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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.494.8748" ID-GBIF-Dataset="e8b2a5fb-bd85-48c3-b33e-121ddc15bec9" ID-PMC="PMC4400378" ID-Pensoft-Pub="1313-2970-494-69" ID-PubMed="25901115" ID-ZBK="F73F468E4DF7479E85C4717466635BF9" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-494-69" ModsDocOrigin="ZooKeys 494" ModsDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" checkinTime="1451244473555" checkinUser="pensoft" docAuthor="Sohn, Jae-Cheon, Davis, Donald R. &amp; Lopez-Vaamonde, Carlos" docDate="2015" docId="5CED1F44916051D9BE42C839F0F893C1" docLanguage="en" docName="ZooKeys 494: 69-106" docOrigin="ZooKeys 494" docSource="http://dx.doi.org/10.3897/zookeys.494.8748" docTitle="Philonome penerivifera Sohn &amp; Davis, sp. n." docType="treatment" docUuid="48A07D44-E6AC-4D0B-AD2F-17863CB24E88" docUuidSource="ZooBank" docVersion="4" lastPageNumber="88" masterDocId="3D1D9C71FF96FFF09C07FFAC9A60FF9D" masterDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" masterLastPageNumber="106" masterPageNumber="69" pageNumber="86" updateTime="1668160171790" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sohn, Jae-Cheon</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davis, Donald R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lopez-Vaamonde, Carlos</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>494</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>69</mods:start>
<mods:end>106</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.494.8748</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.494.8748</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-494-69</mods:identifier>
<mods:identifier type="ZBK">F73F468E4DF7479E85C4717466635BF9</mods:identifier>
<mods:identifier type="ZooBank">F73F468E4DF7479E85C4717466635BF9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152059286" LSID="urn:lsid:zoobank.org:act:48A07D44-E6AC-4D0B-AD2F-17863CB24E88" httpUri="http://treatment.plazi.org/id/5CED1F44916051D9BE42C839F0F893C1" lastPageId="19" lastPageNumber="88" pageId="17" pageNumber="86">
<subSubSection pageId="17" pageNumber="86" type="multiple">
<paragraph pageId="17" pageNumber="86">Taxon classification Animalia Lepidoptera Lyonetiidae</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="86" type="nomenclature">
<paragraph pageId="17" pageNumber="86">
<taxonomicName LSID="http://zoobank.org/48A07D44-E6AC-4D0B-AD2F-17863CB24E88" authority="Sohn &amp; Davis" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="17" pageNumber="86" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera Sohn &amp; Davis</taxonomicName>
<taxonomicNameLabel pageId="17" pageNumber="86">sp. n.</taxonomicNameLabel>
Figs 10, 49-52, 61-62
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="87" type="diagnosis">
<paragraph pageId="18" pageNumber="87">
<pageBreakToken pageId="18" pageNumber="87" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="18" pageNumber="87">
This species is indistinguishable from
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
in external appearance. Both species can be clearly distinguished from each other by the male genitalia (Table 2), including distal margin of cucullus shallowly concave in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera</taxonomicName>
but deeply emarginated in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
; tegumen triangular in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera</taxonomicName>
but subrectangular in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
; and lateral area of vinculum less protruding in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera</taxonomicName>
than in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="87" type="adult">
<paragraph pageId="18" pageNumber="87">Adult</paragraph>
<paragraph pageId="18" pageNumber="87">(Fig. 10). Head: Vertex brownish white or pale orange on posterior 2/3, pale yellowish white on anterior 1/3; scales on interspace between antennal scapes yellowish white; frons lustrous yellowish white; scales on occiput pale orange, with dark brown tips dorsally, pale yellowish white laterally. Antenna 3/5 as long as forewing; scape pale grayish orange dorsally, lustrous yellowish white ventrally; flagellomeres pale orange dorsally, yellowish white ventrally. Labial palpus 1/2 as long as maxillary palpus, dark grayish brown laterally, lustrous yellowish white mesally. Maxillary palpus yellowish white.</paragraph>
<paragraph pageId="18" pageNumber="87">
Thorax: Scales of patagium pale orange, with dark brown tips; tegula reddish brown basally, paler to apex, pale orange apically; mesonotum pale orange, transversely intermixed with dark brown scales at middle. Fore- and midlegs with coxa lustrous yellowish white; femur, tibia, and tarsomeres dark brown dorsally, lustrous yellowish white laterally and ventrally. Hindleg pale brownish gray dorsally, lustrous yellowish white laterally and ventrally. Forewing length 3.2-3.6 mm (n = 2), coloration and patterns similar to
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
. Hindwing dark brownish gray; fringe brownish gray on costal and apical area, yellowish gray along posterior margin.
</paragraph>
<paragraph pageId="18" pageNumber="87">Abdomen: Terga pale grayish orange or pale grayish brown; sterna lustrous, white or pale orange.</paragraph>
<paragraph pageId="18" pageNumber="87">Male genitalia (Figs 49-52): Tegumen triangular, with bifid, setose apex; tuba analis arising from dorsoposterior area of tegumen. Valva divided into two portions; costal portion broadened in basal 1/2, triangular in distal 1/3; distal margin of cucullus slightly emarginated medially, with dense long setae; saccular portion elongate, spatulate, densely setose. Anellus conical, nearly as long as phallus, with minute thorns on interior wall. Vinculum elongate-subrectangular, with semi-elliptical emargination anteromedially; saccus elongate, as long as uncus. Phallus slightly curved at distal 1/3, broadened anteriorly.</paragraph>
<paragraph pageId="18" pageNumber="87">
Female genitalia (Figs 62-62): Apophyses posteriores 2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than apophyses anteriores. Lamella antevaginalis conical, obliquely truncate apically, setose posterolaterally. Ductus bursae as long as corpus bursae, narrow; inception of ductus seminalis at posterior 1/4 of ductus bursae; ductus seminalis coiled. Corpus bursae obovate, with scattered microscopic thorns.
</paragraph>
<caption pageId="18" pageNumber="87">
<paragraph pageId="18" pageNumber="87">
Figures 57-60.
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="genus">Philonome</taxonomicName>
, female genitalia. 57-58
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome nigrescens" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="nigrescens">Philonome nigrescens</taxonomicName>
57 Ventral view 58 Lateral view of segment 8 and sterigma 59-60
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome clemensella" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="clemensella">Philonome clemensella</taxonomicName>
59 Ventral view 60 Lateral view of segment 8 and sterigma.
</paragraph>
</caption>
<caption pageId="18" pageNumber="87">
<paragraph pageId="18" pageNumber="87">
Figures 61-66.
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="genus">Philonome</taxonomicName>
, female genitalia. 61-62
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera</taxonomicName>
61 Ventral view 62 Lateral view of segment 8 and sterigma 63-64
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome kawakitai" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="kawakitai">Philonome kawakitai</taxonomicName>
63 Ventral view 64 Lateral view 65-66
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="18" pageNumber="87" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
65 Ventral view 66 Lateral view of segment 8 and sterigma.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="88" pageId="18" pageNumber="87" type="types">
<paragraph pageId="18" pageNumber="87">Types.</paragraph>
<paragraph lastPageId="19" lastPageNumber="88" pageId="18" pageNumber="87">
Holotype: male,
<normalizedToken originalValue="“Holo-Type”">&quot;Holo-Type&quot;</normalizedToken>
[circular label with red border], &quot;Para Brazil Parish 6 -19.&quot;, &quot;
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="19" pageNumber="88" phylum="Arthropoda" rank="species" species="rivifera">
<pageBreakToken pageId="19" pageNumber="88" start="start">Philonome</pageBreakToken>
rivifera
</taxonomicName>
7/17 Meyr. E. Meyrick det. in Meyrick Coll.&quot; [old label attached before this study], &quot;Meyrick Coll. B.M. 1988-290.&quot;, &quot;B.M. ♂ Genitalia slide No. 32828&quot;. Paratypes (1♂, 4♀, 1 ex.): Brazil: Amazonas: Munaos [= Manaus], 2♀,
<normalizedToken originalValue="“11.19”">&quot;11.19&quot;</normalizedToken>
(Parish), BMNH.
<normalizedToken originalValue="Tefé">Tefe</normalizedToken>
, 1 ex.,
<normalizedToken originalValue="“1.20”">&quot;1.20&quot;</normalizedToken>
(Parish), BMNH. Federal District: Planaltina (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-15.583333">15°35'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-47.7">47°42'W</geoCoordinate>
, alt. 1000m): 1♀, 3 May 1984 (VO Becker), &quot;BECKER 56394&quot;, VOB; 1♀, 15 June 1985, &quot;BECKER 57636&quot;, [GSN] USNM 34615, USNM.
<normalizedToken originalValue="Pará">Para</normalizedToken>
:
<normalizedToken originalValue="Óbidos">Obidos</normalizedToken>
, 1♂,
<normalizedToken originalValue="“9.19”">&quot;9.19&quot;</normalizedToken>
(Parish), BMNH.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="88" type="distribution">
<paragraph pageId="19" pageNumber="88">Distribution.</paragraph>
<paragraph pageId="19" pageNumber="88">
Brazil (Amazonas, Federal District,
<normalizedToken originalValue="Pará">Para</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="88" type="etymology">
<paragraph pageId="19" pageNumber="88">Etymology.</paragraph>
<paragraph pageId="19" pageNumber="88">
The species name is derived from the Latin prefix 'pene (= paene)', meaning
<normalizedToken originalValue="“almost”">&quot;almost&quot;</normalizedToken>
, and the preexisting species name, rivifera, and refers to the overall similarity of this species to
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="19" pageNumber="88" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="88" type="remarks">
<paragraph pageId="19" pageNumber="88">Remarks.</paragraph>
<paragraph pageId="19" pageNumber="88">
The holotype and three paratypes of
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome penerivifera" order="Lepidoptera" pageId="19" pageNumber="88" phylum="Arthropoda" rank="species" species="penerivifera">Philonome penerivifera</taxonomicName>
in the BMNH collection were misidentified as
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome rivifera" order="Lepidoptera" pageId="19" pageNumber="88" phylum="Arthropoda" rank="species" species="rivifera">Philonome rivifera</taxonomicName>
by Edward Meyrick.
</paragraph>
</subSubSection>
</treatment>
</document>