774D40852FFF9C5A57F9EA5692F9FF" box="[849,989,1555,1579]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="cangalha" status="sp. nov.">
|
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|
<emphasis id="B903EAD1852FFF9C5A57F9EA5692F9FF" box="[849,989,1555,1579]" italics="true" pageId="29" pageNumber="480">D. cangalha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="B903EAD1852FFF9C5AE0F9EB513DF9FF" bold="true" box="[998,1138,1554,1579]" pageId="29" pageNumber="480">
|
|||
|
<taxonomicNameLabel id="A23057AA852FFF9C5AE0F9EB513DF9FF" box="[998,1138,1554,1579]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
|
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|
</emphasis>
|
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|
do not present significant difference, the elytral punctures in these species are distinct.
|
|||
|
</paragraph>
|
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|
</subSubSection>
|
|||
|
</treatment>
|
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|
</document>
|