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<document id="10D7B64EE2F0BC28E71185AD60940FA9" ID-CLB-Dataset="25355" ID-DOI="10.11646/zootaxa.4679.1.3" ID-GBIF-Dataset="8b3957bc-d1bb-4d32-a0fe-761ff31060ed" ID-ISSN="1175-5326" ID-Zenodo-Dep="3470037" ID-ZooBank="1CC05FB0-4EDA-4779-874E-8EFF58D6B228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569910544550" checkinUser="plazi" docAuthor="Belousov, Igor A. &amp; Kabak, Ilya I." docDate="2019" docId="7B4987F3FFA2043868D5F4BE9431FE28" docLanguage="en" docName="zootaxa.4679.1.3.pdf" docOrigin="Zootaxa 4679 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus torgaut Belousov &amp; Kabak 2019, sp. n." docType="treatment" docVersion="10" lastPageNumber="55" masterDocId="8770FF8BFFA404306842F70F902AFFA3" masterDocTitle="New species of the genus Trechus Clairville, 1806 from the Chinese Tien Shan (Coleoptera: Carabidae)" masterLastPageNumber="68" masterPageNumber="47" pageNumber="53" updateTime="1698756597006" updateUser="ExternalLinkService">
<mods:mods id="52541410232D2E454F5325F7B09AE22D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CDD873465B048554B3EB8FF86FBFDE0A">
<mods:title id="6DCE8677FE9E60B2FF965522EDFF2E16">New species of the genus Trechus Clairville, 1806 from the Chinese Tien Shan (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="7153B5A7413B8C2BE435229B8BFC0D92" type="personal">
<mods:role id="E7303EA4C89D4556D17E1C896129C689">
<mods:roleTerm id="2A6681489D67748AF4E6E481B37C5F6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24E08924FF84C24476F6A3CBF0894CFA">Belousov, Igor A.</mods:namePart>
</mods:name>
<mods:name id="562EC67EC10DEC2BB789167B47065B83" type="personal">
<mods:role id="BD5590A6ED593F033FB0807616F6695F">
<mods:roleTerm id="CE1E7BF5CD19A5487D2DA9F32E1F7A06">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B74C612CBAA4C3E20C77D172D811FFF">Kabak, Ilya I.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B9A91B455D157BA225F1A29E83DA39CC">text</mods:typeOfResource>
<mods:relatedItem id="DCC9BFE9C751DE5F1F7187D4208B8470" type="host">
<mods:titleInfo id="6042B9599CC824E6621D8C3461609430">
<mods:title id="D68DE9E0C9349C080BC2B180C5035AB0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B8AB33884D4F0D7EB1D40D299B2F7F9E">
<mods:date id="6E869DCA28F50172B888663F42DD7767">2019</mods:date>
<mods:detail id="B2A42A847EA742BEF057DF70860871CB" type="pubDate">
<mods:number id="94B47EBCAE9959B01D3648A149BDF046">2019-10-01</mods:number>
</mods:detail>
<mods:detail id="77DCC3FF02B08B7E77DCB83E18941289" type="volume">
<mods:number id="6AD82F7D8821CED60EEC584B04D36D97">4679</mods:number>
</mods:detail>
<mods:detail id="21486C1EDE9A100F2AF8E7CFFEBF903C" type="issue">
<mods:number id="A6CCBD36181612057D854D2202F22DE1">1</mods:number>
</mods:detail>
<mods:extent id="9DBF0FA53FD23299F0A5BA3573BD4700" unit="page">
<mods:start id="7CB15766C2095C5961C74FF387D28CBF">47</mods:start>
<mods:end id="C6F887F6B95A78DD6FCFE226F7D4AB68">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ADF02E171A0EC9BEC99D79ABFB8A1DCF">journal article</mods:classification>
<mods:identifier id="E1CD31C3CE9BA859178276CD36C7A9DF" type="CLB-Dataset">25355</mods:identifier>
<mods:identifier id="9A0A8F621C1E4EA4AF6AC20A6D44ACCD" type="DOI">10.11646/zootaxa.4679.1.3</mods:identifier>
<mods:identifier id="06E0E79510A475D3C09E4DA8B407EBBC" type="GBIF-Dataset">8b3957bc-d1bb-4d32-a0fe-761ff31060ed</mods:identifier>
<mods:identifier id="6E4A9D2A507BC99C897046DA5E17B3CE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="308CD90852E769447630B95B1188B2E6" type="Zenodo-Dep">3470037</mods:identifier>
<mods:identifier id="D138342301FBF1D7BE4814B24925189F" type="ZooBank">1CC05FB0-4EDA-4779-874E-8EFF58D6B228</mods:identifier>
</mods:mods>
<treatment id="7B4987F3FFA2043868D5F4BE9431FE28" ID-GBIF-Taxon="159609600" LSID="urn:lsid:plazi:treatment:7B4987F3FFA2043868D5F4BE9431FE28" httpUri="http://treatment.plazi.org/id/7B4987F3FFA2043868D5F4BE9431FE28" lastPageId="8" lastPageNumber="55" pageId="6" pageNumber="53">
<subSubSection id="BBFA656EFFA2043668D5F4BE92B7FC68" box="[151,669,945,972]" pageId="6" pageNumber="53" type="nomenclature">
<paragraph id="F35F36E5FFA2043668D5F4BE92B7FC68" blockId="6.[151,669,945,1007]" box="[151,669,945,972]" pageId="6" pageNumber="53">
<heading id="A8178189FFA2043668D5F4BE92B7FC68" bold="true" box="[151,669,945,972]" fontSize="11" level="1" pageId="6" pageNumber="53" reason="1">
<emphasis id="C194EAF7FFA2043668D5F4BE92B7FC68" bold="true" box="[151,669,945,972]" pageId="6" pageNumber="53">
<taxonomicName id="34E04D66FFA2043668D5F4BE9264FC68" ID-CoL="8MKMN" authority="Belousov &amp; Kabak" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[151,590,945,972]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA2043668D5F4BE9173FC68" bold="true" box="[151,345,945,972]" italics="true" pageId="6" pageNumber="53">Trechus torgaut</emphasis>
Belousov &amp; Kabak
</taxonomicName>
,
<taxonomicNameLabel id="DAA7578CFFA204366A19F4BE92B7FC68" box="[603,669,945,971]" pageId="6" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBFA656EFFA2043668D5F4DA9138FC4C" box="[151,274,981,1007]" pageId="6" pageNumber="53" type="description">
<paragraph id="F35F36E5FFA2043668D5F4DA9138FC4C" blockId="6.[151,669,945,1007]" box="[151,274,981,1007]" pageId="6" pageNumber="53">
(
<figureCitation id="6BDB2A60FFA2043668DDF4DA90C5FC4C" box="[159,239,981,1007]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1686,1711]" captionTargetBox="[167,1427,390,1654]" captionTargetId="figure@5.[151,1435,373,1666]" captionTargetPageId="5" captionText="FIGURES 1–2. Trechus Clairville, 1806, habitus, males, paratypes. 1, T. tsanmensis sp. n.; 2, T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470039" httpUri="https://zenodo.org/record/3470039/files/figure.png" pageId="6" pageNumber="53">Figs. 2</figureCitation>
,
<figureCitation id="6BDB2A60FFA2043668B9F4DA9120FC4C" box="[251,266,981,1007]" captionStart="FIGURES 3–4" captionStartId="12.[151,264,1957,1982]" captionTargetBox="[210,1372,183,1923]" captionTargetId="figure@12.[200,1387,166,1937]" captionTargetPageId="12" captionText="FIGURES 3–4. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view, c—paramere. 3—T. tsanmensis sp. n.; 4—T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470041" httpUri="https://zenodo.org/record/3470041/files/figure.png" pageId="6" pageNumber="53">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BBFA656EFFA2043668D5F3129451FAD8" pageId="6" pageNumber="53" type="materials_examined">
<paragraph id="F35F36E5FFA2043668D5F31292DBFA97" blockId="6.[151,1437,1052,2015]" pageId="6" pageNumber="53">
<emphasis id="C194EAF7FFA2043668D5F3129161FB94" bold="true" box="[151,331,1052,1079]" pageId="6" pageNumber="53">Type material:</emphasis>
<materialsCitation id="43883CB8FFA204366914F3129252FBDC" ID-GBIF-Occurrence="2423311293" collectingDate="2001-07-13" collectionCode="ZISP" collectorName="Kabak I. I." country="China" county="Kunges River" elevation="2800" latitude="43.1875" location="Pass" longLatPrecision="19" longitude="84.78889" municipality="Adunkur" pageId="6" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Xinjiang" typeStatus="holotype">
<emphasis id="C194EAF7FFA204366914F31291E9FB94" bold="true" box="[342,451,1053,1079]" pageId="6" pageNumber="53">
<typeStatus id="2C5B8847FFA204366914F31291E9FB94" box="[342,451,1053,1079]" pageId="6" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
: 1(1)
<specimenCount id="E5E6FD6CFFA204366A43F3129230FB94" box="[513,538,1053,1079]" pageId="6" pageNumber="53" type="male">♂</specimenCount>
, “
<collectingCountry id="8BF77675FFA204366A77F3129257FB94" box="[565,637,1053,1079]" name="China" pageId="6" pageNumber="53">China</collectingCountry>
,
<collectingRegion id="3124F807FFA204366ACEF31292DBFB94" box="[652,753,1053,1079]" country="China" name="Xinjiang Uygur" pageId="6" pageNumber="53">Xinjiang</collectingRegion>
, sources of
<collectingCounty id="1A3E4E69FFA204366BC3F3129408FB94" box="[897,1058,1053,1079]" pageId="6" pageNumber="53">Kunges River</collectingCounty>
, left bank near
<collectingMunicipality id="133BAC9FFFA204366C9EF312956AFB94" box="[1244,1344,1053,1079]" pageId="6" pageNumber="53">Adunkur</collectingMunicipality>
1
<location id="F63F603EFFA204366D21F31295BDFB94" LSID="urn:lsid:plazi:treatment:7B4987F3FFA2043868D5F4BE9431FE28:F63F603EFFA204366D21F31295BDFB94" box="[1379,1431,1053,1079]" country="China" county="Kunges River" latitude="43.1875" longLatPrecision="19" longitude="84.78889" municipality="Adunkur" name="Pass" pageId="6" pageNumber="53" stateProvince="Xinjiang">Pass</location>
,
<locationDeviation id="E2ED2FC7FFA2043668D5F34E9198FBF8" box="[151,434,1089,1115]" location="Kunges-Linchan" pageId="6" pageNumber="53">ESE of Kunges-Linchan</locationDeviation>
,
<quantity id="34189B00FFA2043669FDF34F9275FBF8" box="[447,607,1088,1115]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="6" pageNumber="53" unit="m" value="2800.0" valueMax="3100.0" valueMin="2500.0">
<elevation id="78CDD1D6FFA2043669FDF34F9275FBF8" box="[447,607,1088,1115]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="6" pageNumber="53" unit="m" value="2800.0" valueMax="3100.0" valueMin="2500.0">2500–3100 m</elevation>
</quantity>
,
<date id="875E1025FFA204366A2CF34E92C4FBF8" box="[622,750,1089,1115]" pageId="6" pageNumber="53" value="2001-07-13">
<collectingDate id="971AE9CDFFA204366A2CF34E92C4FBF8" box="[622,750,1089,1115]" pageId="6" pageNumber="53" value="2001-07-13">13.07.2001</collectingDate>
</date>
,
<collectorName id="5E155333FFA204366ABFF34E935AFBF8" box="[765,880,1089,1115]" pageId="6" pageNumber="53">Kabak I.I.</collectorName>
leg.” [
<geoCoordinate id="96D45022FFA204366B82F34E9473FBF8" box="[960,1113,1089,1115]" degrees="43" direction="north" minutes="12" orientation="latitude" pageId="6" pageNumber="53" precision="15" seconds="00" value="43.2">43º12´00´´ N</geoCoordinate>
/
<geoCoordinate id="96D45022FFA204366C37F34E9500FBF8" box="[1141,1322,1088,1115]" degrees="84" direction="east" minutes="47" orientation="longitude" pageId="6" pageNumber="53" precision="15" seconds="40" value="84.79444">84º47´40´´ E—</geoCoordinate>
43º11´15´´ N / 84º47´20´´ E] (
<collectionCode id="95F1AE20FFA204366935F36A919EFBDC" box="[375,436,1125,1151]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="53">ZISP</collectionCode>
) {
<figureCitation id="6BDB2A60FFA20436698DF36A9231FBDC" box="[463,539,1125,1151]" captionStart="FIGURE 9" captionStartId="15.[151,250,999,1024]" captionTargetBox="[151,1436,174,979]" captionTargetId="figure@15.[151,1436,174,979]" captionTargetPageId="15" captionText="FIGURE 9. Collecting sites of Trechus species of the uygur group. Yellow rhomb—T. sinus Belousov &amp; Kabak, 2001; red rhomb—T. arshanicus Belousov &amp; Kabak, 2001; yellow square—T. uygur ssp. kuytunicus Belousov &amp; Kabak, 2001; red square—T. uygur ssp. uygur Deuve, 1993; yellow triangle—T. torgaut sp. n.; red triangle—T. tsanmensis sp. n. Locality numbers—as in the “Material” section; those for collecting sites of T. arshanicus and T. sinus—see Belousov &amp; Kabak (2001)." figureDoi="http://doi.org/10.5281/zenodo.3470047" httpUri="https://zenodo.org/record/3470047/files/figure.png" pageId="6" pageNumber="53">Fig. 9</figureCitation>
: 22–23
</materialsCitation>
}.
<materialsCitation id="43883CB8FFA204366ACDF36A93F1FB00" ID-GBIF-Occurrence="2423311322" collectingDate="2001-07-13" collectionCode="IZAS, MPU, BMNH, SMNS, ZISP, ZSM, CAG, CAK, CBK, CDW, CJS, CVZ" collectorName="Kabak I. I." country="China" county="Kunges River" elevation="2800" latitude="43.1875" location="Pass" longLatPrecision="19" longitude="84.78889" municipality="Adunkur" pageId="6" pageNumber="53" specimenCount="51" specimenCount-female="50" specimenCount-male="1" stateProvince="Xinjiang Uygur" typeStatus="paratype">
<emphasis id="C194EAF7FFA204366ACDF36A932DFBDC" bold="true" box="[655,775,1125,1151]" pageId="6" pageNumber="53">
<typeStatus id="2C5B8847FFA204366ACDF36A932DFBDC" box="[655,775,1125,1151]" pageId="6" pageNumber="53" type="paratype">Paratypes</typeStatus>
</emphasis>
: 63(5)
<specimenCount id="E5E6FD6CFFA204366B14F36B93A8FBDC" box="[854,898,1124,1151]" pageId="6" pageNumber="53" type="male">♂♂</specimenCount>
,
<specimenCount id="E5E6FD6CFFA204366BD3F36B93F2FBDC" box="[913,984,1124,1151]" pageId="6" pageNumber="53" type="female">50♀♀</specimenCount>
, collected with holotype (
<collectionCode id="95F1AE20FFA204366D4AF36A9560FBDC" box="[1288,1354,1125,1151]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="6" pageNumber="53">IZAS</collectionCode>
,
<collectionCode id="95F1AE20FFA204366D1BF36A95B3FBDC" box="[1369,1433,1125,1151]" country="France" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="6" pageNumber="53" type="Herbarium">MPU</collectionCode>
,
<collectionCode id="95F1AE20FFA2043668D5F38690C4FB00" box="[151,238,1161,1187]" pageId="6" pageNumber="53">BMNH</collectionCode>
,
<collectionCode id="95F1AE20FFA2043668B8F3869163FB00" box="[250,329,1161,1187]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="6" pageNumber="53">SMNS</collectionCode>
,
<collectionCode id="95F1AE20FFA204366914F38691A5FB00" box="[342,399,1161,1187]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="53">ZISP</collectionCode>
,
<collectionCode id="95F1AE20FFA2043669DEF38691F2FB00" box="[412,472,1161,1187]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="53" type="Museum">ZSM</collectionCode>
,
<collectionCode id="95F1AE20FFA2043669A6F3869208FB00" box="[484,546,1161,1187]" pageId="6" pageNumber="53">CAG</collectionCode>
,
<collectionCode id="95F1AE20FFA204366A6DF3869247FB00" box="[559,621,1161,1187]" pageId="6" pageNumber="53">CAK</collectionCode>
,
<collectionCode id="95F1AE20FFA204366A3BF386929FFB00" box="[633,693,1161,1187]" pageId="6" pageNumber="53">CBK</collectionCode>
,
<collectionCode id="95F1AE20FFA204366A80F386932EFB00" box="[706,772,1161,1187]" pageId="6" pageNumber="53">CDW</collectionCode>
,
<collectionCode id="95F1AE20FFA204366B52F386936AFB00" box="[784,832,1161,1187]" pageId="6" pageNumber="53">CJS</collectionCode>
,
<collectionCode id="95F1AE20FFA204366B0EF38693A3FB00" box="[844,905,1161,1187]" pageId="6" pageNumber="53">CVZ</collectionCode>
); 30(4)
</materialsCitation>
<materialsCitation id="43883CB8FFA204366B9EF386914BFAAC" ID-GBIF-Occurrence="2423610541" collectingDate="2001-07-15" collectionCode="IZAS, MPU, BMNH, SMNS, ZISP, ZSM, CAG, CAK, CBK" collectorName="Kabak I. I." country="China" elevation="3100" latitude="43.25" location="Kunges River" longLatPrecision="19" longitude="84.80139" municipality="Tsagan-Usun River" pageId="6" pageNumber="53" specimenCount="46" specimenCount-female="45" specimenCount-male="1" stateProvince="Xinjiang" typeStatus="paratype">
<specimenCount id="E5E6FD6CFFA204366B9EF386942DFB00" box="[988,1031,1161,1187]" pageId="6" pageNumber="53" type="male">♂♂</specimenCount>
,
<specimenCount id="E5E6FD6CFFA204366C51F3879471FB00" box="[1043,1115,1160,1187]" pageId="6" pageNumber="53" type="female">45♀♀</specimenCount>
, “
<collectingCountry id="8BF77675FFA204366C30F3869490FB00" box="[1138,1210,1161,1187]" name="China" pageId="6" pageNumber="53">China</collectingCountry>
,
<collectingRegion id="3124F807FFA204366C85F3869506FB00" box="[1223,1324,1161,1187]" country="China" name="Xinjiang Uygur" pageId="6" pageNumber="53">Xinjiang</collectingRegion>
, left bank of
<collectingMunicipality id="133BAC9FFFA2043668F7F3A291A4FB64" box="[181,398,1197,1223]" pageId="6" pageNumber="53">Tsagan-Usun River</collectingMunicipality>
(right tributary of
<location id="F63F603EFFA204366A23F3A2932AFB6B" LSID="urn:lsid:plazi:treatment:7B4987F3FFA2043868D5F4BE9431FE28:F63F603EFFA204366A23F3A2932AFB6B" box="[609,768,1197,1224]" country="China" latitude="43.25" longLatPrecision="19" longitude="84.80139" municipality="Tsagan-Usun River" name="Kunges River" pageId="6" pageNumber="53" stateProvince="Xinjiang">Kunges River</location>
),
<quantity id="34189B00FFA204366B56F3A2939BFB64" box="[788,945,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="6" pageNumber="53" unit="m" value="3100.0" valueMax="3400.0" valueMin="2800.0">
<elevation id="78CDD1D6FFA204366B56F3A2939BFB64" box="[788,945,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="6" pageNumber="53" unit="m" value="3100.0" valueMax="3400.0" valueMin="2800.0">2800–3400 m</elevation>
</quantity>
,
<date id="875E1025FFA204366BFFF3A39414FB64" box="[957,1086,1196,1223]" pageId="6" pageNumber="53" value="2001-07-15">
<collectingDate id="971AE9CDFFA204366BFFF3A39414FB64" box="[957,1086,1196,1223]" pageId="6" pageNumber="53" value="2001-07-15">15.07.2001</collectingDate>
</date>
,
<collectorName id="5E155333FFA204366C0BF3A29490FB64" box="[1097,1210,1197,1223]" pageId="6" pageNumber="53">Kabak I.I.</collectorName>
leg.” [
<geoCoordinate id="96D45022FFA204366D44F3A395B6FB64" box="[1286,1436,1196,1223]" degrees="43" direction="north" minutes="15" orientation="latitude" pageId="6" pageNumber="53" precision="15" seconds="30" value="43.258335">43º15´30´´ N</geoCoordinate>
/
<geoCoordinate id="96D45022FFA2043668E4F3DE9172FB48" box="[166,344,1232,1259]" degrees="84" direction="east" minutes="47" orientation="longitude" pageId="6" pageNumber="53" precision="15" seconds="30" value="84.791664">84º47´30´´ E—</geoCoordinate>
43º15´00´´ N / 84º48´05´´ E] (
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,
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,
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) {
<figureCitation id="6BDB2A60FFA204366D29F3DE9082FAAC" captionStart="FIGURE 9" captionStartId="15.[151,250,999,1024]" captionTargetBox="[151,1436,174,979]" captionTargetId="figure@15.[151,1436,174,979]" captionTargetPageId="15" captionText="FIGURE 9. Collecting sites of Trechus species of the uygur group. Yellow rhomb—T. sinus Belousov &amp; Kabak, 2001; red rhomb—T. arshanicus Belousov &amp; Kabak, 2001; yellow square—T. uygur ssp. kuytunicus Belousov &amp; Kabak, 2001; red square—T. uygur ssp. uygur Deuve, 1993; yellow triangle—T. torgaut sp. n.; red triangle—T. tsanmensis sp. n. Locality numbers—as in the “Material” section; those for collecting sites of T. arshanicus and T. sinus—see Belousov &amp; Kabak (2001)." figureDoi="http://doi.org/10.5281/zenodo.3470047" httpUri="https://zenodo.org/record/3470047/files/figure.png" pageId="6" pageNumber="53">Fig. 9</figureCitation>
: 20–21}; 1(1)
<specimenCount id="E5E6FD6CFFA204366900F3FA9171FAAC" box="[322,347,1269,1295]" pageId="6" pageNumber="53" type="male">♂</specimenCount>
,
</materialsCitation>
<materialsCitation id="43883CB8FFA204366925F3FA92C9FA97" ID-GBIF-Occurrence="2423610540" collectingDate="2012-07-18" collectionCode="CBK" collectorName="Kabak I. I." country="China" elevation="3060" latitude="43.2175" location="Narat Mt. Range Dzhergalan River" pageId="6" pageNumber="53" specimenCount="1" specimenCount-female="1" stateProvince="Xinjiang" typeStatus="paratype">
1(1)
<specimenCount id="E5E6FD6CFFA2043669DAF3FA9186FAAC" box="[408,428,1269,1295]" pageId="6" pageNumber="53" type="female">♀</specimenCount>
, “
<collectingCountry id="8BF77675FFA204366981F3FA9221FAAC" box="[451,523,1269,1295]" name="China" pageId="6" pageNumber="53">China</collectingCountry>
,
<collectingRegion id="3124F807FFA204366A54F3FA9251FAAC" box="[534,635,1269,1295]" country="China" name="Xinjiang Uygur" pageId="6" pageNumber="53">Xinjiang</collectingRegion>
,
<location id="F63F603EFFA204366AC4F3FA9425FAAC" LSID="urn:lsid:plazi:treatment:7B4987F3FFA2043868D5F4BE9431FE28:F63F603EFFA204366AC4F3FA9425FAAC" box="[646,1039,1269,1295]" country="China" latitude="43.2175" name="Narat Mt. Range Dzhergalan River" pageId="6" pageNumber="53" stateProvince="Xinjiang">Narat Mt. Range Dzhergalan River</location>
, H=
<quantity id="34189B00FFA204366C7FF3FA94BEFAAC" box="[1085,1172,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="6" pageNumber="53" unit="m" value="3060.0">
<elevation id="78CDD1D6FFA204366C7FF3FA94BEFAAC" box="[1085,1172,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="3.06" pageId="6" pageNumber="53" unit="m" value="3060.0">3060 m</elevation>
</quantity>
,
<date id="875E1025FFA204366CE2F3FA950AFAAC" box="[1184,1312,1269,1295]" pageId="6" pageNumber="53" value="2012-07-18">
<collectingDate id="971AE9CDFFA204366CE2F3FA950AFAAC" box="[1184,1312,1269,1295]" pageId="6" pageNumber="53" value="2012-07-18">18.07.2012</collectingDate>
</date>
,
<collectorName id="5E155333FFA204366D6EF3FA95B6FAAC" box="[1324,1436,1269,1295]" pageId="6" pageNumber="53">Kabak I.I.</collectorName>
leg.” [
<geoCoordinate id="96D45022FFA204366899F216915BFA90" box="[219,369,1305,1331]" degrees="43" direction="north" minutes="13" orientation="latitude" pageId="6" pageNumber="53" precision="15" seconds="03" value="43.2175">43º13´03´´ N</geoCoordinate>
/
<geoCoordinate id="96D45022FFA2043669C5F2179228FA90" box="[391,514,1304,1331]" pageId="6" pageNumber="53">83º26´29´´</geoCoordinate>
] (
<collectionCode id="95F1AE20FFA204366A5AF216927EFA90" box="[536,596,1305,1331]" pageId="6" pageNumber="53">CBK</collectionCode>
) {
<figureCitation id="6BDB2A60FFA204366A2EF216929FFA90" box="[620,693,1305,1331]" captionStart="FIGURE 9" captionStartId="15.[151,250,999,1024]" captionTargetBox="[151,1436,174,979]" captionTargetId="figure@15.[151,1436,174,979]" captionTargetPageId="15" captionText="FIGURE 9. Collecting sites of Trechus species of the uygur group. Yellow rhomb—T. sinus Belousov &amp; Kabak, 2001; red rhomb—T. arshanicus Belousov &amp; Kabak, 2001; yellow square—T. uygur ssp. kuytunicus Belousov &amp; Kabak, 2001; red square—T. uygur ssp. uygur Deuve, 1993; yellow triangle—T. torgaut sp. n.; red triangle—T. tsanmensis sp. n. Locality numbers—as in the “Material” section; those for collecting sites of T. arshanicus and T. sinus—see Belousov &amp; Kabak (2001)." figureDoi="http://doi.org/10.5281/zenodo.3470047" httpUri="https://zenodo.org/record/3470047/files/figure.png" pageId="6" pageNumber="53">Fig. 9</figureCitation>
: 16
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}.
</paragraph>
<paragraph id="F35F36E5FFA204366885F2339451FAD8" blockId="6.[151,1437,1052,2015]" pageId="6" pageNumber="53">
<emphasis id="C194EAF7FFA204366885F2339212FAF4" bold="true" box="[199,568,1340,1367]" pageId="6" pageNumber="53">Additional material examined</emphasis>
:
<materialsCitation id="43883CB8FFA204366A0FF2329447FAD8" ID-GBIF-Occurrence="2423610539" collectingDate="2012-07-18" collectionCode="CBK" collectorName="Kabak I. I." country="China" elevation="3290" latitude="43.20778" location="Narat Mt. Range" municipality="Narat Mt. Range" pageId="6" pageNumber="53" specimenCount="1" specimenCount-female="1" stateProvince="Xinjiang">
1(1)
<specimenCount id="E5E6FD6CFFA204366A3FF23292BAFAF4" box="[637,656,1341,1367]" pageId="6" pageNumber="53" type="female">♀</specimenCount>
(?), “
<collectingCountry id="8BF77675FFA204366A98F2329308FAF4" box="[730,802,1341,1367]" name="China" pageId="6" pageNumber="53">China</collectingCountry>
,
<collectingRegion id="3124F807FFA204366B71F23293B2FAF4" box="[819,920,1341,1367]" country="China" name="Xinjiang Uygur" pageId="6" pageNumber="53">Xinjiang</collectingRegion>
,
<collectingMunicipality id="133BAC9FFFA204366BEAF2329445FAF4" box="[936,1135,1341,1367]" pageId="6" pageNumber="53">Narat Mt. Range</collectingMunicipality>
Tshon-Dzhergalan Basin, H=
<quantity id="34189B00FFA2043668F8F26E9138FAD8" box="[186,274,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="6" pageNumber="53" unit="m" value="3290.0">
<elevation id="78CDD1D6FFA2043668F8F26E9138FAD8" box="[186,274,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="6" pageNumber="53" unit="m" value="3290.0">3290 m</elevation>
</quantity>
,
<date id="875E1025FFA20436695CF26E91B4FAD8" box="[286,414,1377,1403]" pageId="6" pageNumber="53" value="2012-07-18">
<collectingDate id="971AE9CDFFA20436695CF26E91B4FAD8" box="[286,414,1377,1403]" pageId="6" pageNumber="53" value="2012-07-18">18.07.2012</collectingDate>
</date>
,
<collectorName id="5E155333FFA2043669E8F26E9230FAD8" box="[426,538,1377,1403]" pageId="6" pageNumber="53">Kabak I.I.</collectorName>
leg.” [
<geoCoordinate id="96D45022FFA204366A27F26E92D1FAD8" box="[613,763,1377,1403]" degrees="43" direction="north" minutes="12" orientation="latitude" pageId="6" pageNumber="53" precision="15" seconds="28" value="43.20778">43º12´28´´ N</geoCoordinate>
/
<geoCoordinate id="96D45022FFA204366B53F26F93A7FAD8" box="[785,909,1376,1403]" pageId="6" pageNumber="53">83º26´39´´</geoCoordinate>
] (
<collectionCode id="95F1AE20FFA204366BE0F26E93F4FAD8" box="[930,990,1377,1403]" pageId="6" pageNumber="53">CBK</collectionCode>
) {
<figureCitation id="6BDB2A60FFA204366BB4F26E9415FAD8" box="[1014,1087,1377,1403]" captionStart="FIGURE 9" captionStartId="15.[151,250,999,1024]" captionTargetBox="[151,1436,174,979]" captionTargetId="figure@15.[151,1436,174,979]" captionTargetPageId="15" captionText="FIGURE 9. Collecting sites of Trechus species of the uygur group. Yellow rhomb—T. sinus Belousov &amp; Kabak, 2001; red rhomb—T. arshanicus Belousov &amp; Kabak, 2001; yellow square—T. uygur ssp. kuytunicus Belousov &amp; Kabak, 2001; red square—T. uygur ssp. uygur Deuve, 1993; yellow triangle—T. torgaut sp. n.; red triangle—T. tsanmensis sp. n. Locality numbers—as in the “Material” section; those for collecting sites of T. arshanicus and T. sinus—see Belousov &amp; Kabak (2001)." figureDoi="http://doi.org/10.5281/zenodo.3470047" httpUri="https://zenodo.org/record/3470047/files/figure.png" pageId="6" pageNumber="53">Fig. 9</figureCitation>
: 17
</materialsCitation>
}.
</paragraph>
</subSubSection>
<subSubSection id="BBFA656EFFA204376885F28B90C3F818" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53" type="description">
<paragraph id="F35F36E5FFA204366885F28B90E0F98C" blockId="6.[151,1437,1052,2015]" pageId="6" pageNumber="53">
<emphasis id="C194EAF7FFA204366885F28B9173FA3C" bold="true" box="[199,345,1412,1439]" pageId="6" pageNumber="53">Description.</emphasis>
Medium-sized species for the
<taxonomicName id="34E04D66FFA204366AFBF28992D0FA3C" authorityName="Deuve" authorityYear="1993" box="[697,762,1414,1439]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA204366AFBF28992D0FA3C" box="[697,762,1414,1439]" italics="true" pageId="6" pageNumber="53">uygur</emphasis>
</taxonomicName>
species group, body length
<quantity id="34189B00FFA204366C7FF28B94CFFA3C" box="[1085,1253,1412,1439]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.25" metricValueMin="4.65" pageId="6" pageNumber="53" unit="mm" value="4.95" valueMax="5.25" valueMin="4.65">4.65–5.25 mm</quantity>
. Body elongate, constricted at pronotal base, markedly depressed (
<figureCitation id="6BDB2A60FFA204366A8AF2A69327FA60" box="[712,781,1449,1475]" captionStart="FIGURES 1–2" captionStartId="5.[151,264,1686,1711]" captionTargetBox="[167,1427,390,1654]" captionTargetId="figure@5.[151,1435,373,1666]" captionTargetPageId="5" captionText="FIGURES 1–2. Trechus Clairville, 1806, habitus, males, paratypes. 1, T. tsanmensis sp. n.; 2, T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470039" httpUri="https://zenodo.org/record/3470039/files/figure.png" pageId="6" pageNumber="53">Fig. 2</figureCitation>
). Antennae and legs long, especially tarsi. Upper-side reddish dark brown, usually with slightly darker head and disc of elytra. Anterior part of head, disc of pronotum, base, suture and margins of elytra paler, reddish. Legs uniformly brown reddish. Antennae vaguely obscured in apical half.
</paragraph>
<paragraph id="F35F36E5FFA204366885F13690F7F940" blockId="6.[151,1437,1052,2015]" pageId="6" pageNumber="53">Head large, eyes small and subconvex, tempora long and convex, their pubescence sparse and extremely short, often barely distinguishable. Frontal furrows evenly impressed, slightly arcuate, nearly subparallel-sided and rather distant from each other in middle. Supraorbital setae located in lines distinctly convergent posteriad, posterior seta far behind posterior margin of eye. Pores of both anterior and posterior setae foveolate, though posterior one shallower.</paragraph>
<paragraph id="F35F36E5FFA204376885F1E293DCFF74" blockId="6.[151,1437,1052,2015]" lastBlockId="7.[151,1437,153,2015]" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53">Pronotum subcordate, convex on disc, markedly contracted at base, with maximum width in apical third. Its sides broadly arcuate in anterior part, nearly straight in posterior part, deeply sinuate before hind angles, the latter large and acute, produced mostly outwards. Posterior margin slightly bisinuate or straight; anterior margin nearly rectilinear, only slightly longer than posterior margin. Anterior angles broadly rounded and more or less produced anteriad. Lateral margins bordered and widely reflexed, especially so in posterior half. Prebasal transverse impression subparallel to posterior margin, rectangularly curved in basal foveae, rather deep, though not sharply engraved. Basal foveae average in size and depth. Apical transverse impression distinct throughout. Apical portion finely, basal portion roughly rugose. Anterior marginal seta placed in about anterior third of pronotum, posterior seta close to hind angle. Median line evenly impressed, more deeply near basal margin.</paragraph>
<paragraph id="F35F36E5FFA304376885F7EE90F2FDB8" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">Elytra narrow, oblong-oval, widest in apical third, with nearly rectilinear sides, much narrower at level of humeri than at level of subapical sinuation. Humeri broadly rounded, prehumeral margin arcuate. Each elytron sinuate before and rounded at apex. Striation of elytra well-impressed: all striae distinct and continuous, though outer ones much shallower and occasionally shortened anteriad. Inner intervals slightly convex, others flat. Apical recurrent striole short and nearly straight, sinuate and joining stria 5 anteriorly. Stria 3 joining stria 4 on apical slope, stria 2 surpassing level of preapical pore, stria 6 joining stria 7 distinctly before anterior termination of recurrent striole. Parascutellar striole distinct. Apical triangle (three pores on apical slope of elytra) slightly oblong, angulo-apical pore located approximately in middle between exterior pore and apex of elytra. All striae finely but distinctly punctured.</paragraph>
<paragraph id="F35F36E5FFA304376885F52A90CCFD08" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">Microsculpture shallowly engraved, faint medially on disc of both head and pronotum, consisting of isodiametric meshes on parietal area, irregular and slightly transverse meshes on disc of pronotum, and distinctly transverse meshes on elytra. Body finely micropunctured over entire surface, especially densely near anterior margin of pronotum.</paragraph>
<paragraph id="F35F36E5FFA304376885F5BA95ACFD50" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">Two basal segments dilated and clearly transverse in male anterior tarsi, both provided with adhesive appendages beneath. Front tibiae flattened on exterior surface, glabrous on anterior surface, except for one row of setae.</paragraph>
<paragraph id="F35F36E5FFA304376885F5F294C6FC68" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">
Aedeagus (
<figureCitation id="6BDB2A60FFA304376904F5F291A0FCB4" box="[326,394,765,791]" captionStart="FIGURES 3–4" captionStartId="12.[151,264,1957,1982]" captionTargetBox="[210,1372,183,1923]" captionTargetId="figure@12.[200,1387,166,1937]" captionTargetPageId="12" captionText="FIGURES 3–4. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view, c—paramere. 3—T. tsanmensis sp. n.; 4—T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470041" httpUri="https://zenodo.org/record/3470041/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
) of medium size, subcylindrical, slightly S-shaped, with oblique apical disc; sagittal aileron medium to small, nearly effaced in some specimens. Lamella rather short, medium-sized, well-defined, of triangular shape, rounded apically, directed to right side in dorsal view. Parameres slender, nearly straight in apical half; left one clearly longer, with distinctly protruding ventral apophysis; each paramere bearing 5–6 apical setae. Endophallus armature rather small, consisting of two pieces located in apical half of median lobe; larger piece semicylindrical in shape, located near ventral surface of aedeagal tube; second piece located near dorsolateral wall.
</paragraph>
<paragraph id="F35F36E5FFA304376885F4DB937CFB94" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">
<emphasis id="C194EAF7FFA304376885F4DB9193FC4C" bold="true" box="[199,441,980,1007]" pageId="7" pageNumber="54">Sexual dimorphism.</emphasis>
Males differ in smaller head (EW/HW on average, 1.76 vs.
<quantity id="34189B00FFA304376C35F4DA94EDFC4C" box="[1143,1223,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" pageId="7" pageNumber="54" unit="in" value="1.7">1.70 in</quantity>
females, p≤0.001; PW/HW 1.16 vs. 1.14 respectively, p≤0.01), longer elytra (EL/PL, on average, 3.01 vs.
<quantity id="34189B00FFA304376C3DF4F794E5FBB0" box="[1151,1231,1016,1043]" metricMagnitude="-2" metricUnit="m" metricValue="7.493" pageId="7" pageNumber="54" unit="in" value="2.95">2.95 in</quantity>
females, p≤0.05), and longer antennae (AL/EL 1.01 vs.
<quantity id="34189B00FFA304376A7FF31292A1FB94" box="[573,651,1053,1079]" metricMagnitude="-2" metricUnit="m" metricValue="2.5146" pageId="7" pageNumber="54" unit="in" value="0.99">0.99 in</quantity>
females, p≤0.01).
</paragraph>
<paragraph id="F35F36E5FFA304376885F34F9130FAB3" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">
<emphasis id="C194EAF7FFA304376885F34F9112FBF8" bold="true" box="[199,312,1088,1115]" pageId="7" pageNumber="54">Variation</emphasis>
. Externally, the population from the Tsagan-Usun River is larger (mean body length 5.0 mm vs.
<quantity id="34189B00FFA304376D3BF34E90E9FBDC" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="54" unit="mm" value="4.9">4.9 mm</quantity>
in specimens from the
<typeStatus id="2C5B8847FFA30437698EF36A91D6FBDC" box="[460,508,1125,1151]" pageId="7" pageNumber="54">type</typeStatus>
locality, p≤0.05), with pronotum wider (PW/PL, on average, 1.41 vs. 1.38 respec- tively, p≤0.01), anterior lateral seta of pronotum is located more posteriad (28% vs. 25% respectively, p≤0.001), all discal pores of elytra are located closer to the elytral base (discal formula: 26, 60, 89 vs. 27, 64,
<quantity id="34189B00FFA304376CA7F3A29535FB64" box="[1253,1311,1197,1223]" metricMagnitude="0" metricUnit="m" metricValue="2.3114" pageId="7" pageNumber="54" unit="in" value="91.0">91 in</quantity>
specimens from the
<typeStatus id="2C5B8847FFA304376942F3DE911AFB48" box="[256,304,1233,1259]" pageId="7" pageNumber="54">type</typeStatus>
locality), umbilicate pores 1 and 2 are located more posteriorly (10% and 16% vs. 9% and 15% respectively).
</paragraph>
<paragraph id="F35F36E5FFA304376885F217954BFAF4" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">
<emphasis id="C194EAF7FFA304376885F217919BFA90" bold="true" box="[199,433,1304,1331]" pageId="7" pageNumber="54">Comparative notes.</emphasis>
<taxonomicName id="34E04D66FFA3043769FAF215920FFA90" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[440,549,1306,1331]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA3043769FAF215920FFA90" box="[440,549,1306,1331]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376A6FF2169247FA90" bold="true" box="[557,621,1305,1331]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376A6FF2169247FA90" box="[557,621,1305,1331]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from all known members of the
<taxonomicName id="34E04D66FFA304376C6CF2159445FA90" authorityName="Deuve" authorityYear="1993" box="[1070,1135,1306,1331]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376C6CF2159445FA90" box="[1070,1135,1306,1331]" italics="true" pageId="7" pageNumber="54">uygur</emphasis>
</taxonomicName>
species group, in having a medium-sized aedeagus (very large in other species) with a rather short apical portion clearly curved upward.
</paragraph>
<paragraph id="F35F36E5FFA304376885F26D90C3F818" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="54">
<taxonomicName id="34E04D66FFA304376885F26D911FFAD8" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[199,309,1378,1403]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA304376885F26D911FFAD8" box="[199,309,1378,1403]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA30437697CF26E9154FAD8" bold="true" box="[318,382,1377,1403]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA30437697CF26E9154FAD8" box="[318,382,1377,1403]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
seems to be more similar to
<taxonomicName id="34E04D66FFA304376A8BF26D934BFAD8" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[713,865,1378,1403]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFA304376A8BF26D934BFAD8" box="[713,865,1378,1403]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376B2BF26E9380FAD8" bold="true" box="[873,938,1377,1403]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376B2BF26E9380FAD8" box="[873,938,1377,1403]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described above, but differs, apart from the above-listed unique characters, in proportionally larger apical lamella and quite different endophallus armature with dorsal piece much smaller, located less proximally and consisting of two poorly sclerotized symmetrical lobes (
<figureCitation id="6BDB2A60FFA3043768E2F2C290C8FA44" box="[160,226,1485,1511]" captionStart="FIGURES 3–4" captionStartId="12.[151,264,1957,1982]" captionTargetBox="[210,1372,183,1923]" captionTargetId="figure@12.[200,1387,166,1937]" captionTargetPageId="12" captionText="FIGURES 3–4. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view, c—paramere. 3—T. tsanmensis sp. n.; 4—T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470041" httpUri="https://zenodo.org/record/3470041/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
vs.
<figureCitation id="6BDB2A60FFA30437694DF2C29178FA44" box="[271,338,1485,1511]" captionStart="FIGURES 3–4" captionStartId="12.[151,264,1957,1982]" captionTargetBox="[210,1372,183,1923]" captionTargetId="figure@12.[200,1387,166,1937]" captionTargetPageId="12" captionText="FIGURES 3–4. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view, c—paramere. 3—T. tsanmensis sp. n.; 4—T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470041" httpUri="https://zenodo.org/record/3470041/files/figure.png" pageId="7" pageNumber="54">Fig. 3</figureCitation>
). Externally
<taxonomicName id="34E04D66FFA3043769A1F2C19265FA44" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[483,591,1486,1511]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA3043769A1F2C19265FA44" box="[483,591,1486,1511]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376A15F2C292BCFA44" bold="true" box="[599,662,1485,1511]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376A15F2C292BCFA44" box="[599,662,1485,1511]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="34E04D66FFA304376B87F2C19471FA44" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[965,1115,1486,1511]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFA304376B87F2C19471FA44" box="[965,1115,1486,1511]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376C20F2C2948BFA44" bold="true" box="[1122,1185,1485,1511]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376C20F2C2948BFA44" box="[1122,1185,1485,1511]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by its smaller size (on average,
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vs.
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in
<taxonomicName id="34E04D66FFA304376A57F2FD9281F9A8" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[533,683,1522,1547]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFA304376A57F2FD9281F9A8" box="[533,683,1522,1547]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376AF0F2FE92DBF9A8" bold="true" box="[690,753,1521,1547]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376AF0F2FE92DBF9A8" box="[690,753,1521,1547]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), much narrower habitus with proportionally wider head and pronotum (mean for EW/HW ratio is 1.73 vs.
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<taxonomicName id="34E04D66FFA304376AB5F11993A4F98C" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[759,910,1558,1583]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFA304376AB5F11993A4F98C" box="[759,910,1558,1583]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376BD4F11A93FCF98C" bold="true" box="[918,982,1557,1583]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376BD4F11A93FCF98C" box="[918,982,1557,1583]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; PW/PL: 1.40 vs.
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<taxonomicName id="34E04D66FFA304376CBFF11995BFF98C" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[1277,1429,1558,1583]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis">
<emphasis id="C194EAF7FFA304376CBFF11995BFF98C" box="[1277,1429,1558,1583]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
; EW/PW: 1.50 vs.
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<taxonomicName id="34E04D66FFA3043769FAF1359265F9F0" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[440,591,1594,1619]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis">
<emphasis id="C194EAF7FFA3043769FAF1359265F9F0" box="[440,591,1594,1619]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
) and the second discal pores of elytra located mostly at level of umbilicate pores 5 vs. clearly before this level in
<taxonomicName id="34E04D66FFA304376A7CF15192FEF9D4" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[574,724,1630,1655]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFA304376A7CF15192FEF9D4" box="[574,724,1630,1655]" italics="true" pageId="7" pageNumber="54">T. tsanmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAA7578CFFA304376A98F1529332F9D4" box="[730,792,1629,1655]" pageId="7" pageNumber="54" rank="species">
<emphasis id="C194EAF7FFA304376A98F1529338F9D4" bold="true" box="[730,786,1629,1655]" pageId="7" pageNumber="54">sp. n</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="34E04D66FFA304376B5DF15193A1F9D4" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[799,907,1630,1655]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA304376B5DF15193A1F9D4" box="[799,907,1630,1655]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376BD0F15293FBF9D4" bold="true" box="[914,977,1629,1655]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376BD0F15293FBF9D4" box="[914,977,1629,1655]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is also similar to
<emphasis id="C194EAF7FFA304376CD6F15194DEF9D4" box="[1172,1268,1630,1655]" italics="true" pageId="7" pageNumber="54">
<taxonomicName id="34E04D66FFA304376CD6F15194DAF9D4" authorityName="Deuve" authorityYear="1993" box="[1172,1264,1630,1655]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">T. uygur</taxonomicName>
,
</emphasis>
though readily distinguished, first of all, by its shorter apical lamella, which is triangular-shaped in dorsal view vs. very long and parallel-sided, pointed apically in
<taxonomicName id="34E04D66FFA304376A5AF1A9925CF91C" authorityName="Deuve" authorityYear="1993" box="[536,630,1702,1727]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376A5AF1A9925CF91C" box="[536,630,1702,1727]" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
and smaller endophallus armature (
<figureCitation id="6BDB2A60FFA304376C49F1AA9464F91C" box="[1035,1102,1701,1727]" captionStart="FIGURES 3–4" captionStartId="12.[151,264,1957,1982]" captionTargetBox="[210,1372,183,1923]" captionTargetId="figure@12.[200,1387,166,1937]" captionTargetPageId="12" captionText="FIGURES 3–4. Male genitalia of Trechus spp. n., paratypes: a—lateral view, b—dorsal view, c—paramere. 3—T. tsanmensis sp. n.; 4—T. torgaut sp. n." figureDoi="http://doi.org/10.5281/zenodo.3470041" httpUri="https://zenodo.org/record/3470041/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
vs.
<figureCitation id="6BDB2A60FFA304376C3CF1AA94CEF91C" box="[1150,1252,1700,1727]" captionStart="FIGURES 5–6" captionStartId="13.[151,264,1972,1997]" captionTargetBox="[177,1415,179,1937]" captionTargetId="figure@13.[159,1429,166,1952]" captionTargetPageId="13" captionText="FIGURES 5–6. Male genitalia of Trechus uygur (from Belousov &amp; Kabak 2001): a—lateral view, b—dorsal view. 5—T. uygur ssp. uygur Deuve, 1993 from the Ulan-Gelik Pass; 6—T. uygur ssp. kuytunicus Belousov &amp; Kabak, 2001, paratype." figureDoi="http://doi.org/10.5281/zenodo.3470043" httpUri="https://zenodo.org/record/3470043/files/figure.png" pageId="7" pageNumber="54">Figs 5–6</figureCitation>
). Externally, the new species is distinguished from allied taxa by the shape of the pronotum with anterior angles, on average, more distinctly produced anteriad. Morphometrically, the new species differs from
<taxonomicName id="34E04D66FFA304376BBAF1E1947EF8A4" authorityName="Deuve" authorityYear="1993" box="[1016,1108,1774,1799]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376BBAF1E1947EF8A4" box="[1016,1108,1774,1799]" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
in the longer anterior margin of the pronotum (PA/PB ratio is 1.04 vs.
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<taxonomicName id="34E04D66FFA304376AFDF01D9336F888" authorityName="Deuve" authorityYear="1993" box="[703,796,1810,1835]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376AFDF01D9336F888" box="[703,796,1810,1835]" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
, p≤0.001). In addition,
<taxonomicName id="34E04D66FFA304376C68F01D94BDF888" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[1066,1175,1810,1835]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFA304376C68F01D94BDF888" box="[1066,1175,1810,1835]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFA304376CE2F01E94CAF888" bold="true" box="[1184,1248,1809,1835]" pageId="7" pageNumber="54">
<taxonomicNameLabel id="DAA7578CFFA304376CE2F01E94CAF888" box="[1184,1248,1809,1835]" pageId="7" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has wider elytra (means for EL/EW:
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<taxonomicName id="34E04D66FFA30437699BF0399262F8EC" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[473,584,1846,1871]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="torgaut">
<emphasis id="C194EAF7FFA30437699BF0399262F8EC" box="[473,584,1846,1871]" italics="true" pageId="7" pageNumber="54">T. torgaut</emphasis>
</taxonomicName>
vs.
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<taxonomicName id="34E04D66FFA304376A91F0399318F8EC" authorityName="Deuve" authorityYear="1993" box="[723,818,1846,1871]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376A91F0399318F8EC" box="[723,818,1846,1871]" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
, p≤0.001), more transverse pronotum (average values for PW/PL: 1.40 vs. 1.34, p≤0.001), longer antennae (means for EL/AL: 1.00 vs.
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<taxonomicName id="34E04D66FFA304376CD5F05594DFF8D0" authorityName="Deuve" authorityYear="1993" box="[1175,1269,1882,1907]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376CD5F05594DFF8D0" box="[1175,1269,1882,1907]" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
, p≤0.001) and somewhat smaller eyes; all discal setae are slightly shifted posteriad (discal formula: 26, 62, 90 vs. 24, 57,
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<taxonomicName id="34E04D66FFA304376DCAF07190F2F818" authorityName="Deuve" authorityYear="1993" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="uygur">
<emphasis id="C194EAF7FFA304376DCAF07190F2F818" italics="true" pageId="7" pageNumber="54">T. uygur</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="BBFA656EFFA304386885F0CB91EDFEC4" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54" type="distribution">
<paragraph id="F35F36E5FFA304386885F0CB92C9FF58" blockId="7.[151,1437,153,2015]" lastBlockId="8.[151,1437,153,395]" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
<emphasis id="C194EAF7FFA304376885F0CB914BF87C" bold="true" box="[199,353,1988,2015]" pageId="7" pageNumber="54">Distribution.</emphasis>
The species inhabits mountains at sources of the Kunges River,
<collectingRegion id="3124F807FFA304376C05F0CA9486F87C" box="[1095,1196,1989,2015]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="54">Xinjiang</collectingRegion>
,
<collectingCountry id="8BF77675FFA304376CFAF0CA94D5F87C" box="[1208,1279,1989,2015]" name="China" pageId="7" pageNumber="54">China</collectingCountry>
. Additionally, a small series of this species was collected on the northern slopes of the Narat Mountain Range, in the basin of the Tshon-Dzhergalan River, approximately
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west of the
<typeStatus id="2C5B8847FFAC04386B7DF7B29345FF74" box="[831,879,189,215]" pageId="8" pageNumber="55">type</typeStatus>
locality of
<taxonomicName id="34E04D66FFAC04386BAFF7B194A9FF74" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[1005,1155,190,215]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="tsanmensis" status="sp. nov.">
<emphasis id="C194EAF7FFAC04386BAFF7B194A9FF74" box="[1005,1155,190,215]" italics="true" pageId="8" pageNumber="55">T. tsanmensis</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFAC04386CCBF7B294EDFF74" bold="true" box="[1161,1223,189,215]" pageId="8" pageNumber="55">
<taxonomicNameLabel id="DAA7578CFFAC04386CCBF7B294EDFF74" box="[1161,1223,189,215]" pageId="8" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and more than
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west of other known findings of
<taxonomicName id="34E04D66FFAC04386A6DF7ED92B6FF58" authority="Belousov &amp; Kabak, 2019" authorityName="Belousov &amp; Kabak" authorityYear="2019" box="[559,668,226,251]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="torgaut" status="sp. nov.">
<emphasis id="C194EAF7FFAC04386A6DF7ED92B6FF58" box="[559,668,226,251]" italics="true" pageId="8" pageNumber="55">T. torgaut</emphasis>
</taxonomicName>
<emphasis id="C194EAF7FFAC04386AE1F7EE92C9FF58" bold="true" box="[675,739,225,251]" pageId="8" pageNumber="55">
<taxonomicNameLabel id="DAA7578CFFAC04386AE1F7EE92C9FF58" box="[675,739,225,251]" pageId="8" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="F35F36E5FFAC04386885F60B91EDFEC4" blockId="8.[151,1437,153,395]" pageId="8" pageNumber="55">
<emphasis id="C194EAF7FFAC04386885F60B9161FEBC" bold="true" box="[199,331,260,287]" pageId="8" pageNumber="55">Bionomics.</emphasis>
The species occurs in a comparatively wide belt of altitudes between 2500 and
<quantity id="34189B00FFAC04386CACF60A9562FEBC" box="[1262,1352,261,287]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="55" unit="m" value="3400.0">3400 m</quantity>
. It was found in the forest zone as well as in the alpine zone and seems to be associated with humid biotopes, especially strictly at lower elevations.
</paragraph>
</subSubSection>
<subSubSection id="BBFA656EFFAC04386885F67F9431FE28" box="[199,1051,368,395]" pageId="8" pageNumber="55" type="etymology">
<paragraph id="F35F36E5FFAC04386885F67F9431FE28" blockId="8.[151,1437,153,395]" box="[199,1051,368,395]" pageId="8" pageNumber="55">
<emphasis id="C194EAF7FFAC04386885F67F918EFE28" bold="true" box="[199,420,368,395]" pageId="8" pageNumber="55">Derivatio nominis.</emphasis>
The species is named after a tribe of Mongolian people.
</paragraph>
</subSubSection>
</treatment>
</document>
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