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<document ID-DOI="10.11865/zs.2022101" ID-ISSN="2095-6827" ID-Zenodo-Dep="7176146" ID-ZooBank="1C62670D-1CEB-4A0C-8A12-D8824F6B61DA" approvalRequired="14" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="6" checkinTime="1665296205732" checkinUser="felipe" docAuthor="Xin, Yafei, Jiang, Tongyao, Yao, Zhiyuan &amp; Li, Shuqiang" docDate="2022" docId="743FB8540915BC50B1F1CC4BFBFBFD81" docLanguage="en" docName="ZoolSyst.47.1.1–65.pdf" docOrigin="Zoological Systematics 47 (1)" docSource="http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Palpalpaculla phyui Jiang &amp; Li 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="54" masterDocId="8806C02C0926BC65B179CF72FFCEFFBD" masterDocTitle="Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="52" updateTime="1665315257893" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xin, Yafei</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar Corresponding authors, E-mails: yaozy @ synu. edu. cn; lisq @ ioz. ac. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Tongyao</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yao, Zhiyuan</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>65</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11865/zs.2022101</mods:identifier>
<mods:identifier type="ISSN">2095-6827</mods:identifier>
<mods:identifier type="Zenodo-Dep">7176146</mods:identifier>
<mods:identifier type="ZooBank">1C62670D-1CEB-4A0C-8A12-D8824F6B61DA</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172614" ID-Zenodo-Dep="7172614" LSID="urn:lsid:plazi:treatment:743FB8540915BC50B1F1CC4BFBFBFD81" httpUri="http://treatment.plazi.org/id/743FB8540915BC50B1F1CC4BFBFBFD81" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
<subSubSection box="[136,746,825,850]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph blockId="51.[136,746,825,850]" box="[136,746,825,850]" pageId="51" pageNumber="52">
<heading box="[136,746,825,850]" centered="true" fontSize="10" level="2" pageId="51" pageNumber="52" reason="2">
<emphasis bold="true" box="[136,586,825,850]" pageId="51" pageNumber="52">
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[136,489,825,849]" class="Arachnida" family="Tetrablemmidae" genus="Palpalpaculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="phyui" status="sp. nov.">
<emphasis bold="true" box="[136,357,825,849]" italics="true" pageId="51" pageNumber="52">Palpalpaculla phyui</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel box="[501,586,826,850]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[616,738,826,850]" captionStart-0="Figure 49" captionStart-1="Figure 50" captionStart-2="Figure 51" captionStartId-0="51.[155,218,2034,2056]" captionStartId-1="52.[155,218,2025,2047]" captionStartId-2="53.[155,218,2038,2060]" captionTargetBox-0="[189,1399,980,1994]" captionTargetBox-1="[189,1398,473,1977]" captionTargetBox-2="[264,1296,656,1987]" captionTargetId-0="figure-323@51.[189,1399,980,1994]" captionTargetId-1="figure-133@52.[189,1398,473,1977]" captionTargetId-2="figure-251@53.[264,1323,652,1990]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="Figure 49. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo). A. Left palp, dorsal view; B. Right palp, dorsal view. Scale bars=0.05mm." captionText-1="Figure 50. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." captionText-2="Figure 51. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), left palp, dorsal view. Scale bar=0.05mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176258" figureDoi-1="http://doi.org/10.5281/zenodo.7176260" figureDoi-2="http://doi.org/10.5281/zenodo.7176262" httpUri-0="https://zenodo.org/record/7176258/files/figure.png" httpUri-1="https://zenodo.org/record/7176260/files/figure.png" httpUri-2="https://zenodo.org/record/7176262/files/figure.png" pageId="51" pageNumber="52">Figs 49–51</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="materials_examined">
<paragraph blockId="51.[136,1452,881,940]" pageId="51" pageNumber="52">
<materialsCitation county="Hukawng Valley" location="Male" municipality="Late Cretaceous" pageId="51" pageNumber="52" specimenCount="1" typeStatus="holotype">
<typeStatus box="[189,292,881,905]" pageId="51" pageNumber="52">Holotype</typeStatus>
.
<location LSID="urn:lsid:plazi:treatment:743FB8540915BC50B1F1CC4BFBFBFD81:F9495F990915BC56B04DCC03FEA5FC34" box="[308,363,881,905]" county="Hukawng Valley" municipality="Late Cretaceous" name="Male" pageId="51" pageNumber="52">Male</location>
(IZCAS-Ar42701Fo).
<collectingMunicipality box="[622,800,881,905]" pageId="51" pageNumber="52">Late Cretaceous</collectingMunicipality>
amber from
<collectingCounty box="[957,1142,881,905]" pageId="51" pageNumber="52">Hukawng Valley</collectingCounty>
. Syninclusions include two dipterans.
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176258" ID-Zenodo-Dep="7176258" httpUri="https://zenodo.org/record/7176258/files/figure.png" pageId="51" pageNumber="52" startId="51.[155,218,2034,2056]" targetBox="[189,1399,980,1994]" targetPageId="51">
<paragraph blockId="51.[155,1433,2034,2088]" pageId="51" pageNumber="52">
Figure 49.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[262,574,2034,2056]" class="Arachnida" family="Tetrablemmidae" genus="Palpalpaculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="phyui" status="sp. nov.">
<emphasis box="[262,459,2034,2055]" italics="true" pageId="51" pageNumber="52">Palpalpaculla phyui</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[585,662,2034,2055]" pageId="51" pageNumber="52">
<taxonomicNameLabel box="[585,662,2034,2055]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42701Fo). A. Left palp, dorsal view; B. Right palp, dorsal view. Scale bars =0.05 mm.
</paragraph>
</caption>
<paragraph blockId="52.[136,1455,236,399]" box="[189,1301,236,260]" pageId="52" pageNumber="53">Etymology. The species name is a popular Burmese boy’s name and means “white”; noun in apposition.</paragraph>
<paragraph blockId="52.[136,1455,236,399]" pageId="52" pageNumber="53">
Diagnosis. The new species resembles
<taxonomicName authorityName="Wunderlich" authorityYear="2017" box="[610,719,271,295]" class="Arachnida" family="Tetrablemmidae" genus="Palpalpaculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis box="[610,719,271,295]" italics="true" pageId="52" pageNumber="53">P. pulcher</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Wunderlich" box="[777,1026,271,295]" pageId="52" pageNumber="53" refId="ref14002" refString="Wunderlich, J. 2017. New and rare fossil spiders (Araneae) in Mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae. In: Wunderlich, J. (ed.), Beitrage zur Araneologie, Vol. 10. Hirschberg, Germany. pp. 72 - 279." type="book" year="2017">Wunderlich, 2017: 135</bibRefCitation>
, figs 89–91, photos 66–67) by the bulb shape but can be distinguished by the embolus shape (
<figureCitation box="[739,872,306,330]" captionStart="Figure 49" captionStartId="51.[155,218,2034,2056]" captionTargetBox="[189,1399,980,1994]" captionTargetId="figure-323@51.[189,1399,980,1994]" captionTargetPageId="51" captionText="Figure 49. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo). A. Left palp, dorsal view; B. Right palp, dorsal view. Scale bars=0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176258" httpUri="https://zenodo.org/record/7176258/files/figure.png" pageId="52" pageNumber="53">Figs 49A–B</figureCitation>
,
<figureCitation box="[888,917,306,330]" captionStart="Figure 51" captionStartId="53.[155,218,2038,2060]" captionTargetBox="[264,1296,656,1987]" captionTargetId="figure-251@53.[264,1323,652,1990]" captionTargetPageId="53" captionText="Figure 51. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), left palp, dorsal view. Scale bar=0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176262" httpUri="https://zenodo.org/record/7176262/files/figure.png" pageId="52" pageNumber="53">51</figureCitation>
). The embolus is tuber-shaped and straight in
<taxonomicName authorityName="Wunderlich" authorityYear="2017" class="Arachnida" family="Tetrablemmidae" genus="Palpalpaculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis italics="true" pageId="52" pageNumber="53">P. pulcher</emphasis>
</taxonomicName>
and the new species, but the embolus of the new species is more complex distally.
</paragraph>
<paragraph blockId="52.[136,1455,236,399]" lastBlockId="53.[136,1452,236,572]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
Description. Male. Total length 1.96; carapace 0.81 long, 0.52 wide; opisthosoma 1.15 long, 0.65 wide. Left palp: 0.46 (0.16 + 0.09 + 0.21), left leg I: - (- + - + - + 0.42 + 0.32), leg
<collectionCode box="[790,811,237,260]" pageId="53" pageNumber="54">II</collectionCode>
: - (- + - + - + 0.33 + 0.28), leg
<collectionCode box="[1143,1173,237,260]" pageId="53" pageNumber="54">III</collectionCode>
: - (- + - + - + 0.32+ 0.21), leg IV: - (- + - + - + 0.43 + 0.31); right palp: 0.43 (0.15 + 0.09 + 0.19), right leg I: - (0.72 + - + - + 0.41 + 0.33), leg
<collectionCode box="[1409,1430,272,295]" pageId="53" pageNumber="54">II</collectionCode>
: - (0.46 + - + - + 0.35 + 0.33), leg
<collectionCode box="[476,507,306,329]" pageId="53" pageNumber="54">III</collectionCode>
: - (0.41 + - + - + 0.33 + 0.30), leg IV: - (0.54 + - + - + 0.45 + 0.34). Eye sizes: ALE 0.06, AME 0.06, PLE 0.06, eyes interdistances not visible. Habitus as in
<figureCitation box="[842,974,340,364]" captionStart="Figure 50" captionStartId="52.[155,218,2025,2047]" captionTargetBox="[189,1398,473,1977]" captionTargetId="figure-133@52.[189,1398,473,1977]" captionTargetPageId="52" captionText="Figure 50. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." figureDoi="http://doi.org/10.5281/zenodo.7176260" httpUri="https://zenodo.org/record/7176260/files/figure.png" pageId="53" pageNumber="54">Figs 50A–B</figureCitation>
. Carapace (
<figureCitation box="[1095,1191,340,364]" captionStart="Figure 50" captionStartId="52.[155,218,2025,2047]" captionTargetBox="[189,1398,473,1977]" captionTargetId="figure-133@52.[189,1398,473,1977]" captionTargetPageId="52" captionText="Figure 50. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." figureDoi="http://doi.org/10.5281/zenodo.7176260" httpUri="https://zenodo.org/record/7176260/files/figure.png" pageId="53" pageNumber="54">Fig. 50A</figureCitation>
) sclerotized, reticulated, brown; cephalic part moderately raised; six eyes (
<figureCitation box="[683,783,375,399]" captionStart="Figure 50" captionStartId="52.[155,218,2025,2047]" captionTargetBox="[189,1398,473,1977]" captionTargetId="figure-133@52.[189,1398,473,1977]" captionTargetPageId="52" captionText="Figure 50. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." figureDoi="http://doi.org/10.5281/zenodo.7176260" httpUri="https://zenodo.org/record/7176260/files/figure.png" pageId="53" pageNumber="54">Fig. 50A</figureCitation>
) almost ovoid, AME = ALE = PLE, ALE and PLE adjacent; clypeus brown; clypeus vertical anteriorly; labium triangular, distally obtuse; sternum marginally rugose, posterior corner protruded. Opisthosoma (
<figureCitation box="[407,543,444,468]" captionStart="Figure 50" captionStartId="52.[155,218,2025,2047]" captionTargetBox="[189,1398,473,1977]" captionTargetId="figure-133@52.[189,1398,473,1977]" captionTargetPageId="52" captionText="Figure 50. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars=0.50mm." figureDoi="http://doi.org/10.5281/zenodo.7176260" httpUri="https://zenodo.org/record/7176260/files/figure.png" pageId="53" pageNumber="54">Figs 50A–B</figureCitation>
): oval with sclerotized lateral plates, smooth, modified by tiny pits, covered with thin setae; ventral scutum rugose.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176260" ID-Zenodo-Dep="7176260" httpUri="https://zenodo.org/record/7176260/files/figure.png" pageId="52" pageNumber="53" startId="52.[155,218,2025,2047]" targetBox="[189,1398,473,1977]" targetPageId="52">
<paragraph blockId="52.[155,1433,2025,2079]" pageId="52" pageNumber="53">
Figure 50.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[262,571,2025,2047]" class="Arachnida" family="Tetrablemmidae" genus="Palpalpaculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="phyui" status="sp. nov.">
<emphasis box="[262,458,2025,2046]" italics="true" pageId="52" pageNumber="53">Palpalpaculla phyui</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[583,660,2025,2046]" pageId="52" pageNumber="53">
<taxonomicNameLabel box="[583,660,2025,2046]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42701Fo), habitus. A. Dorsal view; B. Ventral view. Scale bars = 0.50 mm.
</paragraph>
</caption>
<paragraph blockId="53.[136,1452,236,572]" pageId="53" pageNumber="54">
Palp (
<figureCitation box="[252,387,514,538]" captionStart="Figure 49" captionStartId="51.[155,218,2034,2056]" captionTargetBox="[189,1399,980,1994]" captionTargetId="figure-323@51.[189,1399,980,1994]" captionTargetPageId="51" captionText="Figure 49. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo). A. Left palp, dorsal view; B. Right palp, dorsal view. Scale bars=0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176258" httpUri="https://zenodo.org/record/7176258/files/figure.png" pageId="53" pageNumber="54">Figs 49A–B</figureCitation>
,
<figureCitation box="[399,429,514,538]" captionStart="Figure 51" captionStartId="53.[155,218,2038,2060]" captionTargetBox="[264,1296,656,1987]" captionTargetId="figure-251@53.[264,1323,652,1990]" captionTargetPageId="53" captionText="Figure 51. Palpalpaculla phyui Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42701Fo), left palp, dorsal view. Scale bar=0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176262" httpUri="https://zenodo.org/record/7176262/files/figure.png" pageId="53" pageNumber="54">51</figureCitation>
). Femur and patella with apophysis; cymbium small; bulb long, strongly swollen distally, bent once; embolus originates from apex of oval palpal bulb; embolus sclerotized, thickened.
</paragraph>
</subSubSection>
</treatment>
</document>
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